Long non-coding RNA uc.291 controls epithelial differentiation by interfering with the ACTL6A/BAF complex.


Journal

EMBO reports
ISSN: 1469-3178
Titre abrégé: EMBO Rep
Pays: England
ID NLM: 100963049

Informations de publication

Date de publication:
04 03 2020
Historique:
received: 13 07 2018
revised: 23 12 2019
accepted: 08 01 2020
pubmed: 6 2 2020
medline: 28 4 2021
entrez: 5 2 2020
Statut: ppublish

Résumé

The mechanisms that regulate the switch between epidermal progenitor state and differentiation are not fully understood. Recent findings indicate that the chromatin remodelling BAF complex (Brg1-associated factor complex or SWI/SNF complex) and the transcription factor p63 mutually recruit one another to open chromatin during epidermal differentiation. Here, we identify a long non-coding transcript that includes an ultraconserved element, uc.291, which physically interacts with ACTL6A and modulates chromatin remodelling to allow differentiation. Loss of uc.291 expression, both in primary keratinocytes and in three-dimensional skin equivalents, inhibits differentiation as indicated by epidermal differentiation complex genes down-regulation. ChIP experiments reveal that upon uc.291 depletion, ACTL6A is bound to the differentiation gene promoters and inhibits BAF complex targeting to induce terminal differentiation genes. In the presence of uc.291, the ACTL6A inhibitory effect is released, allowing chromatin changes to promote the expression of differentiation genes. Thus, uc.291 interacts with ACTL6A to modulate chromatin remodelling activity, allowing the transcription of late differentiation genes.

Identifiants

pubmed: 32017402
doi: 10.15252/embr.201846734
pmc: PMC7054673
doi:

Substances chimiques

ACTL6A protein, human 0
Actins 0
BANF1 protein, human 0
Chromatin 0
Chromosomal Proteins, Non-Histone 0
DNA-Binding Proteins 0
RNA, Long Noncoding 0
Transcription Factors 0

Banques de données

GEO
['GSE103890']

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

e46734

Subventions

Organisme : Fundació la Marató de TV3 (TV3 Marathon Foundation)
ID : PI043296
Pays : International
Organisme : Ministerio de Economía y Competitividad (MEC)
ID : BFU2014-55054-P
Pays : International
Organisme : Medical Research Council
ID : MC_U132670600
Pays : United Kingdom
Organisme : Associazione Italiana per la Ricerca sul Cancro (AIRC)
ID : 22206
Pays : International
Organisme : EC | H2020 | H2020 Priority Excellent Science | H2020 European Research Council (ERC)
ID : RIBOMYLOME_309545
Pays : International
Organisme : Medical Research Council
ID : MC_UU_00025/2
Pays : United Kingdom
Organisme : Ministero della Salute (Ministry of Health, Italy)
ID : IDI-IRCCS-RC3.2
Pays : International

Informations de copyright

© 2020 The Authors. Published under the terms of the CC BY NC ND 4.0 license.

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Auteurs

Emanuele Panatta (E)

Department of Experimental Medicine, University of Rome "Tor Vergata", Rome, Italy.
MRC Toxicology Unit, University of Cambridge, Cambridge, UK.

Anna Maria Lena (AM)

Department of Experimental Medicine, University of Rome "Tor Vergata", Rome, Italy.

Mara Mancini (M)

IDI-IRCCS, Rome, Italy.

Artem Smirnov (A)

Department of Experimental Medicine, University of Rome "Tor Vergata", Rome, Italy.

Alberto Marini (A)

Department of Experimental Medicine, University of Rome "Tor Vergata", Rome, Italy.
MRC Toxicology Unit, University of Cambridge, Cambridge, UK.

Riccardo Delli Ponti (R)

Centre for Genomic Regulation (CRG), The Barcelona Institute for Science and Technology, Barcelona, Spain.
Universitat Pompeu Fabra (UPF), Barcelona, Spain.

Teresa Botta-Orfila (T)

Centre for Genomic Regulation (CRG), The Barcelona Institute for Science and Technology, Barcelona, Spain.
Universitat Pompeu Fabra (UPF), Barcelona, Spain.

Gian Gaetano Tartaglia (GG)

Centre for Genomic Regulation (CRG), The Barcelona Institute for Science and Technology, Barcelona, Spain.
Universitat Pompeu Fabra (UPF), Barcelona, Spain.
Institucio Catalana de Recerca i Estudis Avançats (ICREA), Barcelona, Spain.

Alessandro Mauriello (A)

Department of Experimental Medicine, University of Rome "Tor Vergata", Rome, Italy.

Xinna Zhang (X)

The Center for RNA Interference and Non-coding RNAs, The University of Texas MD Anderson Cancer Center, Houston, TX, USA.

George A Calin (GA)

The Center for RNA Interference and Non-coding RNAs, The University of Texas MD Anderson Cancer Center, Houston, TX, USA.

Gerry Melino (G)

Department of Experimental Medicine, University of Rome "Tor Vergata", Rome, Italy.
MRC Toxicology Unit, University of Cambridge, Cambridge, UK.

Eleonora Candi (E)

Department of Experimental Medicine, University of Rome "Tor Vergata", Rome, Italy.
IDI-IRCCS, Rome, Italy.

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Classifications MeSH