Evaluating the effect of spaceflight on the host-pathogen interaction between human intestinal epithelial cells and Salmonella Typhimurium.


Journal

NPJ microgravity
ISSN: 2373-8065
Titre abrégé: NPJ Microgravity
Pays: United States
ID NLM: 101703605

Informations de publication

Date de publication:
09 Mar 2021
Historique:
received: 27 08 2020
accepted: 03 02 2021
entrez: 22 3 2021
pubmed: 23 3 2021
medline: 23 3 2021
Statut: epublish

Résumé

Spaceflight uniquely alters the physiology of both human cells and microbial pathogens, stimulating cellular and molecular changes directly relevant to infectious disease. However, the influence of this environment on host-pathogen interactions remains poorly understood. Here we report our results from the STL-IMMUNE study flown aboard Space Shuttle mission STS-131, which investigated multi-omic responses (transcriptomic, proteomic) of human intestinal epithelial cells to infection with Salmonella Typhimurium when both host and pathogen were simultaneously exposed to spaceflight. To our knowledge, this was the first in-flight infection and dual RNA-seq analysis using human cells.

Identifiants

pubmed: 33750813
doi: 10.1038/s41526-021-00136-w
pii: 10.1038/s41526-021-00136-w
pmc: PMC7943786
doi:

Types de publication

Journal Article

Langues

eng

Pagination

9

Subventions

Organisme : NASA | Ames Research Center
ID : NNX09AH40G
Organisme : NASA | Ames Research Center
ID : NNX09AH40G
Organisme : NASA | Kennedy Space Center (KSC, Inc.)
ID : 80NSSC18K1478
Organisme : NASA | Kennedy Space Center (KSC, Inc.)
ID : 80NSSC18K1478

