Multi-ancestry genome-wide gene-smoking interaction study of 387,272 individuals identifies new loci associated with serum lipids.


Journal

Nature genetics
ISSN: 1546-1718
Titre abrégé: Nat Genet
Pays: United States
ID NLM: 9216904

Informations de publication

Date de publication:
04 2019
Historique:
received: 08 01 2018
accepted: 07 02 2019
entrez: 31 3 2019
pubmed: 31 3 2019
medline: 20 4 2019
Statut: ppublish

Résumé

The concentrations of high- and low-density-lipoprotein cholesterol and triglycerides are influenced by smoking, but it is unknown whether genetic associations with lipids may be modified by smoking. We conducted a multi-ancestry genome-wide gene-smoking interaction study in 133,805 individuals with follow-up in an additional 253,467 individuals. Combined meta-analyses identified 13 new loci associated with lipids, some of which were detected only because association differed by smoking status. Additionally, we demonstrate the importance of including diverse populations, particularly in studies of interactions with lifestyle factors, where genomic and lifestyle differences by ancestry may contribute to novel findings.

Identifiants

pubmed: 30926973
doi: 10.1038/s41588-019-0378-y
pii: 10.1038/s41588-019-0378-y
pmc: PMC6467258
mid: NIHMS1521094
doi:

Substances chimiques

Lipids 0

Types de publication

Journal Article Meta-Analysis Research Support, N.I.H., Extramural Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

636-648

Subventions

Organisme : NIDDK NIH HHS
ID : R01 DK093757
Pays : United States
Organisme : NIEHS NIH HHS
ID : P30 ES007048
Pays : United States
Organisme : NIA NIH HHS
ID : U01 AG009740
Pays : United States
Organisme : Intramural NIH HHS
ID : Z01 HG200362
Pays : United States
Organisme : Medical Research Council
ID : MC_UU_12015/1
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : R01 HL142302
Pays : United States
Organisme : Medical Research Council
ID : MR/L01341X/1
Pays : United Kingdom
Organisme : NIA NIH HHS
ID : U01 AG023746
Pays : United States
Organisme : NCI NIH HHS
ID : U01 CA182913
Pays : United States
Organisme : Intramural NIH HHS
ID : ZIA HG200362-02
Pays : United States
Organisme : Biotechnology and Biological Sciences Research Council
ID : BB/F019394/1
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : K01 HL135405
Pays : United States
Organisme : NHLBI NIH HHS
ID : R01 HL120393
Pays : United States
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ID : R01 HL046380
Pays : United States
Organisme : NHLBI NIH HHS
ID : U01 HL120393
Pays : United States
Organisme : NHLBI NIH HHS
ID : R01 HL113338
Pays : United States
Organisme : NIDDK NIH HHS
ID : R01 DK117445
Pays : United States
Organisme : NIDDK NIH HHS
ID : R01 DK072193
Pays : United States
Organisme : NIDDK NIH HHS
ID : P30 DK079626
Pays : United States
Organisme : NIDDK NIH HHS
ID : R01 DK107786
Pays : United States
Organisme : NCI NIH HHS
ID : UM1 CA173640
Pays : United States
Organisme : NHLBI NIH HHS
ID : R01 HL119443
Pays : United States
Organisme : NHLBI NIH HHS
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Organisme : NHLBI NIH HHS
ID : R01 HL105756
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Organisme : NHLBI NIH HHS
ID : R35 HL135818
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Organisme : Medical Research Council
ID : MR/R023484/1
Pays : United Kingdom
Organisme : Intramural NIH HHS
ID : Z99 HG999999
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ID : MR/L01632X/1
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ID : MR/K002414/1
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ID : U01 HL137162
Pays : United States
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ID : MC_UU_00007/10
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ID : U01 DK062370
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Organisme : Medical Research Council
ID : MC_UP_1605/7
Pays : United Kingdom
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ID : R21 HL123677
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ID : R01 MD012765
Pays : United States
Organisme : NCI NIH HHS
ID : UM1 CA182910
Pays : United States
Organisme : NICHD NIH HHS
ID : P2C HD050924
Pays : United States

