Targeted Killing of Pseudomonas aeruginosa by Pyocin G Occurs via the Hemin Transporter Hur.


Journal

Journal of molecular biology
ISSN: 1089-8638
Titre abrégé: J Mol Biol
Pays: Netherlands
ID NLM: 2985088R

Informations de publication

Date de publication:
12 06 2020
Historique:
received: 06 03 2020
revised: 21 04 2020
accepted: 21 04 2020
pubmed: 28 4 2020
medline: 29 12 2020
entrez: 28 4 2020
Statut: ppublish

Résumé

Pseudomonas aeruginosa is a priority pathogen for the development of new antibiotics, particularly because multi-drug-resistant strains of this bacterium cause serious nosocomial infections and are the leading cause of death in cystic fibrosis patients. Pyocins, bacteriocins of P. aeruginosa, are potent and diverse protein antibiotics that are deployed during bacterial competition. Pyocins are produced by more than 90% of P. aeruginosa strains and may have utility as last resort antibiotics against this bacterium. In this study, we explore the antimicrobial activity of a newly discovered pyocin called pyocin G (PyoG). We demonstrate that PyoG has broad killing activity against a collection of clinical P. aeruginosa isolates and is active in a Galleria mellonella infection model. We go on to identify cell envelope proteins that are necessary for the import of PyoG and its killing activity. PyoG recognizes bacterial cells by binding to Hur, an outer-membrane TonB-dependent transporter. Both pyocin and Hur interact with TonB1, which in complex with ExbB-ExbD links the proton motive force generated across the inner membrane with energy-dependent pyocin translocation across the outer membrane. Inner-membrane translocation of PyoG is dependent on the conserved inner-membrane AAA+ ATPase/protease, FtsH. We also report a functional exploration of the PyoG receptor. We demonstrate that Hur can bind to hemin in vitro and that this interaction is blocked by PyoG, confirming the role of Hur in hemin acquisition.

Identifiants

pubmed: 32339530
pii: S0022-2836(20)30308-9
doi: 10.1016/j.jmb.2020.04.020
pmc: PMC7322526
pii:
doi:

Substances chimiques

Anti-Bacterial Agents 0
Bacterial Proteins 0
Bacteriocins 0
Membrane Proteins 0
Pyocins 0
tonB protein, Bacteria 0
Hemin 743LRP9S7N
ATPases Associated with Diverse Cellular Activities EC 3.6.4.-

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

3869-3880

Subventions

Organisme : Wellcome Trust
ID : 201505/Z/16/Z
Pays : United Kingdom

Informations de copyright

Copyright © 2020 The Authors. Published by Elsevier Ltd.. All rights reserved.

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Auteurs

Iva Atanaskovic (I)

Department of Biochemistry, University of Oxford, South Parks Road, Oxford OX1 3QU, UK.

Khedidja Mosbahi (K)

Institute of Infection, Immunity, and Inflammation, College of Medical, Veterinary, and Life Sciences, University of Glasgow, G12 8QQ Glasgow, UK.

Connor Sharp (C)

Department of Zoology, University of Oxford, 11a Mansfield Rd, Oxford OX1 3SZ, UK.

Nicholas G Housden (NG)

Department of Biochemistry, University of Oxford, South Parks Road, Oxford OX1 3QU, UK.

Renata Kaminska (R)

Department of Biochemistry, University of Oxford, South Parks Road, Oxford OX1 3QU, UK.

Daniel Walker (D)

Institute of Infection, Immunity, and Inflammation, College of Medical, Veterinary, and Life Sciences, University of Glasgow, G12 8QQ Glasgow, UK.

Colin Kleanthous (C)

Department of Biochemistry, University of Oxford, South Parks Road, Oxford OX1 3QU, UK. Electronic address: colin.kleanthous@bioch.ox.ac.uk.

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Classifications MeSH