Références

Mcpee, J. C. & Charles, J. B. Human Health and Performance Risks of Space Exploration Missions. (US National Aeronautics and Space Admin; NASA SP-2009-3405 edition, 2010).
Crucian, B. E. et al. Immune system dysregulation during spaceflight: potential countermeasures for deep space exploration missions. Front Immunol. 9, 1437 (2018).
pubmed: 30018614 pmcid: 6038331 doi: 10.3389/fimmu.2018.01437
Voorhies, A. A. & Lorenzi, H. A. The challenge of maintaining a healthy microbiome during long-duration space missions. Front. Astron. Space Sci. 3, https://doi.org/10.3389/fspas.2016.00023 (2016).
Crucian, B. et al. Alterations in adaptive immunity persist during long-duration spaceflight. NPJ Microgravity 1, 15013 (2015).
pubmed: 28725716 pmcid: 5515498 doi: 10.1038/npjmgrav.2015.13
Castro, V. A., Thrasher, A. N., Healy, M., Ott, C. M. & Pierson, D. L. Microbial characterization during the early habitation of the International Space Station. Micro. Ecol. 47, 119–126 (2004).
doi: 10.1007/s00248-003-1030-y
Yamaguchi, N. et al. Microbial monitoring of crewed habitats in space-current status and future perspectives. Microbes Environ. 29, 250–260 (2014).
pubmed: 25130885 pmcid: 4159036 doi: 10.1264/jsme2.ME14031
Kish, A. L. et al. Biostability and Microbiological Analysis Of Shuttle Crew Refuse. (SAE Technical Paper #2002-01-2356, 2002).
Singh, N. K., Wood, J. M., Karouia, F. & Venkateswaran, K. Succession and persistence of microbial communities and antimicrobial resistance genes associated with International Space Station environmental surfaces. Microbiome 6, 204 (2018).
pubmed: 30424821 pmcid: 6234677 doi: 10.1186/s40168-018-0585-2
Wilson, J. W. et al. Space flight alters bacterial gene expression and virulence and reveals a role for global regulator Hfq. Proc. Natl Acad. Sci. USA 104, 16299–16304 (2007).
doi: 10.1073/pnas.0707155104 pubmed: 17901201 pmcid: 2042201
Wilson, J. W. et al. Media ion composition controls regulatory and virulence response of Salmonella in spaceflight. PLoS ONE 3, e3923 (2008).
pubmed: 19079590 pmcid: 2592540 doi: 10.1371/journal.pone.0003923
Wilson, J. W. et al. Low-Shear modeled microgravity alters the Salmonella enterica serovar typhimurium stress response in an RpoS-independent manner. Appl. Environ. Microbiol. 68, 5408–5416 (2002).
pubmed: 12406731 pmcid: 129924 doi: 10.1128/AEM.68.11.5408-5416.2002
Wilson, J. W. et al. Microarray analysis identifies Salmonella genes belonging to the low-shear modeled microgravity regulon. Proc. Natl Acad. Sci. USA 99, 13807–13812 (2002).
doi: 10.1073/pnas.212387899 pubmed: 12370447 pmcid: 129779
Gilbert, R. et al. Spaceflight and simulated microgravity conditions increase virulence of Serratia marcescens in the Drosophila melanogaster infection model. npj Microgravity 6, 4 (2020).
Nickerson, C. A, Pellis, N. R. & Ott, C. M. Effect of Spaceflight and Spaceflight Analogue Culture on Human and Microbial Cells: Novel Insights into Disease Mechanisms (Springer, 2016).
Horneck, G., Klaus, D. M. & Mancinelli, R. L. Space microbiology. Microbiol. Mol. Biol. Rev. 74, 121–156, (2010).
Mermel, L. A. Infection prevention and control during prolonged human space travel. Clin. Infect. Dis. 56, 123–130 (2013).
pubmed: 23051761 doi: 10.1093/cid/cis861
Barrila, J. et al. Modeling host-pathogen interactions in the context of the microenvironment: three-dimensional cell culture comes of age. Infect. Immun. 86, https://doi.org/10.1128/IAI.00282-18 (2018).