Références

Willer, C. J. et al. Discovery and refinement of loci associated with lipid levels. Nat. Genet. 45, 1274–1283 (2013).
doi: 10.1038/ng.2797
Do, R. et al. Common variants associated with plasma triglycerides and risk for coronary artery disease. Nat. Genet. 45, 1345–1352 (2013).
doi: 10.1038/ng.2795
Peloso, G. M. et al. Association of low-frequency and rare coding-sequence variants with blood lipids and coronary heart disease in 56,000 whites and blacks. Am. J. Hum. Genet. 94, 223–232 (2014).
doi: 10.1016/j.ajhg.2014.01.009
Spracklen, C. N. et al. Association analyses of East Asian individuals and trans-ancestry analyses with European individuals reveal new loci associated with cholesterol and triglyceride levels. Hum. Mol. Genet. 26, 1770–1784 (2017).
doi: 10.1093/hmg/ddx062
Teslovich, T. M. et al. Biological, clinical and population relevance of 95 loci for blood lipids. Nature 466, 707–713 (2010).
doi: 10.1038/nature09270
Kathiresan, S. et al. Polymorphisms associated with cholesterol and risk of cardiovascular events. N. Engl. J. Med. 358, 1240–1249 (2008).
doi: 10.1056/NEJMoa0706728
Kar, D. et al. Relationship of cardiometabolic parameters in non-smokers, current smokers, and quitters in diabetes: a systematic review and meta-analysis. Cardiovasc. Diabetol. 15, 158 (2016).
doi: 10.1186/s12933-016-0475-5
Zong, C. et al. Cigarette smoke exposure impairs reverse cholesterol transport which can be minimized by treatment of hydrogen-saturated saline. Lipids Health Dis. 14, 159 (2015).
doi: 10.1186/s12944-015-0160-9
Manning, A. K. et al. Meta-analysis of gene–environment interaction: joint estimation of SNP and SNP × environment regression coefficients. Genet. Epidemiol. 35, 11–18 (2011).
doi: 10.1002/gepi.20546
Psaty, B. M. et al. Cohorts for Heart and Aging Research in Genomic Epidemiology (CHARGE) Consortium: design of prospective meta-analyses of genome-wide association studies from five cohorts. Circ. Cardiovasc. Genet. 2, 73–80 (2009).
doi: 10.1161/CIRCGENETICS.108.829747
Rao, D. C. et al. Multiancestry study of gene–lifestyle interactions for cardiovascular traits in 610 475 individuals from 124 cohorts: design and rationale. Circ. Cardiovasc. Genet. 10, e001649 (2017).
Lanktree, M. B. et al. Genetic meta-analysis of 15,901 African Americans identifies variation in EXOC3L1 is associated with HDL concentration. J. Lipid Res. 56, 1781–1786 (2015).
doi: 10.1194/jlr.P059477
Pers, T. H. et al. Biological interpretation of genome-wide association studies using predicted gene functions. Nat. Commun. 6, 5890–5890 (2015).
doi: 10.1038/ncomms6890
Suzuki, J., Imanishi, E. & Nagata, S. Xkr8 phospholipid scrambling complex in apoptotic phosphatidylserine exposure. Proc. Natl Acad. Sci. USA 113, 9509–9514 (2016).
doi: 10.1073/pnas.1610403113
Wang, J. et al. Genome-wide expression analysis reveals diverse effects of acute nicotine exposure on neuronal function-related genes and pathways. Front. Psychiatry 2, 5 (2011).
pubmed: 21556275 pmcid: 3089989
International Parkinson Disease Genomics Consortium. Imputation of sequence variants for identification of genetic risks for Parkinson’s disease: a meta-analysis of genome-wide association studies. Lancet 377, 641–649 (2011).
doi: 10.1016/S0140-6736(10)62345-8
Ng, M. C. Y. et al. Meta-analysis of genome-wide association studies in African Americans provides insights into the genetic architecture of type 2 diabetes. PLoS Genet. 10, e1004517 (2014).
doi: 10.1371/journal.pgen.1004517
Cai, K., Lucki, N. C. & Sewer, M. B. Silencing diacylglycerol kinase-θ expression reduces steroid hormone biosynthesis and cholesterol metabolism in human adrenocortical cells. Biochim. Biophys. Acta 1841, 552–562 (2014).
doi: 10.1016/j.bbalip.2013.12.005
Cai, K. & Sewer, M. B. Diacylglycerol kinase θ couples farnesoid X receptor–dependent bile acid signalling to Akt activation and glucose homoeostasis in hepatocytes. Biochem. J. 454, 267–274 (2013).
doi: 10.1042/BJ20130609
Cordell, H. J. et al. International genome-wide meta-analysis identifies new primary biliary cirrhosis risk loci and targetable pathogenic pathways. Nat. Commun. 6, 8019 (2015).
doi: 10.1038/ncomms9019
Edwards, T. L. et al. Genome-wide association study confirms SNPs in SNCA and the MAPT region as common risk factors for Parkinson disease. Ann. Hum. Genet. 74, 97–109 (2010).
doi: 10.1111/j.1469-1809.2009.00560.x
Lill, C. M. et al. Comprehensive research synopsis and systematic meta-analyses in Parkinson’s disease genetics: the PDGene Database. PLoS Genet. 8, e1002548 (2012).
doi: 10.1371/journal.pgen.1002548
Nalls, M. A. et al. Large-scale meta-analysis of genome-wide association data identifies six new risk loci for Parkinson’s disease. Nat. Genet. 46, 989–993 (2014).
doi: 10.1038/ng.3043
Pankratz, N. et al. Meta-analysis of Parkinson disease: identification of a novel locus, RIT2. Ann. Neurol. 71, 370–384 (2012).
doi: 10.1002/ana.22687
Wang, J. et al. Phlegm-dampness constitution: genomics, susceptibility, adjustment and treatment with traditional Chinese medicine. Am. J. Chin. Med. 41, 253–262 (2013).
doi: 10.1142/S0192415X13500183
Choi, J.-H. et al. Variations in TAS1R taste receptor gene family modify food intake and gastric cancer risk in a Korean population. Mol. Nutr. Food Res. 60, 2433–2445 (2016).
doi: 10.1002/mnfr.201600145
Hoffmann, T. J. et al. A large electronic-health-record-based genome-wide study of serum lipids. Nat. Genet. 50, 401–413 (2018).
doi: 10.1038/s41588-018-0064-5
Klarin, D. et al. Genetics of blood lipids among ~300,000 multi-ethnic participants of the Million Veteran Program. Nat. Genet. 50, 1514–1523 (2018).
doi: 10.1038/s41588-018-0222-9
Liu, D. J. et al. Exome-wide association study of plasma lipids in ~300,000 individuals. Nat. Genet. 49, 1758 (2017).
doi: 10.1038/ng.3977
Ward, L. D. & Kellis, M. HaploReg: a resource for exploring chromatin states, conservation, and regulatory motif alterations within sets of genetically linked variants. Nucleic Acids Res. 40, D930–D934 (2012).
doi: 10.1093/nar/gkr917
Tobacco Use Among U.S. Racial/Ethnic Minority Groups—African Americans, American Indians and Alaska Natives, Asian Americans and Pacific Islanders, and Hispanics: a Report of the Surgeon General (US Department of Health and Human Services, Centers for Disease Control and Prevention, National Center for Chronic Disease Prevention and Health Promotion, Office on Smoking and Health, 1998).
Villanti, A. C. et al. Changes in the prevalence and correlates of menthol cigarette use in the USA, 2004–2014. Tobacco Control 25, ii14 (2016).
doi: 10.1136/tobaccocontrol-2016-053329
Ross, K. C., Dempsey, D. A., St.Helen, G., Delucchi, K. & Benowitz, N. L. The influence of puff characteristics, nicotine dependence, and rate of nicotine metabolism on daily nicotine exposure in African American smokers. Cancer Epidemiol. Bomarkers Prev. 25, 936–943 (2016).
doi: 10.1158/1055-9965.EPI-15-1034
Ton, H. T. et al. Menthol enhances the desensitization of human α3β4 nicotinic acetylcholine receptors. Mol. Pharmacol. 88, 256–264 (2015).
doi: 10.1124/mol.115.098285
Alexander, L. A. et al. Why we must continue to investigate menthol’s role in the African American smoking paradox. Nicotine Tobacco Res. 18, S91–S101 (2016).
doi: 10.1093/ntr/ntv209
Jones, M. R., Tellez-Plaza, M. & Navas-Acien, A. Smoking, menthol cigarettes and all-cause, cancer and cardiovascular mortality: evidence from the National Health and Nutrition Examination Survey (NHANES) and a meta-analysis. PLoS One 8, e77941 (2013).
doi: 10.1371/journal.pone.0077941
Munro, H. M., Tarone, R. E., Wang, T. J. & Blot, W. J. Menthol and nonmenthol cigarette smoking: all-cause deaths, cardiovascular disease deaths, and other causes of death among blacks and whites. Circulation 133, 1861–1866 (2016).
doi: 10.1161/CIRCULATIONAHA.115.020536
Murray, R. P., Connett, J. E., Skeans, M. A. & Tashkin, D. P. Menthol cigarettes and health risks in lung health study data. Nicotine Tobacco Res. 9, 101–107 (2007).
doi: 10.1080/14622200601078418
Vozoris, N. T. Mentholated cigarettes and cardiovascular and pulmonary diseases: a population-based study. Arch. Intern. Med. 172, 590–593 (2012).
doi: 10.1001/archinternmed.2012.320
Pérez-Stable, E. J., Herrera, B., Jacob, I. P. & Benowitz, N. L. Nicotine metabolism and intake in black and white smokers. J. Am. Med. Assoc. 280, 152–156 (1998).
doi: 10.1001/jama.280.2.152
Khariwala, S. S. et al. Cotinine and tobacco-specific carcinogen exposure among nondaily smokers in a multiethnic sample. Nicotine Tobacco Res. 16, 600–605 (2014).
doi: 10.1093/ntr/ntt194
Jain, R. B. Distributions of selected urinary metabolites of volatile organic compounds by age, gender, race/ethnicity, and smoking status in a representative sample of U.S. adults. Environ. Toxicol. Pharmacol. 40, 471–479 (2015).
doi: 10.1016/j.etap.2015.07.018
Benowitz, N. L., Dains, K. M., Dempsey, D., Wilson, M. & Jacob, P. Racial differences in the relationship between number of cigarettes smoked and nicotine and carcinogen exposure. Nicotine Tobacco Res. 13, 772–783 (2011).
doi: 10.1093/ntr/ntr072
The Health Consequences of Smoking—50 Years of Progress: a Report of the Surgeon General (US Department of Health and Human Services, Centers for Disease Control and Prevention, National Center for Chronic Disease Prevention and Health Promotion, Office on Smoking and Health, 2014).
Ito, S. et al. Nicotine-induced expression of low-density lipoprotein receptor in oral epithelial cells. PLoS One 8, e82563 (2013).
doi: 10.1371/journal.pone.0082563
Dullaart, R. P., Hoogenberg, K., Dikkeschei, B. D. & van Tol, A. Higher plasma lipid transfer protein activities and unfavorable lipoprotein changes in cigarette-smoking men. Arterioscler. Thromb. 14, 1581–1585 (1994).
doi: 10.1161/01.ATV.14.10.1581
Frondelius, K. et al. Lifestyle and dietary determinants of serum apolipoprotein A1 and apolipoprotein B concentrations: cross-sectional analyses within a Swedish cohort of 24,984 individuals. Nutrients 9, 211 (2017).
doi: 10.3390/nu9030211
Onat, A. et al. Preheparin serum lipoprotein lipase mass interacts with gender, gene polymorphism and, positively, with smoking. Clin. Chem. Lab. Med. 47, 208 (2009).
pubmed: 19191728
Winkler, T. W. et al. Quality control and conduct of genome-wide association meta-analyses. Nat. Protoc. 9, 1192–1212 (2014).
doi: 10.1038/nprot.2014.071
Winkler, T. W. et al. EasyStrata: evaluation and visualization of stratified genome-wide association meta-analysis data. Bioinformatics 31, 259–261 (2015).
doi: 10.1093/bioinformatics/btu621
Marchini, J. & Howie, B. Genotype imputation for genome-wide association studies. Nat. Rev. Genet. 11, 499–511 (2010).
doi: 10.1038/nrg2796
Kraft, P., Yen, Y. C., Stram, D. O., Morrison, J. & Gauderman, W. J. Exploiting gene–environment interaction to detect genetic associations. Hum. Hered. 63, 111–119 (2007).
doi: 10.1159/000099183
Skol, A. D., Scott, L. J., Abecasis, G. R. & Boehnke, M. Joint analysis is more efficient than replication-based analysis for two-stage genome-wide association studies. Nat. Genet. 38, 209–213 (2006).
doi: 10.1038/ng1706
Das, S. et al. Next-generation genotype imputation service and methods. Nat. Genet. 48, 1284 (2016).
doi: 10.1038/ng.3656
Winkler, T. W. et al. Approaches to detect genetic effects that differ between two strata in genome-wide meta-analyses: recommendations based on a systematic evaluation. PLoS One 12, e0181038 (2017).
doi: 10.1371/journal.pone.0181038
Nikpay, M. et al. A comprehensive 1000 Genomes–based genome-wide association meta-analysis of coronary artery disease. Nat. Genet. 47, 1121–1130 (2015).
doi: 10.1038/ng.3396
Wood, A. R. et al. Defining the role of common variation in the genomic and biological architecture of adult human height. Nat. Genet. 46, 1173–1186 (2014).
doi: 10.1038/ng.3097
Locke, A. E. et al. Genetic studies of body mass index yield new insights for obesity biology. Nature 518, 197 (2015).
doi: 10.1038/nature14177
Shungin, D. et al. New genetic loci link adipose and insulin biology to body fat distribution. Nature 518, 187 (2015).
doi: 10.1038/nature14132
Justice, A. E. et al. Genome-wide meta-analysis of 241,258 adults accounting for smoking behaviour identifies novel loci for obesity traits. Nat. Commun. 8, 14977 (2017).
doi: 10.1038/ncomms14977
Manning, A. K. et al. A genome-wide approach accounting for body mass index identifies genetic variants influencing fasting glycemic traits and insulin resistance. Nat. Genet. 44, 659–669 (2012).
doi: 10.1038/ng.2274
Liu, C. T. et al. Trans-ethnic meta-analysis and functional annotation illuminates the genetic architecture of fasting glucose and insulin. Am. J. Hum. Genet. 99, 56–75 (2016).
doi: 10.1016/j.ajhg.2016.05.006
Gaulton, K. J. et al. Genetic fine-mapping and genomic annotation defines causal mechanisms at type 2 diabetes susceptibility loci. Nat. Genet. 47, 1415–1425 (2015).
doi: 10.1038/ng.3437
Genome-wide trans-ancestry meta-analysis provides insight into the genetic architecture of type 2 diabetes susceptibility. Nat. Genet. 46, 234–244 (2014).
Mahajan, A. et al. Trans-ethnic fine mapping highlights kidney-function genes linked to salt sensitivity. Am. J. Hum. Genet. 99, 636–646 (2016).
doi: 10.1016/j.ajhg.2016.07.012
Joehanes, R. et al. Integrated genome-wide analysis of expression quantitative trait loci aids interpretation of genomic association studies. Genome Biol. 18, 16 (2017).
doi: 10.1186/s13059-016-1142-6
Blake, J. A. et al. The Mouse Genome Database: integration of and access to knowledge about the laboratory mouse. Nucleic Acids Res. 42, D810–D817 (2014).
doi: 10.1093/nar/gkt1225
Lage, K. et al. A human phenome-interactome network of protein complexes implicated in genetic disorders. Nat. Biotechnol. 25, 309–316 (2007).
doi: 10.1038/nbt1295

