Fibrillar form of α-synuclein-specific scFv antibody inhibits α-synuclein seeds induced aggregation and toxicity.


Journal

Scientific reports
ISSN: 2045-2322
Titre abrégé: Sci Rep
Pays: England
ID NLM: 101563288

Informations de publication

Date de publication:
18 05 2020
Historique:
received: 19 07 2019
accepted: 24 04 2020
entrez: 20 5 2020
pubmed: 20 5 2020
medline: 15 12 2020
Statut: epublish

Résumé

Synucleinopathies including Parkinson's disease (PD), dementia with Lewy bodies (DLB), and multiple system atrophy (MSA) are characterized by pathological accumulation of α-synuclein (α-syn). Amongst the various approaches attempting to tackle the pathological features of synucleinopathies, antibody-based immunotherapy holds much promise. However, the large size of antibodies and corresponding difficulty in crossing the blood-brain barrier has limited development in this area. To overcome this issue, we engineered single-chain variable fragments (scFvs) against fibrillar α-syn, a putative disease-relevant form of α-syn. The purified scFvs showed specific activity towards α-syn fibrils and oligomers in comparison to monomers and recognized intracellular inclusions in human post-mortem brain tissue of Lewy body disease cases, but not aged controls. In vitro studies indicated scFvs inhibit the seeding of α-syn aggregation in a time-dependent manner, decreased α-syn seed-induced toxicity in a cell model of PD, and reduced the production of insoluble α-syn phosphorylated at Ser-129 (pS129-α-syn). These results suggest that our α-syn fibril-specific scFvs recognize α-syn pathology and can inhibit the aggregation of α-syn in vitro and prevent seeding-dependent toxicity. Therefore, the scFvs described here have considerable potential to be utilized towards immunotherapy in synucleinopathies and may also have applications in ante-mortem imaging modalities.

Identifiants

pubmed: 32424162
doi: 10.1038/s41598-020-65035-8
pii: 10.1038/s41598-020-65035-8
pmc: PMC7235225
doi:

Substances chimiques

Protein Aggregates 0
Single-Chain Antibodies 0
alpha-Synuclein 0

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

8137

Subventions

Organisme : Medical Research Council
ID : G0400074
Pays : United Kingdom
Organisme : Medical Research Council
ID : G0502157
Pays : United Kingdom
Organisme : Medical Research Council
ID : G0900652
Pays : United Kingdom
Organisme : Medical Research Council
ID : G1100540
Pays : United Kingdom