Bissell, M. J. & Aggeler, J. Dynamic reciprocity: how do extracellular matrix and hormones direct gene expression? Prog. Clin. Biol. Res 249, 251–262 (1987).
pubmed: 3671428
Ingber, D. E. How cells (might) sense microgravity. FASEB J. 13, S3–S15 (1999).
pubmed: 10352140 doi: 10.1096/fasebj.13.9001.s3
Hughes-Fulford, M. Function of the cytoskeleton in gravisensing during spaceflight. Adv. Space Res. 32, 1585–1593 (2003).
pubmed: 15002415 doi: 10.1016/S0273-1177(03)90399-1
Lewis, M. L. The cytoskeleton, apoptosis, and gene expression in T lymphocytes and other mammalian cells exposed to altered gravity. Adv. Space Biol. Med. 8, 77–128 (2002).
pubmed: 12951694 doi: 10.1016/S1569-2574(02)08016-4
Ibarra, J. A. & Steele-Mortimer, O. Salmonella-the ultimate insider. Salmonella virulence factors that modulate intracellular survival. Cell Microbiol. 11, 1579–1586 (2009).
pubmed: 19775254 pmcid: 2774479 doi: 10.1111/j.1462-5822.2009.01368.x
Gruenheid, S. & Finlay, B. B. Microbial pathogenesis and cytoskeletal function. Nature 422, 775–781 (2003).
pubmed: 12700772 doi: 10.1038/nature01603
Nickerson, C. A. et al. Microgravity as a novel environmental signal affecting Salmonella enterica Serovar typhimurium virulence. Infect. Immun. 68, 3147–3152 (2000).
pubmed: 10816456 pmcid: 97548 doi: 10.1128/IAI.68.6.3147-3152.2000
Castro, S. L., Nelman-Gonzalez, M., Nickerson, C. A. & Ott, C. M. Induction of attachment-independent biofilm formation and repression of Hfq expression by low-fluid-shear culture of Staphylococcus aureus. Appl. Environ. Microbiol. 77, 6368–6378 (2011).
pubmed: 21803898 pmcid: 3187170 doi: 10.1128/AEM.00175-11
Crabbé, A. et al. Response of Pseudomonas aeruginosa PAO1 to low shear modelled microgravity involves AlgU regulation. Environ. Microbiol. 12, 1545–1564 (2010).
pubmed: 20236169
Crabbe, A. et al. Transcriptional and proteomic responses of Pseudomonas aeruginosa PAO1 to spaceflight conditions involve Hfq regulation and reveal a role for oxygen. Appl. Environ. Microbiol. 77, 1221–1230 (2011).
pubmed: 21169425 doi: 10.1128/AEM.01582-10
Grant, K. C., Khodadad, C. L. M. & Foster, J. S. Role of Hfq in an animal–microbe symbiosis under simulated microgravity conditions. Int. J. Astrobiol. 13, 53–61 (2014).
doi: 10.1017/S1473550413000359
Larsen, S. B., Cowley, C. J. & Fuchs, E. Epithelial cells: liaisons of immunity. Curr. Opin. Immunol. 62, 45–53 (2020).
pubmed: 31874430 doi: 10.1016/j.coi.2019.11.004
Allaire, J. M. et al. The intestinal epithelium: central coordinator of mucosal immunity. Trends Immunol. 39, 677–696, (2018).
pubmed: 29716793 doi: 10.1016/j.it.2018.04.002
Carregaro, F., Stefanini, A. C., Henrique, T. & Tajara, E. H. Study of small proline-rich proteins (SPRRs) in health and disease: a review of the literature. Arch. Dermatol. Res. 305, 857–866 (2013).
pubmed: 24085571 doi: 10.1007/s00403-013-1415-9
Pradervand, S. et al. Small proline-rich protein 1A is a gp130 pathway- and stress-inducible cardioprotective protein. EMBO J. 23, 4517–4525 (2004).
pubmed: 15510217 pmcid: 526469 doi: 10.1038/sj.emboj.7600454
Pyle, A. L. et al. Regulation of the atheroma-enriched protein, SPRR3, in vascular smooth muscle cells through cyclic strain is dependent on integrin alpha1beta1/collagen interaction. Am. J. Pathol. 173, 1577–1588 (2008).
pubmed: 18832573 pmcid: 2570146 doi: 10.2353/ajpath.2008.080042
Pecaut, M. J. et al. Is spaceflight-induced immune dysfunction linked to systemic changes in metabolism? PLOS ONE 12, e0174174 (2017).
