Differential localization of histone variant TH2B during the first round compared with subsequent rounds of spermatogenesis.

Hist1h2aa Hist1h2ba ID4-eGFP TH2A TH2B WIN 18446 ZBTB16 chromatin germ cells neonatal spermatocyte spermatogenesis spermatogonia testis

Journal

Developmental dynamics : an official publication of the American Association of Anatomists
ISSN: 1097-0177
Titre abrégé: Dev Dyn
Pays: United States
ID NLM: 9201927

Informations de publication

Date de publication:
06 2019
Historique:
received: 11 09 2018
revised: 20 03 2019
accepted: 29 03 2019
pubmed: 3 4 2019
medline: 20 2 2020
entrez: 3 4 2019
Statut: ppublish

Résumé

Male germ cells are unique because they express a substantial number of variants of the general DNA binding proteins, known as histones, yet the biological significance of these variants is still unknown. In the present study, we aimed to address the expression pattern of the testis-specific histone H2B variant (TH2B) and the testis-specific histone H2A variant (TH2A) within the neonatal mouse testis. We demonstrate that TH2B and TH2A are present in a testis-enriched for undifferentiated spermatogonia. Co-localization studies with an undifferentiated marker, ZBTB16, revealed that TH2B and ZBTB16 co-localize in the neonatal testis. Upon the appearance of the primary spermatocytes, TH2B no longer co-localized with the ZBTB16 positive spermatogonia but were instead detected within the differentiating spermatogonia. This pattern of expression where TH2B and ZBTB16 no longer co-localize was maintained in the adult testis. These findings are in contrast to previous studies, which demonstrated that TH2B and TH2A were found only in adult spermatocytes. Our data are in support of a switch in the expression of these variants following the first round of spermatogonial differentiation. These studies reinforce current understandings that spermatogonia within the neonatal mouse testis are inherently different from those residing within the adult testis.

Sections du résumé

BACKGROUND
Male germ cells are unique because they express a substantial number of variants of the general DNA binding proteins, known as histones, yet the biological significance of these variants is still unknown. In the present study, we aimed to address the expression pattern of the testis-specific histone H2B variant (TH2B) and the testis-specific histone H2A variant (TH2A) within the neonatal mouse testis.
RESULTS
We demonstrate that TH2B and TH2A are present in a testis-enriched for undifferentiated spermatogonia. Co-localization studies with an undifferentiated marker, ZBTB16, revealed that TH2B and ZBTB16 co-localize in the neonatal testis. Upon the appearance of the primary spermatocytes, TH2B no longer co-localized with the ZBTB16 positive spermatogonia but were instead detected within the differentiating spermatogonia. This pattern of expression where TH2B and ZBTB16 no longer co-localize was maintained in the adult testis.
CONCLUSION
These findings are in contrast to previous studies, which demonstrated that TH2B and TH2A were found only in adult spermatocytes. Our data are in support of a switch in the expression of these variants following the first round of spermatogonial differentiation. These studies reinforce current understandings that spermatogonia within the neonatal mouse testis are inherently different from those residing within the adult testis.

Identifiants

pubmed: 30939211
doi: 10.1002/dvdy.33
pmc: PMC6545161
mid: NIHMS1022072
doi:

Substances chimiques

Histones 0
histone H2B type 1-A 0

Types de publication

Journal Article Research Support, N.I.H., Extramural

Langues

eng

Sous-ensembles de citation

IM

Pagination

488-500

Subventions

Organisme : NICHD NIH HHS
ID : R01 HD010808
Pays : United States
Organisme : NICHD NIH HHS
ID : R37 HD010808
Pays : United States

Informations de copyright

© 2019 The Authors. Developmental Dynamics published by Wiley Periodicals, Inc. on behalf of American Association of Anatomists.

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Auteurs

My-Thanh Beedle (MT)

School of Molecular Biosciences and Center for Reproductive Biology, Washington State University, Pullman, Washington.

Traci Topping (T)

School of Molecular Biosciences and Center for Reproductive Biology, Washington State University, Pullman, Washington.

Cathryn Hogarth (C)

School of Molecular Biosciences and Center for Reproductive Biology, Washington State University, Pullman, Washington.

Michael Griswold (M)

School of Molecular Biosciences and Center for Reproductive Biology, Washington State University, Pullman, Washington.

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Classifications MeSH