Références

Spillantini, M. G. & Goedert, M. The alpha-synucleinopathies: Parkinson’s disease, dementia with Lewy bodies, and multiple system atrophy. Ann. N. Y. Acad. Sci. 920, 16–27 (2000).
doi: 10.1111/j.1749-6632.2000.tb06900.x
McKeith, I. G. et al. Diagnosis and management of dementia with Lewy bodies: Fourth consensus report of the DLB Consortium. Neurology 89, 88–100, https://doi.org/10.1212/WNL.0000000000004058 (2017).
doi: 10.1212/WNL.0000000000004058 pubmed: 28592453 pmcid: 5496518
Spillantini, M. G. et al. Alpha-synuclein in Lewy bodies. Nature 388, 839–840, https://doi.org/10.1038/42166 (1997).
doi: 10.1038/42166 pubmed: 9278044 pmcid: 9278044
Papp, M. I., Kahn, J. E. & Lantos, P. L. Glial cytoplasmic inclusions in the CNS of patients with multiple system atrophy (striatonigral degeneration, olivopontocerebellar atrophy and Shy-Drager syndrome). J. Neurol. Sci. 94, 79–100 (1989).
doi: 10.1016/0022-510X(89)90219-0
Cykowski, M. D. et al. Expanding the spectrum of neuronal pathology in multiple system atrophy. Brain: a J. Neurol. 138, 2293–2309, https://doi.org/10.1093/brain/awv114 (2015).
doi: 10.1093/brain/awv114
Masliah, E. et al. Dopaminergic loss and inclusion body formation in alpha-synuclein mice: implications for neurodegenerative disorders. Science 287, 1265–1269 (2000).
doi: 10.1126/science.287.5456.1265
Feany, M. B. & Bender, W. W. A Drosophila model of Parkinson’s disease. Nature 404, 394–398, https://doi.org/10.1038/35006074 (2000).
doi: 10.1038/35006074 pubmed: 10746727
Chen, L. & Feany, M. B. Alpha-synuclein phosphorylation controls neurotoxicity and inclusion formation in a Drosophila model of Parkinson disease. Nat. Neurosci. 8, 657–663, https://doi.org/10.1038/nn1443 (2005).
doi: 10.1038/nn1443 pubmed: 15834418
Singleton, A. B. et al. alpha-Synuclein locus triplication causes Parkinson’s disease. Science 302, 841, https://doi.org/10.1126/science.1090278 (2003).
doi: 10.1126/science.1090278 pubmed: 14593171 pmcid: 14593171
Zarranz, J. J. et al. The new mutation, E46K, of alpha-synuclein causes Parkinson and Lewy body dementia. Ann. Neurol. 55, 164–173, https://doi.org/10.1002/ana.10795 (2004).
doi: 10.1002/ana.10795 pubmed: 14755719
Polymeropoulos, M. H. et al. Mutation in the alpha-synuclein gene identified in families with Parkinson’s disease. Science 276, 2045–2047 (1997).
doi: 10.1126/science.276.5321.2045
Galvin, J. E., Uryu, K., Lee, V. M. & Trojanowski, J. Q. Axon pathology in Parkinson’s disease and Lewy body dementia hippocampus contains alpha-, beta-, and gamma-synuclein. Proc. Natl Acad. Sci. U S Am. 96, 13450–13455 (1999).
doi: 10.1073/pnas.96.23.13450
Mittal, S. et al. beta2-Adrenoreceptor is a regulator of the alpha-synuclein gene driving risk of Parkinson’s disease. Science 357, 891–898, https://doi.org/10.1126/science.aaf3934 (2017).
doi: 10.1126/science.aaf3934 pubmed: 5761666 pmcid: 5761666
Iwai, A. et al. The precursor protein of non-A beta component of Alzheimer’s disease amyloid is a presynaptic protein of the central nervous system. Neuron 14, 467–475 (1995).
doi: 10.1016/0896-6273(95)90302-X
Stefanis, L. alpha-Synuclein in Parkinson’s disease. Cold Spring Harb. Perspect. Med. 2, a009399, https://doi.org/10.1101/cshperspect.a009399 (2012).
doi: 10.1101/cshperspect.a009399 pubmed: 22355802 pmcid: 3281589
Landrigan, P. J. et al. Early environmental origins of neurodegenerative disease in later life. Environ. health Perspect. 113, 1230–1233, https://doi.org/10.1289/ehp.7571 (2005).
doi: 10.1289/ehp.7571 pubmed: 16140633 pmcid: 1280407