McNab, F., Mayer-Barber, K., Sher, A., Wack, A. & O’Garra, A. Type I interferons in infectious disease. Nat. Rev. Immunol. 15, 87–103 (2015).
pubmed: 25614319 pmcid: 7162685 doi: 10.1038/nri3787
Kotredes, K. P., Thomas, B. & Gamero, A. M. The protective role of type I interferons in the gastrointestinal tract. Front. Immunol. 8, 410 (2017).
pubmed: 28428788 pmcid: 5382159 doi: 10.3389/fimmu.2017.00410
Katlinskaya, Y. V. et al. Type I interferons control proliferation and function of the intestinal epithelium. Mol. Cell Biol. 36, 1124–1135 (2016).
pubmed: 26811327 pmcid: 4800802 doi: 10.1128/MCB.00988-15
Shi, L. et al. Spaceflight and simulated microgravity suppresses macrophage development via altered RAS/ERK/NFkappaB and metabolic pathways. Cell. Mol. Immunol. https://doi.org/10.1038/s41423-019-0346-6 (2020).
Fritz, V. & Fajas, L. Metabolism and proliferation share common regulatory pathways in cancer cells. Oncogene 29, 4369–4377 (2010).
pubmed: 20514019 pmcid: 3004916 doi: 10.1038/onc.2010.182
Leschner, S. & Weiss, S. Salmonella-allies in the fight against cancer. J. Mol. Med. 88, 763–773 (2010).
pubmed: 20526574 doi: 10.1007/s00109-010-0636-z
Honer zu Bentrup, K. et al. Three-dimensional organotypic models of human colonic epithelium to study the early stages of enteric salmonellosis. Microbes Infect. 8, 1813–1825 (2006).
pubmed: 16730210 doi: 10.1016/j.micinf.2006.02.020
Barker, N., Tan, S. & Clevers, H. Lgr proteins in epithelial stem cell biology. Development 140, 2484–2494 (2013).
pubmed: 23715542 doi: 10.1242/dev.083113
Field, C. J., Johnson, I. R. & Schley, P. D. Nutrients and their role in host resistance to infection. J. Leukoc. Biol. 71, 16–32 (2002).
pubmed: 11781377 doi: 10.1189/jlb.71.1.16
Steeb, B. et al. Parallel exploitation of diverse host nutrients enhances Salmonella virulence. PLoS Pathog. 9, e1003301 (2013).
pubmed: 23633950 pmcid: 3636032 doi: 10.1371/journal.ppat.1003301
Jessup, J. M. et al. Microgravity culture reduces apoptosis and increases the differentiation of a human colorectal carcinoma cell line. Vitr. Cell Dev. Biol. Anim. 36, 367–373 (2000).
doi: 10.1290/1071-2690(2000)036<0367:MCRAAI>2.0.CO;2
Kumar, S., Suman, S., Fornace, A. J. Jr & Datta, K. Space radiation triggers persistent stress response, increases senescent signaling, and decreases cell migration in mouse intestine. Proc. Natl Acad. Sci. USA 115, E9832–E9841 (2018).
pubmed: 30275302 doi: 10.1073/pnas.1807522115 pmcid: 6196540
Moreno-Villanueva, M., Wong, M., Lu, T., Zhang, Y. & Wu, H. Interplay of space radiation and microgravity in DNA damage and DNA damage response. NPJ Microgravity 3, 14 (2017).
pubmed: 28649636 pmcid: 5460239 doi: 10.1038/s41526-017-0019-7
Garrett-Bakelman, F. E. et al. The NASA Twins Study: a multidimensional analysis of a year-long human spaceflight. Science 364. https://doi.org/10.1126/science.aau8650 (2019).
George, K., Rhone, J., Beitman, A. & Cucinotta, F. A. Cytogenetic damage in the blood lymphocytes of astronauts: effects of repeat long-duration space missions. Mutat. Res. 756, 165–169 (2013).
pubmed: 23639573 doi: 10.1016/j.mrgentox.2013.04.007
Barrila, J. et al. Spaceflight modulates gene expression in the whole blood of astronauts. npj Microgravity 2, 16039 (2016).
pubmed: 28725744 pmcid: 5515525 doi: 10.1038/npjmgrav.2016.39
Simpson-Haidaris, P. J. et al. Induction of fibrinogen expression in the lung epithelium during Pneumocystis carinii Pneumonia. Infect. Immun. 66, 4431–4439 (1998).
pubmed: 9712798 pmcid: 108536 doi: 10.1128/IAI.66.9.4431-4439.1998