Auteurs

Amy R Bentley (AR)

Center for Research on Genomics and Global Health, National Human Genome Research Institute, US National Institutes of Health, Bethesda, MD, USA. amy.bentley@nih.gov.

Yun J Sung (YJ)

Division of Biostatistics, Washington University School of Medicine, St. Louis, MO, USA.

Michael R Brown (MR)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, University of Texas Health Science Center at Houston, Houston, TX, USA.

Thomas W Winkler (TW)

Department of Genetic Epidemiology, University of Regensburg, Regensburg, Germany.

Aldi T Kraja (AT)

Division of Statistical Genomics, Department of Genetics, Washington University School of Medicine, St. Louis, MO, USA.

Ioanna Ntalla (I)

Clinical Pharmacology, William Harvey Research Institute, Barts and The London School of Medicine and Dentistry, Queen Mary University of London, London, UK.

Karen Schwander (K)

Division of Biostatistics, Washington University School of Medicine, St. Louis, MO, USA.

Daniel I Chasman (DI)

Division of Preventive Medicine, Brigham and Women's Hospital, Boston, MA, USA.
Harvard Medical School, Boston, MA, USA.

Elise Lim (E)

Department of Biostatistics, Boston University School of Public Health, Boston, MA, USA.

Xuan Deng (X)

Department of Biostatistics, Boston University School of Public Health, Boston, MA, USA.

Xiuqing Guo (X)

Institute for Translational Genomics and Population Sciences, Department of Pediatrics, Los Angeles Biomedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.

Jingmin Liu (J)

Women's Health Initiative Clinical Coordinating Center, Fred Hutchinson Cancer Research Center, Seattle, WA, USA.

Yingchang Lu (Y)

Charles Bronfman Institute for Personalized Medicine, Icahn School of Medicine at Mount Sinai, New York, NY, USA.

Ching-Yu Cheng (CY)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, Singapore.
Centre for Quantitative Medicine, Academic Medicine Research Institute, Ophthalmology and Visual Sciences Academic Clinical Program (Eye ACP), Duke-NUS Medical School, Singapore, Singapore.
Department of Ophthalmology, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.

Xueling Sim (X)

Saw Swee Hock School of Public Health, National University of Singapore and National University Health System, Singapore, Singapore.

Dina Vojinovic (D)

Department of Epidemiology, Erasmus University Medical Center, Rotterdam, the Netherlands.

Jennifer E Huffman (JE)

Medical Research Council Human Genetics Unit, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, UK.

Solomon K Musani (SK)

Jackson Heart Study, Department of Medicine, University of Mississippi Medical Center, Jackson, MS, USA.

Changwei Li (C)

Epidemiology and Biostatistics, University of Georgia at Athens College of Public Health, Athens, GA, USA.

Mary F Feitosa (MF)

Division of Statistical Genomics, Department of Genetics, Washington University School of Medicine, St. Louis, MO, USA.

Melissa A Richard (MA)

Brown Foundation Institute of Molecular Medicine, University of Texas Health Science Center at Houston, Houston, TX, USA.

Raymond Noordam (R)

Internal Medicine, Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands.

Jenna Baker (J)

Center for Research on Genomics and Global Health, National Human Genome Research Institute, US National Institutes of Health, Bethesda, MD, USA.

Guanjie Chen (G)

Center for Research on Genomics and Global Health, National Human Genome Research Institute, US National Institutes of Health, Bethesda, MD, USA.

Hugues Aschard (H)

Centre de Bioinformatique, Biostatistique, et Biologie Intégrative (C3BI), Institut Pasteur, Paris, France.
Department of Epidemiology, Harvard T. H. Chan School of Public Health, Boston, MA, USA.

Traci M Bartz (TM)

Cardiovascular Health Research Unit, Department of Biostatistics, University of Washington, Seattle, WA, USA.
Cardiovascular Health Research Unit, Department of Medicine, University of Washington, Seattle, WA, USA.

Jingzhong Ding (J)

Center on Diabetes, Obesity, and Metabolism, Gerontology and Geriatric Medicine, Wake Forest University Health Sciences, Winston-Salem, NC, USA.

Rajkumar Dorajoo (R)

Genome Institute of Singapore, Agency for Science, Technology and Research, Singapore, Singapore.

Alisa K Manning (AK)

Clinical and Translational Epidemiology Unit, Massachusetts General Hospital, Boston, MA, USA.
Department of Medicine, Harvard Medical School, Boston, MA, USA.

Tuomo Rankinen (T)

Human Genomics Laboratory, Pennington Biomedical Research Center, Baton Rouge, LA, USA.

Albert V Smith (AV)

Icelandic Heart Association, Kopavogur, Iceland.
Faculty of Medicine, University of Iceland, Reykjavik, Iceland.

Salman M Tajuddin (SM)

Health Disparities Research Section, Laboratory of Epidemiology and Population Sciences, National Institute on Aging, US National Institutes of Health, Baltimore, MD, USA.

Wei Zhao (W)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.

Mariaelisa Graff (M)

Department of Epidemiology, Gillings School of Global Public Health, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.

Maris Alver (M)

Estonian Genome Center, Institute of Genomics, University of Tartu, Tartu, Estonia.

Mathilde Boissel (M)

CNRS UMR 8199, European Genomic Institute for Diabetes (EGID), Institut Pasteur de Lille, University of Lille, Lille, France.