Betarbet, R. et al. Chronic systemic pesticide exposure reproduces features of Parkinson’s disease. Nat. Neurosci. 3, 1301–1306, https://doi.org/10.1038/81834 (2000).
doi: 10.1038/81834 pubmed: 11100151
Bartels, T., Choi, J. G. & Selkoe, D. J. alpha-Synuclein occurs physiologically as a helically folded tetramer that resists aggregation. Nature 477, 107–110, https://doi.org/10.1038/nature10324 (2011).
doi: 10.1038/nature10324 pubmed: 21841800 pmcid: 3166366
Ghosh, D., Mehra, S., Sahay, S., Singh, P. K. & Maji, S. K. alpha-synuclein aggregation and its modulation. Int. J. Biol. Macromol. 100, 37–54, https://doi.org/10.1016/j.ijbiomac.2016.10.021 (2017).
doi: 10.1016/j.ijbiomac.2016.10.021 pubmed: 27737778
El-Agnaf, O. M., Walsh, D. M. & Allsop, D. Soluble oligomers for the diagnosis of neurodegenerative diseases. Lancet. Neurol. 2, 461–462 (2003).
doi: 10.1016/S1474-4422(03)00481-2
Atik, A., Stewart, T. & Zhang, J. Alpha-synuclein as a biomarker for Parkinson’s disease. Brain Pathol. 26, 410–418 (2016).
doi: 10.1111/bpa.12370
el-Agnaf, O. M. & Irvine, G. B. Aggregation and neurotoxicity of alpha-synuclein and related peptides. Biochemical Society transactions 30, 559–565, 10.1042 (2002).
Chia, K. Y., Ng, K. Y., Koh, R. Y. & Chye, S. M. Single-chain Fv Antibodies for Targeting. Neurodegenerative Diseases. CNS neurological Disord. drug. targets 17, 671–679, https://doi.org/10.2174/1871527317666180315161626 (2018).
doi: 10.2174/1871527317666180315161626
Nannenga, B. L., Zameer, A. & Sierks, M. R. Anti-oligomeric single chain variable domain antibody differentially affects huntingtin and alpha-synuclein aggregates. FEBS Lett. 582, 517–522, https://doi.org/10.1016/j.febslet.2008.01.014 (2008).
doi: 10.1016/j.febslet.2008.01.014 pubmed: 18230361
Emadi, S., Kasturirangan, S., Wang, M. S., Schulz, P. & Sierks, M. R. Detecting morphologically distinct oligomeric forms of alpha-synuclein. J. Biol. Chem. 284, 11048–11058, https://doi.org/10.1074/jbc.M806559200 (2009).
doi: 10.1074/jbc.M806559200 pubmed: 19141614 pmcid: 2670110
Maguire-Zeiss, K. A. et al. Identification of human alpha-synuclein specific single chain antibodies. Biochemical biophysical Res. Commun. 349, 1198–1205, https://doi.org/10.1016/j.bbrc.2006.08.127 (2006).
doi: 10.1016/j.bbrc.2006.08.127
Yuan, B. & Sierks, M. R. Intracellular targeting and clearance of oligomeric alpha-synuclein alleviates toxicity in mammalian cells. Neurosci. Lett. 459, 16–18, https://doi.org/10.1016/j.neulet.2009.04.046 (2009).
doi: 10.1016/j.neulet.2009.04.046 pubmed: 19394405
Huang, L., Su, X. & Federoff, H. J. Single-chain fragment variable passive immunotherapies for neurodegenerative diseases. Int. J. Mol. Sci. 14, 19109–19127, https://doi.org/10.3390/ijms140919109 (2013).
doi: 10.3390/ijms140919109 pubmed: 24048248 pmcid: 3794823
Spencer, B. et al. alpha-synuclein conformational antibodies fused to penetratin are effective in models of Lewy body disease. Ann. Clin. Transl. Neurol. 3, 588–606, https://doi.org/10.1002/acn3.321 (2016).
doi: 10.1002/acn3.321 pubmed: 27606342 pmcid: 4999592
Vaikath, N. N. et al. Generation and characterization of novel conformation-specific monoclonal antibodies for alpha-synuclein pathology. Neurobiol. Dis. 79, 81–99, https://doi.org/10.1016/j.nbd.2015.04.009 (2015).
doi: 10.1016/j.nbd.2015.04.009 pubmed: 25937088
El-Agnaf, O. et al. Differential effects of immunotherapy with antibodies targeting alpha-synuclein oligomers and fibrils in a transgenic model of synucleinopathy. Neurobiol. Dis. 104, 85–96, https://doi.org/10.1016/j.nbd.2017.05.002 (2017).
doi: 10.1016/j.nbd.2017.05.002 pubmed: 28476636 pmcid: 5954414