Molmenti, E. P., Ziambaras, T. & Perlmutter, D. H. Evidence for an acute phase response in human intestinal epithelial cells. J. Biol. Chem. 268, 14116–14124 (1993).
pubmed: 7686149 doi: 10.1016/S0021-9258(19)85216-X
Andoh, A. et al. Detection of complement C3 and factor B gene expression in normal colorectal mucosa, adenomas and carcinomas. Clin. Exp. Immunol. 111, 477–483 (1998).
pubmed: 9528886 pmcid: 1904873 doi: 10.1046/j.1365-2249.1998.00496.x
Kulkarni, H. S., Liszewski, M. K., Brody, S. L. & Atkinson, J. P. The complement system in the airway epithelium: an overlooked host defense mechanism and therapeutic target? J. Allergy Clin. Immunol. 141, 1582–1586 e1581 (2018).
pubmed: 29339260 pmcid: 5955701 doi: 10.1016/j.jaci.2017.11.046
Wei, Z., Batagov, A. O., Carter, D. R. & Krichevsky, A. M. Fetal bovine serum RNA interferes with the cell culture derived extracellular RNA. Sci. Rep. 6, 31175 (2016).
pubmed: 27503761 pmcid: 4977539 doi: 10.1038/srep31175
Schleker, S. et al. The current Salmonella-host interactome. Proteom. Clin. Appl. 6, 117–133 (2012).
doi: 10.1002/prca.201100083
Maier, T., Guell, M. & Serrano, L. Correlation of mRNA and protein in complex biological samples. FEBS Lett. 583, 3966–3973 (2009).
pubmed: 19850042 doi: 10.1016/j.febslet.2009.10.036
Broz, P., Ohlson, M. B. & Monack, D. M. Innate immune response to Salmonella typhimurium, a model enteric pathogen. Gut Microbes 3, 62–70 (2012).
pubmed: 22198618 pmcid: 3370950 doi: 10.4161/gmic.19141
Keating, S. E., Maloney, G. M., Moran, E. M. & Bowie, A. G. IRAK-2 participates in multiple toll-like receptor signaling pathways to NFkappaB via activation of TRAF6 ubiquitination. J. Biol. Chem. 282, 33435–33443 (2007).
pubmed: 17878161 doi: 10.1074/jbc.M705266200
Muller, A. et al. IkappaBzeta is a key transcriptional regulator of IL-36-driven psoriasis-related gene expression in keratinocytes. Proc. Natl Acad. Sci. USA 115, 10088–10093 (2018).
pubmed: 30224457 doi: 10.1073/pnas.1801377115 pmcid: 6176600
Yamazaki, S., Muta, T. & Takeshige, K. A novel IkappaB protein, IkappaB-zeta, induced by proinflammatory stimuli, negatively regulates nuclear factor-kappaB in the nuclei. J. Biol. Chem. 276, 27657–27662 (2001).
pubmed: 11356851 doi: 10.1074/jbc.M103426200
Chen, X. et al. RelB sustains IkappaBalpha expression during endotoxin tolerance. Clin. Vaccin. Immunol. 16, 104–110 (2009).
doi: 10.1128/CVI.00320-08
Le Negrate, G. et al. Salmonella secreted factor L deubiquitinase of Salmonella typhimurium inhibits NF-kappaB, suppresses IkappaBalpha ubiquitination and modulates innate immune responses. J. Immunol. 180, 5045–5056 (2008).
pubmed: 18354230 doi: 10.4049/jimmunol.180.7.5045
Sun, H., Kamanova, J., Lara-Tejero, M. & Galan, J. E. A family of Salmonella type IIi secretion effector proteins selectively targets the NF-kappaB signaling pathway to preserve host homeostasis. PLoS Pathog. 12, e1005484 (2016).
pubmed: 26933955 pmcid: 4775039 doi: 10.1371/journal.ppat.1005484
Kaur, I., Simons, E. R., Kapadia, A. S., Ott, C. M. & Pierson, D. L. Effect of spaceflight on ability of monocytes to respond to endotoxins of gram-negative bacteria. Clin. Vaccin. Immunol. 15, 1523–1528 (2008).
doi: 10.1128/CVI.00065-08
Jin, M. et al. Responses of intestinal mucosal barrier functions of rats to simulated weightlessness. Front. Physiol. 9, 729 (2018).
pubmed: 29962963 pmcid: 6011188 doi: 10.3389/fphys.2018.00729
Mednieks, M. H. A. In Oral Tissue Responses to Travel in Space. Beyond LEO - Human Health Issues for Deep Space Exploration (ed Reynolds, R. J.) (IntechOpen, 2019).
