FFAT motif phosphorylation controls formation and lipid transfer function of inter-organelle contacts.


Journal

The EMBO journal
ISSN: 1460-2075
Titre abrégé: EMBO J
Pays: England
ID NLM: 8208664

Informations de publication

Date de publication:
01 12 2020
Historique:
received: 28 12 2019
revised: 01 09 2020
accepted: 07 09 2020
pubmed: 31 10 2020
medline: 13 4 2021
entrez: 30 10 2020
Statut: ppublish

Résumé

Organelles are physically connected in membrane contact sites. The endoplasmic reticulum possesses three major receptors, VAP-A, VAP-B, and MOSPD2, which interact with proteins at the surface of other organelles to build contacts. VAP-A, VAP-B, and MOSPD2 contain an MSP domain, which binds a motif named FFAT (two phenylalanines in an acidic tract). In this study, we identified a non-conventional FFAT motif where a conserved acidic residue is replaced by a serine/threonine. We show that phosphorylation of this serine/threonine is critical for non-conventional FFAT motifs (named Phospho-FFAT) to be recognized by the MSP domain. Moreover, structural analyses of the MSP domain alone or in complex with conventional and Phospho-FFAT peptides revealed new mechanisms of interaction. Based on these new insights, we produced a novel prediction algorithm, which expands the repertoire of candidate proteins with a Phospho-FFAT that are able to create membrane contact sites. Using a prototypical tethering complex made by STARD3 and VAP, we showed that phosphorylation is instrumental for the formation of ER-endosome contacts, and their sterol transfer function. This study reveals that phosphorylation acts as a general switch for inter-organelle contacts.

Identifiants

pubmed: 33124732
doi: 10.15252/embj.2019104369
pmc: PMC7705450
doi:

Substances chimiques

Lipids 0
MOSPD2 protein, human 0
Membrane Proteins 0
Receptors, Chemokine 0
VAPA protein, human 0
VAPB protein, human 0
Vesicular Transport Proteins 0

Banques de données

PDB
['6TQR', '6TQS', '6TQT', '6TQU']

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

e104369

Subventions

Organisme : Biotechnology and Biological Sciences Research Council
ID : BB/M011801/1
Pays : United Kingdom
Organisme : Agence Nationale de la Recherche (ANR)
ID : ANR-19-CE44-0003
Organisme : Institut National Du Cancer (INCa)
ID : INCA_9269
Organisme : Biotechnology and Biological Sciences Research Council
ID : BB/P003818/1
Pays : United Kingdom
Organisme : French Infrastructure for Integrated Structural Biology (FRISBI)
ID : ANR-10-INSB-05-01
Organisme : Ligue Contre le Cancer

Informations de copyright

© 2020 The Authors. Published under the terms of the CC BY NC ND 4.0 license.

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Auteurs

Thomas Di Mattia (T)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Arthur Martinet (A)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Souade Ikhlef (S)

Institut de Pharmacologie Moléculaire et Cellulaire, Université Côte d'Azur, CNRS, Valbonne, France.

Alastair G McEwen (AG)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Yves Nominé (Y)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Corinne Wendling (C)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Pierre Poussin-Courmontagne (P)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Laetitia Voilquin (L)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Pascal Eberling (P)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Frank Ruffenach (F)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Jean Cavarelli (J)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

John Slee (J)

UCL Institute of Ophthalmology, London, UK.

Timothy P Levine (TP)

UCL Institute of Ophthalmology, London, UK.

Guillaume Drin (G)

Institut de Pharmacologie Moléculaire et Cellulaire, Université Côte d'Azur, CNRS, Valbonne, France.

Catherine Tomasetto (C)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

Fabien Alpy (F)

Institut de Génétique et de Biologie Moléculaire et Cellulaire (IGBMC), Illkirch, France.
Institut National de la Santé et de la Recherche Médicale (INSERM), U 1258, Illkirch, France.
Centre National de la Recherche Scientifique (CNRS), UMR 7104, Illkirch, France.
Université de Strasbourg, Illkirch, France.

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