Singh, S. M. & Panda, A. K. Solubilization and refolding of bacterial inclusion body proteins. J. Biosci. Bioeng. 99, 303–310, https://doi.org/10.1263/jbb.99.303 (2005).
doi: 10.1263/jbb.99.303 pubmed: 16233795
Jarrett, J. T. & Lansbury, P. T. Jr. Amyloid fibril formation requires a chemically discriminating nucleation event: studies of an amyloidogenic sequence from the bacterial protein OsmB. Biochemistry 31, 12345–12352 (1992).
doi: 10.1021/bi00164a008
Harper, J. D., Wong, S. S., Lieber, C. M. & Lansbury, P. T. Jr. Assembly of A beta amyloid protofibrils: an in vitro model for a possible early event in Alzheimer’s disease. Biochemistry 38, 8972–8980, https://doi.org/10.1021/bi9904149 (1999).
doi: 10.1021/bi9904149
Jarrett, J. T. & Lansbury, P. T. Jr. Seeding “one-dimensional crystallization” of amyloid: a pathogenic mechanism in Alzheimer’s disease and scrapie? Cell 73, 1055–1058 (1993).
doi: 10.1016/0092-8674(93)90635-4
Harper, J. D. & Lansbury, P. T. Jr. Models of amyloid seeding in Alzheimer’s disease and scrapie: mechanistic truths and physiological consequences of the time-dependent solubility of amyloid proteins. Annu. Rev. Biochem. 66, 385–407, https://doi.org/10.1146/annurev.biochem.66.1.385 (1997).
doi: 10.1146/annurev.biochem.66.1.385
Volpicelli-Daley, L. A., Luk, K. C. & Lee, V. M. Addition of exogenous alpha-synuclein preformed fibrils to primary neuronal cultures to seed recruitment of endogenous alpha-synuclein to Lewy body and Lewy neurite-like aggregates. Nat. Protoc. 9, 2135–2146, https://doi.org/10.1038/nprot.2014.143 (2014).
doi: 10.1038/nprot.2014.143 pubmed: 25122523 pmcid: 4372899
Luk, K. C. et al. Exogenous alpha-synuclein fibrils seed the formation of Lewy body-like intracellular inclusions in cultured cells. Proc. Natl Acad. Sci. U S Am. 106, 20051–20056, https://doi.org/10.1073/pnas.0908005106 (2009).
doi: 10.1073/pnas.0908005106
Ardah, M. T. et al. Ginsenoside Rb1 inhibits fibrillation and toxicity of alpha-synuclein and disaggregates preformed fibrils. Neurobiol. Dis. 74, 89–101, https://doi.org/10.1016/j.nbd.2014.11.007 (2015).
doi: 10.1016/j.nbd.2014.11.007 pubmed: 25449909
Ardah, M. T. et al. Structure activity relationship of phenolic acid inhibitors of alpha-synuclein fibril formation and toxicity. Front. aging Neurosci. 6, 197, https://doi.org/10.3389/fnagi.2014.00197 (2014).
doi: 10.3389/fnagi.2014.00197 pubmed: 25140150 pmcid: 4122169
Spencer, B. et al. Selective targeting of 3 repeat Tau with brain penetrating single chain antibodies for the treatment of neurodegenerative disorders. Acta neuropathologica 136, 69–87, https://doi.org/10.1007/s00401-018-1869-0 (2018).
doi: 10.1007/s00401-018-1869-0 pubmed: 29934874 pmcid: 6112111
Skrlj, N. et al. Recombinant single-chain antibody with the Trojan peptide penetratin positioned in the linker region enables cargo transfer across the blood-brain barrier. Appl. Biochem. Biotechnol. 169, 159–169, https://doi.org/10.1007/s12010-012-9962-7 (2013).
doi: 10.1007/s12010-012-9962-7 pubmed: 23160949
Wong, Y. C. & Krainc, D. alpha-synuclein toxicity in neurodegeneration: mechanism and therapeutic strategies. Nat. Med. 23, 1–13, https://doi.org/10.1038/nm.4269 (2017).
doi: 10.1038/nm.4269 pubmed: 28170377
Winner, B. et al. In vivo demonstration that alpha-synuclein oligomers are toxic. Proc. Natl Acad. Sci. U S Am. 108, 4194–4199, https://doi.org/10.1073/pnas.1100976108 (2011).
doi: 10.1073/pnas.1100976108
Mahul-Mellier, A. L. et al. Fibril growth and seeding capacity play key roles in alpha-synuclein-mediated apoptotic cell death. Cell death Differ. 22, 2107–2122, https://doi.org/10.1038/cdd.2015.79 (2015).