Jin Fang Chai (JF)

Saw Swee Hock School of Public Health, National University of Singapore and National University Health System, Singapore, Singapore.

Xu Chen (X)

Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden.

Jasmin Divers (J)

Department of Biostatistics and Data Science, Division of Public Health Sciences, Wake Forest School of Medicine, Winston-Salem, NC, USA.

Evangelos Evangelou (E)

Department of Epidemiology and Biostatistics, School of Public Health, Imperial College, London, UK.
Department of Hygiene and Epidemiology, University of Ioannina Medical School, Ioannina, Greece.

Chuan Gao (C)

Molecular Genetics and Genomics Program, Wake Forest School of Medicine, Winston-Salem, NC, USA.

Anuj Goel (A)

Division of Cardiovascular Medicine, Radcliffe Department of Medicine, University of Oxford, Oxford, UK.
Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.

Yanick Hagemeijer (Y)

University of Groningen, University Medical Center Groningen, Department of Cardiology, Groningen, the Netherlands.

Sarah E Harris (SE)

Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, UK.
Medical Genetics Section, Centre for Genomic and Experimental Medicine, University of Edinburgh, Edinburgh, UK.

Fernando P Hartwig (FP)

Postgraduate Programme in Epidemiology, Federal University of Pelotas, Pelotas, Brazil.
Medical Research Council Integrative Epidemiology Unit, University of Bristol, Bristol, UK.

Meian He (M)

Department of Occupational and Environmental Health and State Key Laboratory of Environmental Health for Incubating, Tongji Medical College, Huazhong University of Science and Technology, Wuhan, China.

Andrea R V R Horimoto (ARVR)

Laboratory of Genetics and Molecular Cardiology, Heart Institute (InCor), University of São Paulo Medical School, São Paulo, Brazil.

Fang-Chi Hsu (FC)

Department of Biostatistics and Data Science, Division of Public Health Sciences, Wake Forest School of Medicine, Winston-Salem, NC, USA.

Yi-Jen Hung (YJ)

Endocrinology and Metabolism, Tri-Service General Hospital, Taipei, Taiwan.
School of Medicine, National Defense Medical Center, Taipei, Taiwan.

Anne U Jackson (AU)

Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, MI, USA.

Anuradhani Kasturiratne (A)

Department of Public Health, Faculty of Medicine, University of Kelaniya, Ragama, Sri Lanka.

Pirjo Komulainen (P)

Foundation for Research in Health, Exercise, and Nutrition, Kuopio Research Institute of Exercise Medicine, Kuopio, Finland.

Brigitte Kühnel (B)

Research Unit of Molecular Epidemiology, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.
Institute of Epidemiology, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.

Karin Leander (K)

Unit of Cardiovascular Epidemiology, Institute of Environmental Medicine, Karolinska Institutet, Stockholm, Sweden.

Keng-Hung Lin (KH)

Ophthalmology, Taichung Veterans General Hospital, Taichung, Taiwan.

Jian'an Luan (J)

Medical Research Council Epidemiology Unit, University of Cambridge, Cambridge, UK.

Leo-Pekka Lyytikäinen (LP)

Department of Clinical Chemistry, Fimlab Laboratories, Tampere, Finland.
Department of Clinical Chemistry, Finnish Cardiovascular Research Center-Tampere, Faculty of Medicine and Technology, Tampere University, Tampere, Finland.

Nana Matoba (N)

Laboratory for Statistical Analysis, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan.

Ilja M Nolte (IM)

University of Groningen, University Medical Center Groningen, Department of Epidemiology, Groningen, the Netherlands.

Maik Pietzner (M)

DZHK (German Centre for Cardiovascular Health), Partner Site Greifswald, Greifswald, Germany.
Institute of Clinical Chemistry and Laboratory Medicine, University Medicine Greifswald, Greifswald, Germany.

Bram Prins (B)

Human Genetics, Wellcome Trust Sanger Institute, Hinxton, UK.

Muhammad Riaz (M)

Department of Cardiovascular Sciences, University of Leicester, Leicester, UK.
NIHR Leicester Biomedical Research Centre, Glenfield Hospital, Leicester, UK.

Antonietta Robino (A)

Institute for Maternal and Child Health, IRCCS 'Burlo Garofolo', Trieste, Italy.

M Abdullah Said (MA)

University of Groningen, University Medical Center Groningen, Department of Cardiology, Groningen, the Netherlands.

Nicole Schupf (N)

Taub Institute for Research on Alzheimer's Disease and the Aging Brain, Columbia University Medical Center, New York, NY, USA.

Robert A Scott (RA)

Medical Research Council Epidemiology Unit, University of Cambridge, Cambridge, UK.

Tamar Sofer (T)

Department of Medicine, Harvard Medical School, Boston, MA, USA.
Division of Sleep and Circadian Disorders, Brigham and Women's Hospital, Boston, MA, USA.

Alena Stancáková (A)

Institute of Clinical Medicine, Internal Medicine, University of Eastern Finland, Kuopio, Finland.

Fumihiko Takeuchi (F)

Department of Gene Diagnostics and Therapeutics, Research Institute, National Center for Global Health and Medicine, Tokyo, Japan.

Bamidele O Tayo (BO)

Department of Public Health Sciences, Loyola University Chicago, Maywood, IL, USA.

Peter J van der Most (PJ)

University of Groningen, University Medical Center Groningen, Department of Epidemiology, Groningen, the Netherlands.

Tibor V Varga (TV)

Department of Clinical Sciences, Genetic and Molecular Epidemiology Unit, Lund University Diabetes Centre, Skåne University Hospital, Malmö, Sweden.

Tzung-Dau Wang (TD)

Cardiovascular Center, National Taiwan University Hospital, Taipei, Taiwan.
National Taiwan University College of Medicine, Taipei, Taiwan.
Division of Cardiology, Department of Internal Medicine, National Taiwan University Hospital, Taipei, Taiwan.

Yajuan Wang (Y)

Department of Population Quantitative and Health Sciences, Case Western Reserve University, Cleveland, OH, USA.

Erin B Ware (EB)

Survey Research Center, Institute for Social Research, University of Michigan, Ann Arbor, MI, USA.

Wanqing Wen (W)

Division of Epidemiology, Department of Medicine, Vanderbilt University School of Medicine, Nashville, TN, USA.

Yong-Bing Xiang (YB)

SKLORG and Department of Epidemiology, Shanghai Cancer Institute, Renji Hospital, Shanghai Jiaotong University School of Medicine, Shanghai, China.

Lisa R Yanek (LR)

Division of General Internal Medicine, Department of Medicine, Johns Hopkins University School of Medicine, Baltimore, MD, USA.

Weihua Zhang (W)

MRC-PHE Centre for Environment and Health, School of Public Health, Imperial College, London, UK.
Department of Cardiology, Ealing Hospital, Middlesex, UK.

Jing Hua Zhao (JH)

Medical Research Council Epidemiology Unit, University of Cambridge, Cambridge, UK.

Adebowale Adeyemo (A)

Center for Research on Genomics and Global Health, National Human Genome Research Institute, US National Institutes of Health, Bethesda, MD, USA.

Saima Afaq (S)

MRC-PHE Centre for Environment and Health, School of Public Health, Imperial College, London, UK.

Najaf Amin (N)

Department of Epidemiology, Erasmus University Medical Center, Rotterdam, the Netherlands.

Marzyeh Amini (M)

University of Groningen, University Medical Center Groningen, Department of Epidemiology, Groningen, the Netherlands.

Dan E Arking (DE)

McKusick-Nathans Institute of Genetic Medicine, Johns Hopkins University School of Medicine, Baltimore, MD, USA.

Zorayr Arzumanyan (Z)

Institute for Translational Genomics and Population Sciences, Department of Pediatrics, Los Angeles Biomedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.

Tin Aung (T)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, Singapore.
Department of Ophthalmology, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.
Ophthalmology and Visual Sciences Academic Clinical Program (Eye ACP), Duke-NUS Medical School, Singapore, Singapore.

Christie Ballantyne (C)

Section of Cardiovascular Research, Baylor College of Medicine, Houston, TX, USA.
Houston Methodist Debakey Heart and Vascular Center, Houston, TX, USA.

R Graham Barr (RG)

Departments of Medicine and Epidemiology, Columbia University Medical Center, New York, NY, USA.

Lawrence F Bielak (LF)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.