Spielmann, G. et al. B cell homeostasis is maintained during long-duration spaceflight. J. Appl. Physiol. 126, 469–476 (2018).
pubmed: 30496712 pmcid: 6397409 doi: 10.1152/japplphysiol.00789.2018
Aase, A. et al. Salivary IgA from the sublingual compartment as a novel noninvasive proxy for intestinal immune induction. Mucosal Immunol. 9, 884–893 (2016).
pubmed: 26509875 doi: 10.1038/mi.2015.107
Externest, D., Meckelein, B., Schmidt, M. A. & Frey, A. Correlations between antibody immune responses at different mucosal effector sites are controlled by antigen type and dosage. Infect. Immun. 68, 3830–3839 (2000).
pubmed: 10858191 pmcid: 101655 doi: 10.1128/IAI.68.7.3830-3839.2000
Ding, X. Z. et al. HSP-70 mitigates LPS/SKI-induced cell damage by increasing sphingosine kinase 1 (SK1). Prostaglandins Other Lipid Mediat. 92, 1–7 (2010).
pubmed: 20123033 doi: 10.1016/j.prostaglandins.2009.12.006
Zupanska, A. K., Denison, F. C., Ferl, R. J. & Paul, A. L. Spaceflight engages heat shock protein and other molecular chaperone genes in tissue culture cells of Arabidopsis thaliana. Am. J. Bot. 100, 235–248 (2013).
pubmed: 23258370 doi: 10.3732/ajb.1200343
Audy, J., Mathieu, O., Belvis, J. & Tompkins, T. A. Transcriptomic response of immune signalling pathways in intestinal epithelial cells exposed to lipopolysaccharides, Gram-negative bacteria or potentially probiotic microbes. Benef. Microbes 3, 273–286 (2012).
pubmed: 23234729 doi: 10.3920/BM2012.0027
Crucian, B., Stowe, R., Quiriarte, H., Pierson, D. & Sams, C. Monocyte phenotype and cytokine production profiles are dysregulated by short-duration spaceflight. Aviat. Space Environ. Med. 82, 857–862 (2011).
pubmed: 21888268 doi: 10.3357/ASEM.3047.2011
Chakraborty, N. et al. An integrated omics analysis: impact of microgravity on host response to lipopolysaccharide in vitro. BMC Genom. 15, 659 (2014).
doi: 10.1186/1471-2164-15-659
Vogel, J. & Luisi, B. F. Hfq and its constellation of RNA. Nat. Rev. Microbiol. 9, 578–589 (2011).
pubmed: 21760622 pmcid: 4615618 doi: 10.1038/nrmicro2615
Morita, T. & Aiba, H. Mechanism and physiological significance of autoregulation of the Escherichia coli hfq gene. RNA 25, 264–276 (2019).
pubmed: 30487269 pmcid: 6348989 doi: 10.1261/rna.068106.118
Voorhies, A. A. et al. Study of the impact of long-duration space missions at the International Space Station on the astronaut microbiome. Sci. Rep. 9, 9911 (2019).
pubmed: 31289321 pmcid: 6616552 doi: 10.1038/s41598-019-46303-8
Castro-Wallace, S., Stahl, S., Voorhies, A., Lorenzi, H. & Douglas, G. L. Response of Lactobacillus acidophilus ATCC 4356 to low-shear modeled microgravity. Acta Astronautica 139, 463–468 (2017).
doi: 10.1016/j.actaastro.2017.07.033
Shao, D. et al. Simulated microgravity affects some biological characteristics of Lactobacillus acidophilus. Appl. Microbiol. Biotechnol. 101, 3439–3449 (2017).
pubmed: 28013406 doi: 10.1007/s00253-016-8059-6
Turroni, S. et al. Gut microbiome and space travelers’ health: state of the art and possible pro/prebiotic strategies for long-term space missions. Front. Physiol. 11, https://doi.org/10.3389/fphys.2020.553929 (2020).
Hansmeier, N., Chao, T. C., Goldman, L. R., Witter, F. R. & Halden, R. U. Prioritization of biomarker targets in human umbilical cord blood: identification of proteins in infant blood serving as validated biomarkers in adults. Environ. Health Perspect. 120, 764–769 (2012).
pubmed: 22538116 pmcid: 3346780 doi: 10.1289/ehp.1104190