doi: 10.1038/cdd.2015.79 pubmed: 26138444 pmcid: 4816119
El-Agnaf, O. M. et al. Aggregates from mutant and wild-type alpha-synuclein proteins and NAC peptide induce apoptotic cell death in human neuroblastoma cells by formation of beta-sheet and amyloid-like filaments. FEBS Lett. 440, 71–75 (1998).
doi: 10.1016/S0014-5793(98)01418-5
Grassi, D. et al. Identification of a highly neurotoxic alpha-synuclein species inducing mitochondrial damage and mitophagy in Parkinson’s disease. Proc. Natl Acad. Sci. U S Am. 115, E2634–E2643, https://doi.org/10.1073/pnas.1713849115 (2018).
doi: 10.1073/pnas.1713849115
Anderson, J. P. et al. Phosphorylation of Ser-129 is the dominant pathological modification of alpha-synuclein in familial and sporadic Lewy body disease. J. Biol. Chem. 281, 29739–29752, https://doi.org/10.1074/jbc.M600933200 (2006).
doi: 10.1074/jbc.M600933200
Iyer, A. et al. The Impact of N-terminal Acetylation of alpha-Synuclein on Phospholipid Membrane Binding and Fibril Structure. J. Biol. Chem. 291, 21110–21122, https://doi.org/10.1074/jbc.M116.726612 (2016).
doi: 10.1074/jbc.M116.726612 pubmed: 27531743 pmcid: 5076520
Paxinou, E. et al. Induction of alpha-synuclein aggregation by intracellular nitrative insult. J. neuroscience: Off. J. Soc. Neurosci. 21, 8053–8061 (2001).
doi: 10.1523/JNEUROSCI.21-20-08053.2001
Hasegawa, M. et al. Phosphorylated alpha-synuclein is ubiquitinated in alpha-synucleinopathy lesions. J. Biol. Chem. 277, 49071–49076, https://doi.org/10.1074/jbc.M208046200 (2002).
doi: 10.1074/jbc.M208046200 pubmed: 12377775
Paleologou, K. E. et al. Phosphorylation at S87 is enhanced in synucleinopathies, inhibits alpha-synuclein oligomerization, and influences synuclein-membrane interactions. J. neuroscience: Off. J. Soc. Neurosci. 30, 3184–3198, https://doi.org/10.1523/JNEUROSCI.5922-09.2010 (2010).
doi: 10.1523/JNEUROSCI.5922-09.2010
Vicente Miranda, H. et al. Glycation potentiates alpha-synuclein-associated neurodegeneration in synucleinopathies. Brain: a J. Neurol. 140, 1399–1419, https://doi.org/10.1093/brain/awx056 (2017).
doi: 10.1093/brain/awx056
Giasson, B. I. et al. Oxidative damage linked to neurodegeneration by selective alpha-synuclein nitration in synucleinopathy lesions. Science 290, 985–989 (2000).
doi: 10.1126/science.290.5493.985
Dorval, V. & Fraser, P. E. Small ubiquitin-like modifier (SUMO) modification of natively unfolded proteins tau and alpha-synuclein. J. Biol. Chem. 281, 9919–9924, https://doi.org/10.1074/jbc.M510127200 (2006).
doi: 10.1074/jbc.M510127200 pubmed: 16464864
Gelb, D. J., Oliver, E. & Gilman, S. Diagnostic criteria for Parkinson disease. Arch. Neurol. 56, 33–39 (1999).
doi: 10.1001/archneur.56.1.33
Ma, M. R., Hu, Z. W., Zhao, Y. F., Chen, Y. X. & Li, Y. M. Phosphorylation induces distinct alpha-synuclein strain formation. Sci. Rep. 6, 37130, https://doi.org/10.1038/srep37130 (2016).
doi: 10.1038/srep37130 pubmed: 27853185 pmcid: 5112567
El-Agnaf, O. M. & Irvine, G. B. Review: formation and properties of amyloid-like fibrils derived from alpha-synuclein and related proteins. J. Struct. Biol. 130, 300–309, https://doi.org/10.1006/jsbi.2000.4262 (2000).
doi: 10.1006/jsbi.2000.4262 pubmed: 10940234
Burre, J., Sharma, M. & Sudhof, T. C. Cell Biology and Pathophysiology of alpha-Synuclein. Cold Spring Harb Perspect Med 8, https://doi.org/10.1101/cshperspect.a024091 (2018).
van Steenoven, I. et al. alpha-Synuclein species as potential cerebrospinal fluid biomarkers for dementia with lewy bodies. Mov. disorders: Off. J. Mov. Disord. Soc. 33, 1724–1733, https://doi.org/10.1002/mds.111 (2018).