Eric Boerwinkle (E)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, University of Texas Health Science Center at Houston, Houston, TX, USA.
Human Genome Sequencing Center, Baylor College of Medicine, Houston, TX, USA.

Erwin P Bottinger (EP)

Charles Bronfman Institute for Personalized Medicine, Icahn School of Medicine at Mount Sinai, New York, NY, USA.

Ulrich Broeckel (U)

Section of Genomic Pediatrics, Departments of Pediatrics, Medicine, and Physiology, Medical College of Wisconsin, Milwaukee, WI, USA.

Morris Brown (M)

Clinical Pharmacology, William Harvey Research Institute, Barts and The London School of Medicine and Dentistry, Queen Mary University of London, London, UK.
NIHR Barts Cardiovascular Biomedical Research Centre, Queen Mary University of London, London, UK.

Brian E Cade (BE)

Division of Sleep and Circadian Disorders, Brigham and Women's Hospital, Boston, MA, USA.

Archie Campbell (A)

Centre for Genomic and Experimental Medicine, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, UK.

Mickaël Canouil (M)

CNRS UMR 8199, European Genomic Institute for Diabetes (EGID), Institut Pasteur de Lille, University of Lille, Lille, France.

Sabanayagam Charumathi (S)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, Singapore.
Centre for Quantitative Medicine, Academic Medicine Research Institute, Ophthalmology and Visual Sciences Academic Clinical Program (Eye ACP), Duke-NUS Medical School, Singapore, Singapore.

Yii-Der Ida Chen (YI)

Institute for Translational Genomics and Population Sciences, Department of Pediatrics, Los Angeles Biomedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.

Kaare Christensen (K)

Danish Aging Research Center, Institute of Public Health, University of Southern Denmark, Odense, Denmark.

Maria Pina Concas (MP)

Institute for Maternal and Child Health, IRCCS 'Burlo Garofolo', Trieste, Italy.

John M Connell (JM)

Ninewells Hospital and Medical School, University of Dundee, Dundee, UK.

Lisa de Las Fuentes (L)

Division of Biostatistics, Washington University School of Medicine, St. Louis, MO, USA.
Cardiovascular Division, Department of Medicine, Washington University School of Medicine, St. Louis, MO, USA.

H Janaka de Silva (HJ)

Department of Medicine, Faculty of Medicine, University of Kelaniya, Ragama, Sri Lanka.

Paul S de Vries (PS)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, University of Texas Health Science Center at Houston, Houston, TX, USA.

Ayo Doumatey (A)

Center for Research on Genomics and Global Health, National Human Genome Research Institute, US National Institutes of Health, Bethesda, MD, USA.

Qing Duan (Q)

Department of Genetics, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.

Charles B Eaton (CB)

Department of Family Medicine and Epidemiology, Alpert Medical School of Brown University, Providence, RI, USA.

Ruben N Eppinga (RN)

University of Groningen, University Medical Center Groningen, Department of Cardiology, Groningen, the Netherlands.

Jessica D Faul (JD)

Survey Research Center, Institute for Social Research, University of Michigan, Ann Arbor, MI, USA.

James S Floyd (JS)

Cardiovascular Health Research Unit, Department of Medicine, University of Washington, Seattle, WA, USA.
Department of Epidemiology, University of Washington, Seattle, WA, USA.

Nita G Forouhi (NG)

Medical Research Council Epidemiology Unit, University of Cambridge, Cambridge, UK.

Terrence Forrester (T)

UWI Solutions for Developing Countries, University of the West Indies, Kingston, Jamaica.

Yechiel Friedlander (Y)

Braun School of Public Health, Hebrew University-Hadassah Medical Center, Jerusalem, Israel.

Ilaria Gandin (I)

Department of Medical Sciences, University of Trieste, Trieste, Italy.

He Gao (H)

Department of Epidemiology and Biostatistics, School of Public Health, Imperial College, London, UK.

Mohsen Ghanbari (M)

Department of Epidemiology, Erasmus University Medical Center, Rotterdam, the Netherlands.
Department of Genetics, School of Medicine, Mashhad University of Medical Sciences, Mashhad, Iran.

Sina A Gharib (SA)

Computational Medicine Core, Center for Lung Biology, UW Medicine Sleep Center, Department of Medicine, University of Washington, Seattle, WA, USA.

Bruna Gigante (B)

Unit of Cardiovascular Epidemiology, Institute of Environmental Medicine, Karolinska Institutet, Stockholm, Sweden.

Franco Giulianini (F)

Division of Preventive Medicine, Brigham and Women's Hospital, Boston, MA, USA.

Hans J Grabe (HJ)

Department of Psychiatry and Psychotherapy, University Medicine Greifswald, Greifswald, Germany.

C Charles Gu (CC)

Division of Biostatistics, Washington University School of Medicine, St. Louis, MO, USA.

Tamara B Harris (TB)

Laboratory of Epidemiology and Population Sciences, National Institute on Aging, US National Institutes of Health, Bethesda, MD, USA.

Sami Heikkinen (S)

Institute of Clinical Medicine, Internal Medicine, University of Eastern Finland, Kuopio, Finland.
Institute of Biomedicine, School of Medicine, University of Eastern Finland, Kuopio, Finland.

Chew-Kiat Heng (CK)

Department of Paediatrics, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.
Khoo Teck Puat-National University Children's Medical Institute, National University Health System, Singapore, Singapore.

Makoto Hirata (M)

Laboratory of Genome Technology, Human Genome Center, Institute of Medical Science, University of Tokyo, Minato-ku, Japan.

James E Hixson (JE)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, University of Texas Health Science Center at Houston, Houston, TX, USA.

M Arfan Ikram (MA)

Department of Epidemiology, Erasmus University Medical Center, Rotterdam, the Netherlands.
Department of Radiology and Nuclear Medicine, Erasmus University Medical Center, Rotterdam, the Netherlands.
Department of Neurology, Erasmus University Medical Center, Rotterdam, the Netherlands.

Yucheng Jia (Y)

Institute for Translational Genomics and Population Sciences, Department of Pediatrics, Los Angeles Biomedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.

Roby Joehanes (R)

Hebrew SeniorLife, Beth Israel Deaconess Medical Center, Harvard Medical School, Boston, MA, USA.
Framingham Heart Study, National Heart, Lung, and Blood Institute, US National Institutes of Health, Bethesda, MD, USA.

Craig Johnson (C)

Collaborative Health Studies Coordinating Center, University of Washington, Seattle, WA, USA.

Jost Bruno Jonas (JB)

Department of Ophthalmology, Medical Faculty Mannheim, University of Heidelberg, Mannheim, Germany.
Beijing Institute of Ophthalmology, Beijing Ophthalmology and Visual Science Key Laboratory, Beijing Tongren Eye Center, Capital Medical University, Beijing, China.

Anne E Justice (AE)

Department of Epidemiology, Gillings School of Global Public Health, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.

Tomohiro Katsuya (T)

Department of Clinical Gene Therapy, Osaka University Graduate School of Medicine, Suita, Japan.
Department of Geriatric and General Medicine, Osaka University Graduate School of Medicine, Suita, Japan.

Chiea Chuen Khor (CC)

Genome Institute of Singapore, Agency for Science, Technology and Research, Singapore, Singapore.

Tuomas O Kilpeläinen (TO)

Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark.
Department of Environmental Medicine and Public Health, Icahn School of Medicine at Mount Sinai, New York, NY, USA.

Woon-Puay Koh (WP)

Saw Swee Hock School of Public Health, National University of Singapore and National University Health System, Singapore, Singapore.
Duke-NUS Medical School, Singapore, Singapore.

Ivana Kolcic (I)

Department of Public Health, Department of Medicine, University of Split, Split, Croatia.

Charles Kooperberg (C)

Fred Hutchinson Cancer Research Center, University of Washington School of Public Health, Seattle, WA, USA.

Jose E Krieger (JE)

Laboratory of Genetics and Molecular Cardiology, Heart Institute (InCor), University of São Paulo Medical School, São Paulo, Brazil.

Stephen B Kritchevsky (SB)

Sticht Center for Healthy Aging and Alzheimer's Prevention, Department of Internal Medicine, Wake Forest School of Medicine, Winston-Salem, NC, USA.

Michiaki Kubo (M)

RIKEN Center for Integrative Medical Sciences, Yokohama, Japan.

Johanna Kuusisto (J)

Institute of Clinical Medicine, Internal Medicine, University of Eastern Finland, Kuopio, Finland.