Auteurs

Jennifer Barrila (J)

Biodesign Center for Fundamental and Applied Microbiomics, Arizona State University, Tempe, AZ, USA.
Biodesign Center for Infectious Diseases and Vaccinology, Arizona State University, Tempe, AZ, USA.

Shameema F Sarker (SF)

Biodesign Center for Infectious Diseases and Vaccinology, Arizona State University, Tempe, AZ, USA.

Nicole Hansmeier (N)

Swette Center for Environmental Biotechnology, The Biodesign Institute, Arizona State University, Tempe, AZ, USA.
Luther College at University of Regina, Department of Biology, Regina, Saskatchewan, Canada.

Shanshan Yang (S)

Bioinformatics Core Facility, Bioscience, Knowledge Enterprise, Arizona State University, Tempe, AZ, USA.

Kristina Buss (K)

Bioinformatics Core Facility, Bioscience, Knowledge Enterprise, Arizona State University, Tempe, AZ, USA.

Natalia Briones (N)

Bioinformatics Core Facility, Bioscience, Knowledge Enterprise, Arizona State University, Tempe, AZ, USA.
Integrated Cancer Genomics Division, The Translational Genomics Research Institute, Phoenix, AZ, USA.

Jin Park (J)

Bioinformatics Core Facility, Bioscience, Knowledge Enterprise, Arizona State University, Tempe, AZ, USA.

Richard R Davis (RR)

Biodesign Center for Fundamental and Applied Microbiomics, Arizona State University, Tempe, AZ, USA.
Biodesign Center for Infectious Diseases and Vaccinology, Arizona State University, Tempe, AZ, USA.

Rebecca J Forsyth (RJ)

Biodesign Center for Infectious Diseases and Vaccinology, Arizona State University, Tempe, AZ, USA.

C Mark Ott (CM)

Biomedical Research and Environmental Sciences Division, NASA Johnson Space Center, Houston, TX, USA.

Kevin Sato (K)

NASA Ames Research Center, Mountain View, CA, USA.

Cristine Kosnik (C)

Tissue Genesis, Inc, Honolulu, HI, USA.

Anthony Yang (A)

Tissue Genesis, Inc, Honolulu, HI, USA.

Cheryl Shimoda (C)

Tissue Genesis, Inc, Honolulu, HI, USA.

Nicole Rayl (N)

NASA Ames Research Center, Mountain View, CA, USA.
NASA Headquarters, Washington, D.C., USA.

Diana Ly (D)

NASA Ames Research Center, Mountain View, CA, USA.

Aaron Landenberger (A)

DoD Space Test Program, Houston, TX, USA.

Stephanie D Wilson (SD)

Astronaut Office, NASA Johnson Space Center, Houston, TX, USA.

Naoko Yamazaki (N)

Space Biomedical Research Office, Human Space Technology and Astronauts Department, Japan Aerospace Exploration Agency (JAXA), Tokyo, Japan.

Jason Steel (J)

Bioinformatics Core Facility, Bioscience, Knowledge Enterprise, Arizona State University, Tempe, AZ, USA.

Camila Montano (C)

Bioinformatics Core Facility, Bioscience, Knowledge Enterprise, Arizona State University, Tempe, AZ, USA.

Rolf U Halden (RU)

Biodesign Center for Environmental Health Engineering, School of Sustainable Engineering and the Built Environment, Arizona State University, Tempe, AZ, USA.

Tom Cannon (T)

Tissue Genesis, Inc, Honolulu, HI, USA.

Sarah L Castro-Wallace (SL)

Biomedical Research and Environmental Sciences Division, NASA Johnson Space Center, Houston, TX, USA.

Cheryl A Nickerson (CA)

Biodesign Center for Fundamental and Applied Microbiomics, Arizona State University, Tempe, AZ, USA. Cheryl.Nickerson@asu.edu.
Biodesign Center for Infectious Diseases and Vaccinology, Arizona State University, Tempe, AZ, USA. Cheryl.Nickerson@asu.edu.
School of Life Sciences, Arizona State University, Tempe, AZ, USA. Cheryl.Nickerson@asu.edu.

Classifications MeSH