doi: 10.1002/mds.111
Zella, S. M. A. et al. Emerging Immunotherapies for Parkinson Disease. Neurology and therapy, https://doi.org/10.1007/s40120-018-0122-z (2018).
Freskgard, P. O. & Urich, E. Antibody therapies in CNS diseases. Neuropharmacology 120, 38–55, https://doi.org/10.1016/j.neuropharm.2016.03.014 (2017).
doi: 10.1016/j.neuropharm.2016.03.014 pubmed: 26972827
Xi, H. et al. Fusion Peptide Improves Stability and Bioactivity of Single Chain Antibody against Rabies Virus. J. microbiology Biotechnol. 27, 718–724, https://doi.org/10.4014/jmb.1611.11062 (2017).
doi: 10.4014/jmb.1611.11062
Zhou, C., Emadi, S., Sierks, M. R. & Messer, A. A human single-chain Fv intrabody blocks aberrant cellular effects of overexpressed alpha-synuclein. Mol. therapy: J. Am. Soc. Gene Ther. 10, 1023–1031, https://doi.org/10.1016/j.ymthe.2004.08.019 (2004).
doi: 10.1016/j.ymthe.2004.08.019
Lynch, S. M., Zhou, C. & Messer, A. An scFv intrabody against the nonamyloid component of alpha-synuclein reduces intracellular aggregation and toxicity. J. Mol. Biol. 377, 136–147, https://doi.org/10.1016/j.jmb.2007.11.096 (2008).
doi: 10.1016/j.jmb.2007.11.096 pubmed: 18237741
Joshi, S. N., Butler, D. C. & Messer, A. Fusion to a highly charged proteasomal retargeting sequence increases soluble cytoplasmic expression and efficacy of diverse anti-synuclein intrabodies. mAbs 4, 686–693, https://doi.org/10.4161/mabs.21696 (2012).
doi: 10.4161/mabs.21696 pubmed: 22929188 pmcid: 3502235
Fang, X. T. et al. High detection sensitivity with antibody-based PET radioligand for amyloid beta in brain. NeuroImage 184, 881–888, https://doi.org/10.1016/j.neuroimage.2018.10.011 (2019).
doi: 10.1016/j.neuroimage.2018.10.011 pubmed: 30300753
Manoutcharian, K., Perez-Garmendia, R. & Gevorkian, G. Recombinant Antibody Fragments for Neurodegenerative Diseases. Curr. Neuropharmacol. 15, 779–788, https://doi.org/10.2174/1570159X01666160930121647 (2017).
doi: 10.2174/1570159X01666160930121647 pubmed: 27697033 pmcid: 5771054
Pieri, L., Madiona, K. & Melki, R. Structural and functional properties of prefibrillar alpha-synuclein oligomers. Sci. Rep. 6, 24526, https://doi.org/10.1038/srep24526 (2016).
doi: 10.1038/srep24526 pubmed: 27075649 pmcid: 4830946

Auteurs

Vijay Gupta (V)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar.

Safa Salim (S)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar.

Issam Hmila (I)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar.

Nishant N Vaikath (NN)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar.

Indulekha P Sudhakaran (IP)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar.

Simona S Ghanem (SS)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar.

Nour K Majbour (NK)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar.

Sara A Abdulla (SA)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar.

Mohamed M Emara (MM)

Basic Medical Sciences Department, College of Medicine, QU Health, Qatar University, Doha, Qatar.

Houari B Abdesselem (HB)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar.

Tamas Lukacsovich (T)

Brain Research Institute, University of Zürich, Zürich, Switzerland.

Daniel Erskine (D)

Translational and Clinical Research Institute, Newcastle University, Newcastle upon Tyne, United Kingdom.

Omar M A El-Agnaf (OMA)

Neurological Disorder Research Center, Qatar Biomedical Research Institute (QBRI), Hamad Bin Khalifa University (HBKU), Qatar Foundation, Doha, Qatar. oelagnaf@qf.org.qa.

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