Timo A Lakka (TA)

Foundation for Research in Health, Exercise, and Nutrition, Kuopio Research Institute of Exercise Medicine, Kuopio, Finland.
Institute of Biomedicine, School of Medicine, University of Eastern Finland, Kuopio, Finland.
Department of Clinical Physiology and Nuclear Medicine, Kuopio University Hospital, Kuopio, Finland.

Carl D Langefeld (CD)

Department of Biostatistics and Data Science, Division of Public Health Sciences, Wake Forest School of Medicine, Winston-Salem, NC, USA.

Claudia Langenberg (C)

Medical Research Council Epidemiology Unit, University of Cambridge, Cambridge, UK.

Lenore J Launer (LJ)

Laboratory of Epidemiology and Population Sciences, National Institute on Aging, US National Institutes of Health, Bethesda, MD, USA.

Benjamin Lehne (B)

Institute of Clinical Sciences, Department of Molecular Sciences, Imperial College, London, UK.

Cora E Lewis (CE)

Department of Medicine, University of Alabama at Birmingham, Birmingham, AL, USA.

Yize Li (Y)

Division of Biostatistics, Washington University School of Medicine, St. Louis, MO, USA.

Jingjing Liang (J)

Department of Population Quantitative and Health Sciences, Case Western Reserve University, Cleveland, OH, USA.

Shiow Lin (S)

Division of Statistical Genomics, Department of Genetics, Washington University School of Medicine, St. Louis, MO, USA.

Ching-Ti Liu (CT)

Department of Biostatistics, Boston University School of Public Health, Boston, MA, USA.

Jianjun Liu (J)

Genome Institute of Singapore, Agency for Science, Technology and Research, Singapore, Singapore.
Department of Medicine, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.

Kiang Liu (K)

Epidemiology, Department of Preventive Medicine, Northwestern University Feinberg School of Medicine, Chicago, IL, USA.

Marie Loh (M)

MRC-PHE Centre for Environment and Health, School of Public Health, Imperial College, London, UK.
Translational Laboratory in Genetic Medicine, Agency for Science, Technology, and Research, Singapore, Singapore.
Department of Biochemistry, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.
Lee Kong Chian School of Medicine, Nanyang Technological University, Singapore, Singapore.

Kurt K Lohman (KK)

Department of Biostatistics and Data Science, Division of Public Health Sciences, Wake Forest School of Medicine, Winston-Salem, NC, USA.

Tin Louie (T)

Department of Biostatistics, University of Washington, Seattle, WA, USA.

Anna Luzzi (A)

Institute for Translational Genomics and Population Sciences, Department of Pediatrics, Los Angeles Biomedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.

Reedik Mägi (R)

Estonian Genome Center, Institute of Genomics, University of Tartu, Tartu, Estonia.

Anubha Mahajan (A)

Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.

Ani W Manichaikul (AW)

Center for Public Health Genomics, University of Virginia, Charlottesville, VA, USA.

Colin A McKenzie (CA)

Tropical Metabolism Research Unit, Caribbean Institute for Health Research, University of the West Indies, Mona, Jamaica.

Thomas Meitinger (T)

Institute of Human Genetics, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.
Institute of Human Genetics, Technische Universität München, Munich, Germany.
Technische Universität München, Munich, Germany.

Andres Metspalu (A)

Estonian Genome Center, Institute of Genomics, University of Tartu, Tartu, Estonia.

Yuri Milaneschi (Y)

Department of Psychiatry, Amsterdam Neuroscience and Amsterdam Public Health Research Institute, Amsterdam University Medical Center, Vrije Universiteit, Amsterdam, the Netherlands.

Lili Milani (L)

Estonian Genome Center, Institute of Genomics, University of Tartu, Tartu, Estonia.

Karen L Mohlke (KL)

Department of Genetics, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.

Yukihide Momozawa (Y)

Laboratory for Genotyping Development, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan.

Andrew P Morris (AP)

Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.
Department of Biostatistics, University of Liverpool, Liverpool, UK.

Alison D Murray (AD)

Institute of Medical Sciences, Aberdeen Biomedical Imaging Centre, University of Aberdeen, Aberdeen, UK.

Mike A Nalls (MA)

Data Tecnica International, Glen Echo, MD, USA.
Laboratory of Neurogenetics, National Institute on Aging, Bethesda, MD, USA.

Matthias Nauck (M)

DZHK (German Centre for Cardiovascular Health), Partner Site Greifswald, Greifswald, Germany.
Institute of Clinical Chemistry and Laboratory Medicine, University Medicine Greifswald, Greifswald, Germany.

Christopher P Nelson (CP)

Department of Cardiovascular Sciences, University of Leicester, Leicester, UK.
NIHR Leicester Biomedical Research Centre, Glenfield Hospital, Leicester, UK.

Kari E North (KE)

Department of Epidemiology, Gillings School of Global Public Health, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.

Jeffrey R O'Connell (JR)

Division of Endocrinology, Diabetes, and Nutrition, University of Maryland School of Medicine, Baltimore, MD, USA.
Program for Personalized and Genomic Medicine, University of Maryland School of Medicine, Baltimore, MD, USA.

Nicholette D Palmer (ND)

Biochemistry, Wake Forest School of Medicine, Winston-Salem, NC, USA.

George J Papanicolau (GJ)

Epidemiology Branch, Division of Cardiovascular Sciences, National Heart, Lung, and Blood Institute, US National Institutes of Health, Bethesda, MD, USA.

Nancy L Pedersen (NL)

Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden.

Annette Peters (A)

Institute of Epidemiology, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.
Epidemiology, Faculty of Medicine, Institute for Medical Information Processing, Biometry, and Epidemiology, Ludwig Maximilian University, Munich, Germany.

Patricia A Peyser (PA)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.

Ozren Polasek (O)

Department of Public Health, Department of Medicine, University of Split, Split, Croatia.
Psychiatric Hospital 'Sveti Ivan', Zagreb, Croatia.
Gen-info, Ltd, Zagreb, Croatia.

Neil Poulter (N)

School of Public Health, Imperial College, London, UK.

Olli T Raitakari (OT)

Department of Clinical Physiology and Nuclear Medicine, Turku University Hospital, Turku, Finland.
Research Centre of Applied and Preventive Cardiovascular Medicine, University of Turku, Turku, Finland.

Alex P Reiner (AP)

Fred Hutchinson Cancer Research Center, University of Washington School of Public Health, Seattle, WA, USA.

Frida Renström (F)

Department of Clinical Sciences, Genetic and Molecular Epidemiology Unit, Lund University Diabetes Centre, Skåne University Hospital, Malmö, Sweden.
Department of Biobank Research, Umeå University, Umeå, Sweden.

Treva K Rice (TK)

Division of Biostatistics, Washington University School of Medicine, St. Louis, MO, USA.

Stephen S Rich (SS)

Center for Public Health Genomics, University of Virginia, Charlottesville, VA, USA.

Jennifer G Robinson (JG)

Department of Epidemiology and Medicine, University of Iowa, Iowa City, IA, USA.

Lynda M Rose (LM)

Division of Preventive Medicine, Brigham and Women's Hospital, Boston, MA, USA.

Frits R Rosendaal (FR)

Clinical Epidemiology, Leiden University Medical Center, Leiden, the Netherlands.

Igor Rudan (I)

Centre for Global Health Research, Usher Institute of Population Health Sciences and Informatics, University of Edinburgh, Edinburgh, UK.

Carsten O Schmidt (CO)

Institute for Community Medicine, University Medicine Greifswald, Greifswald, Germany.

Pamela J Schreiner (PJ)

Division of Epidemiology and Community Health, School of Public Health, University of Minnesota, Minneapolis, MN, USA.

William R Scott (WR)

Institute of Clinical Sciences, Department of Molecular Sciences, Imperial College, London, UK.
National Heart and Lung Institute, Imperial College, London, UK.

Peter Sever (P)

National Heart and Lung Institute, Imperial College, London, UK.

Yuan Shi (Y)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, Singapore.

Stephen Sidney (S)

Division of Research, Kaiser Permanente Northern California, Oakland, CA, USA.

Mario Sims (M)

Jackson Heart Study, Department of Medicine, University of Mississippi Medical Center, Jackson, MS, USA.

Jennifer A Smith (JA)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.
Survey Research Center, Institute for Social Research, University of Michigan, Ann Arbor, MI, USA.

Harold Snieder (H)

University of Groningen, University Medical Center Groningen, Department of Epidemiology, Groningen, the Netherlands.

John M Starr (JM)

Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, UK.
Alzheimer Scotland Dementia Research Centre, University of Edinburgh, Edinburgh, UK.

Konstantin Strauch (K)

Genetic Epidemiology, Faculty of Medicine, Institute for Medical Information Processing, Biometry, and Epidemiology, Ludwig Maximilian University, Munich, Germany.
Institute of Genetic Epidemiology, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.

Heather M Stringham (HM)

Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, MI, USA.

Nicholas Y Q Tan (NYQ)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, Singapore.

Hua Tang (H)

Department of Genetics, Stanford University, Stanford, CA, USA.

Kent D Taylor (KD)

Institute for Translational Genomics and Population Sciences, Department of Pediatrics, Los Angeles Biomedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.

Yik Ying Teo (YY)

Saw Swee Hock School of Public Health, National University of Singapore and National University Health System, Singapore, Singapore.
Genome Institute of Singapore, Agency for Science, Technology and Research, Singapore, Singapore.
Department of Statistics and Applied Probability, National University of Singapore, Singapore, Singapore.
Life Sciences Institute, National University of Singapore, Singapore, Singapore.
NUS Graduate School for Integrative Science and Engineering, National University of Singapore, Singapore, Singapore.

Yih Chung Tham (YC)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, Singapore.

Henning Tiemeier (H)

Department of Epidemiology, Erasmus University Medical Center, Rotterdam, the Netherlands.
Department of Social and Behavioral Sciences, Harvard T. H. Chan School of Public Health, Boston, MA, USA.

Stephen T Turner (ST)

Division of Nephrology and Hypertension, Mayo Clinic, Rochester, MN, USA.

André G Uitterlinden (AG)

Department of Epidemiology, Erasmus University Medical Center, Rotterdam, the Netherlands.
Department of Internal Medicine, Erasmus University Medical Center, Rotterdam, the Netherlands.

Diana van Heemst (D)

Internal Medicine, Gerontology and Geriatrics, Leiden University Medical Center, Leiden, the Netherlands.

Melanie Waldenberger (M)

Research Unit of Molecular Epidemiology, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.
Institute of Epidemiology, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.

Heming Wang (H)

Department of Medicine, Harvard Medical School, Boston, MA, USA.
Division of Sleep and Circadian Disorders, Brigham and Women's Hospital, Boston, MA, USA.

Lan Wang (L)

Department of Biostatistics, Boston University School of Public Health, Boston, MA, USA.

Lihua Wang (L)

Division of Statistical Genomics, Department of Genetics, Washington University School of Medicine, St. Louis, MO, USA.

Wen Bin Wei (WB)

Beijing Tongren Eye Center, Beijing Tongren Hospital, Capital Medical University, Beijing, China.

Christine A Williams (CA)

Division of Statistical Genomics, Department of Genetics, Washington University School of Medicine, St. Louis, MO, USA.

Gregory Wilson (G)

Jackson Heart Study, School of Public Health, Jackson State University, Jackson, MS, USA.

Mary K Wojczynski (MK)

Division of Statistical Genomics, Department of Genetics, Washington University School of Medicine, St. Louis, MO, USA.

Jie Yao (J)

Institute for Translational Genomics and Population Sciences, Department of Pediatrics, Los Angeles Biomedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.

Kristin Young (K)

Department of Epidemiology, Gillings School of Global Public Health, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.

Caizheng Yu (C)

Department of Occupational and Environmental Health and State Key Laboratory of Environmental Health for Incubating, Tongji Medical College, Huazhong University of Science and Technology, Wuhan, China.

Jian-Min Yuan (JM)

Department of Epidemiology, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA, USA.
Division of Cancer Control and Population Sciences, UPMC Hillman Cancer, University of Pittsburgh, Pittsburgh, PA, USA.

Jie Zhou (J)

Center for Research on Genomics and Global Health, National Human Genome Research Institute, US National Institutes of Health, Bethesda, MD, USA.

Alan B Zonderman (AB)

Behavioral Epidemiology Section, Laboratory of Epidemiology and Population Sciences, National Institute on Aging, US National Institutes of Health, Baltimore, MD, USA.

Diane M Becker (DM)

Division of General Internal Medicine, Department of Medicine, Johns Hopkins University School of Medicine, Baltimore, MD, USA.

Michael Boehnke (M)

Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, MI, USA.

Donald W Bowden (DW)

Biochemistry, Wake Forest School of Medicine, Winston-Salem, NC, USA.

John C Chambers (JC)

Department of Epidemiology and Biostatistics, School of Public Health, Imperial College, London, UK.
MRC-PHE Centre for Environment and Health, School of Public Health, Imperial College, London, UK.
Department of Cardiology, Ealing Hospital, Middlesex, UK.
Lee Kong Chian School of Medicine, Nanyang Technological University, Singapore, Singapore.
Imperial College Healthcare NHS Trust, London, UK.

Richard S Cooper (RS)

Department of Public Health Sciences, Loyola University Chicago, Maywood, IL, USA.

Ulf de Faire (U)

Unit of Cardiovascular Epidemiology, Institute of Environmental Medicine, Karolinska Institutet, Stockholm, Sweden.

Ian J Deary (IJ)

Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, UK.
Psychology, University of Edinburgh, Edinburgh, UK.

Paul Elliott (P)

MRC-PHE Centre for Environment and Health, School of Public Health, Imperial College, London, UK.

Tõnu Esko (T)

Estonian Genome Center, Institute of Genomics, University of Tartu, Tartu, Estonia.
Broad Institute of MIT and Harvard, Boston, MA, USA.

Martin Farrall (M)

Division of Cardiovascular Medicine, Radcliffe Department of Medicine, University of Oxford, Oxford, UK.
Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.

Paul W Franks (PW)

Department of Clinical Sciences, Genetic and Molecular Epidemiology Unit, Lund University Diabetes Centre, Skåne University Hospital, Malmö, Sweden.
Department of Nutrition, Harvard T. H. Chan School of Public Health, Harvard University, Boston, MA, USA.
Department of Public Health and Clinical Medicine, Umeå University, Umeå, Sweden.
OCDEM, Radcliffe Department of Medicine, University of Oxford, Oxford, UK.

Barry I Freedman (BI)

Nephrology, Internal Medicine, Wake Forest School of Medicine, Winston-Salem, NC, USA.

Philippe Froguel (P)

CNRS UMR 8199, European Genomic Institute for Diabetes (EGID), Institut Pasteur de Lille, University of Lille, Lille, France.
Department of Genomics of Common Disease, Imperial College, London, UK.

Paolo Gasparini (P)

Institute for Maternal and Child Health, IRCCS 'Burlo Garofolo', Trieste, Italy.
Department of Medical Sciences, University of Trieste, Trieste, Italy.

Christian Gieger (C)

Institute of Epidemiology, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.
German Center for Diabetes Research (DZD), Neuherberg, Germany.

Bernardo L Horta (BL)

Postgraduate Programme in Epidemiology, Federal University of Pelotas, Pelotas, Brazil.

Jyh-Ming Jimmy Juang (JJ)

National Taiwan University College of Medicine, Taipei, Taiwan.
Division of Cardiology, Department of Internal Medicine, National Taiwan University Hospital, Taipei, Taiwan.

Yoichiro Kamatani (Y)

Laboratory for Statistical Analysis, RIKEN Center for Integrative Medical Sciences, Yokohama, Japan.

Candace M Kammerer (CM)

Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA, USA.

Norihiro Kato (N)

Department of Gene Diagnostics and Therapeutics, Research Institute, National Center for Global Health and Medicine, Tokyo, Japan.

Jaspal S Kooner (JS)

MRC-PHE Centre for Environment and Health, School of Public Health, Imperial College, London, UK.
Department of Cardiology, Ealing Hospital, Middlesex, UK.
National Heart and Lung Institute, Imperial College, London, UK.
Imperial College Healthcare NHS Trust, London, UK.

Markku Laakso (M)

Institute of Clinical Medicine, Internal Medicine, University of Eastern Finland, Kuopio, Finland.

Cathy C Laurie (CC)

Department of Biostatistics, University of Washington, Seattle, WA, USA.

I-Te Lee (IT)

Endocrinology and Metabolism, Internal Medicine, Taichung Veterans General Hospital, Taichung, Taiwan.
School of Medicine, Chung Shan Medical University, Taichung, Taiwan.
School of Medicine, National Yang-Ming University, Taipei, Taiwan.

Terho Lehtimäki (T)

Department of Clinical Chemistry, Fimlab Laboratories, Tampere, Finland.
Department of Clinical Chemistry, Finnish Cardiovascular Research Center-Tampere, Faculty of Medicine and Technology, Tampere University, Tampere, Finland.

Patrik K E Magnusson (PKE)

Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden.

Albertine J Oldehinkel (AJ)

University of Groningen, University Medical Center Groningen, Department of Psychiatry, Groningen, the Netherlands.

Brenda W J H Penninx (BWJH)

Department of Psychiatry, Amsterdam Neuroscience and Amsterdam Public Health Research Institute, Amsterdam University Medical Center, Vrije Universiteit, Amsterdam, the Netherlands.

Alexandre C Pereira (AC)

Laboratory of Genetics and Molecular Cardiology, Heart Institute (InCor), University of São Paulo Medical School, São Paulo, Brazil.

Rainer Rauramaa (R)

Foundation for Research in Health, Exercise, and Nutrition, Kuopio Research Institute of Exercise Medicine, Kuopio, Finland.

Susan Redline (S)

Division of Sleep and Circadian Disorders, Brigham and Women's Hospital, Boston, MA, USA.

Nilesh J Samani (NJ)

Department of Cardiovascular Sciences, University of Leicester, Leicester, UK.
NIHR Leicester Biomedical Research Centre, Glenfield Hospital, Leicester, UK.

James Scott (J)

National Heart and Lung Institute, Imperial College, London, UK.

Xiao-Ou Shu (XO)

Division of Epidemiology, Department of Medicine, Vanderbilt University School of Medicine, Nashville, TN, USA.

Pim van der Harst (P)

University of Groningen, University Medical Center Groningen, Department of Cardiology, Groningen, the Netherlands.
University of Groningen, University Medical Center Groningen, Department of Genetics, Groningen, the Netherlands.

Lynne E Wagenknecht (LE)

Public Health Sciences, Wake Forest School of Medicine, Winston-Salem, NC, USA.

Jun-Sing Wang (JS)

Endocrinology and Metabolism, Internal Medicine, Taichung Veterans General Hospital, Taichung, Taiwan.
School of Medicine, National Yang-Ming University, Taipei, Taiwan.

Ya Xing Wang (YX)

Beijing Institute of Ophthalmology, Beijing Ophthalmology and Visual Science Key Laboratory, Beijing Tongren Eye Center, Capital Medical University, Beijing, China.

Nicholas J Wareham (NJ)

Medical Research Council Epidemiology Unit, University of Cambridge, Cambridge, UK.

Hugh Watkins (H)

Division of Cardiovascular Medicine, Radcliffe Department of Medicine, University of Oxford, Oxford, UK.
Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.

David R Weir (DR)

Survey Research Center, Institute for Social Research, University of Michigan, Ann Arbor, MI, USA.

Ananda R Wickremasinghe (AR)

Department of Public Health, Faculty of Medicine, University of Kelaniya, Ragama, Sri Lanka.

Tangchun Wu (T)

Department of Occupational and Environmental Health and State Key Laboratory of Environmental Health for Incubating, Tongji Medical College, Huazhong University of Science and Technology, Wuhan, China.

Eleftheria Zeggini (E)

Human Genetics, Wellcome Trust Sanger Institute, Hinxton, UK.
Institute of Translational Genomics, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.

Wei Zheng (W)

Division of Epidemiology, Department of Medicine, Vanderbilt University School of Medicine, Nashville, TN, USA.

Claude Bouchard (C)

Human Genomics Laboratory, Pennington Biomedical Research Center, Baton Rouge, LA, USA.

Michele K Evans (MK)

Health Disparities Research Section, Laboratory of Epidemiology and Population Sciences, National Institute on Aging, US National Institutes of Health, Baltimore, MD, USA.

Vilmundur Gudnason (V)

Icelandic Heart Association, Kopavogur, Iceland.
Faculty of Medicine, University of Iceland, Reykjavik, Iceland.

Sharon L R Kardia (SLR)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.

Yongmei Liu (Y)

Public Health Sciences, Epidemiology and Prevention, Wake Forest University Health Sciences, Winston-Salem, NC, USA.

Bruce M Psaty (BM)

Cardiovascular Health Research Unit, Department of Medicine, University of Washington, Seattle, WA, USA.
Department of Epidemiology, University of Washington, Seattle, WA, USA.
Department of Health Services, University of Washington, Seattle, WA, USA.
Kaiser Permanente Washington Health Research Institute, Seattle, WA, USA.

Paul M Ridker (PM)

Division of Preventive Medicine, Brigham and Women's Hospital, Boston, MA, USA.
Harvard Medical School, Boston, MA, USA.

Rob M van Dam (RM)

Saw Swee Hock School of Public Health, National University of Singapore and National University Health System, Singapore, Singapore.
Department of Medicine, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.

Dennis O Mook-Kanamori (DO)

Clinical Epidemiology, Leiden University Medical Center, Leiden, the Netherlands.
Public Health and Primary Care, Leiden University Medical Center, Leiden, the Netherlands.

Myriam Fornage (M)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, University of Texas Health Science Center at Houston, Houston, TX, USA.
Brown Foundation Institute of Molecular Medicine, University of Texas Health Science Center at Houston, Houston, TX, USA.

Michael A Province (MA)

Division of Statistical Genomics, Department of Genetics, Washington University School of Medicine, St. Louis, MO, USA.

Tanika N Kelly (TN)

Epidemiology, Tulane University School of Public Health and Tropical Medicine, New Orleans, LA, USA.

Ervin R Fox (ER)

Cardiology, Medicine, University of Mississippi Medical Center, Jackson, MS, USA.

Caroline Hayward (C)

Medical Research Council Human Genetics Unit, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, UK.

Cornelia M van Duijn (CM)

Department of Epidemiology, Erasmus University Medical Center, Rotterdam, the Netherlands.
Nuffield Department of Population Health, University of Oxford, Oxford, UK.

E Shyong Tai (ES)

Saw Swee Hock School of Public Health, National University of Singapore and National University Health System, Singapore, Singapore.
Duke-NUS Medical School, Singapore, Singapore.
Department of Medicine, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.

Tien Yin Wong (TY)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, Singapore.
Department of Ophthalmology, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, Singapore.
Ophthalmology and Visual Sciences Academic Clinical Program (Eye ACP), Duke-NUS Medical School, Singapore, Singapore.

Ruth J F Loos (RJF)

Charles Bronfman Institute for Personalized Medicine, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
Mindich Child Health Development Institute, Icahn School of Medicine at Mount Sinai, New York, NY, USA.

Nora Franceschini (N)

Department of Epidemiology, Gillings School of Global Public Health, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.

Jerome I Rotter (JI)

Institute for Translational Genomics and Population Sciences, Department of Pediatrics, Los Angeles Biomedical Research Institute at Harbor-UCLA Medical Center, Torrance, CA, USA.

Xiaofeng Zhu (X)

Department of Population Quantitative and Health Sciences, Case Western Reserve University, Cleveland, OH, USA.

Laura J Bierut (LJ)

Psychiatry, Washington University School of Medicine, St. Louis, MO, USA.

W James Gauderman (WJ)

Biostatistics, Department of Preventive Medicine, University of Southern California, Los Angeles, CA, USA.

Kenneth Rice (K)

Department of Biostatistics, University of Washington, Seattle, WA, USA.

Patricia B Munroe (PB)

Clinical Pharmacology, William Harvey Research Institute, Barts and The London School of Medicine and Dentistry, Queen Mary University of London, London, UK.
NIHR Barts Cardiovascular Biomedical Research Centre, Queen Mary University of London, London, UK.

Alanna C Morrison (AC)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, University of Texas Health Science Center at Houston, Houston, TX, USA.

Dabeeru C Rao (DC)

Division of Biostatistics, Washington University School of Medicine, St. Louis, MO, USA.

Charles N Rotimi (CN)

Center for Research on Genomics and Global Health, National Human Genome Research Institute, US National Institutes of Health, Bethesda, MD, USA. rotimic@mail.nih.gov.

L Adrienne Cupples (LA)

Department of Biostatistics, Boston University School of Public Health, Boston, MA, USA. adrienne@bu.edu.
Framingham Heart Study, National Heart, Lung, and Blood Institute, US National Institutes of Health, Bethesda, MD, USA. adrienne@bu.edu.

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