Ageing affects subtelomeric DNA methylation in blood cells from a large European population enrolled in the MARK-AGE study.


Journal

GeroScience
ISSN: 2509-2723
Titre abrégé: Geroscience
Pays: Switzerland
ID NLM: 101686284

Informations de publication

Date de publication:
06 2021
Historique:
received: 09 12 2020
accepted: 23 02 2021
pubmed: 20 4 2021
medline: 2 7 2021
entrez: 19 4 2021
Statut: ppublish

Résumé

Ageing leaves characteristic traces in the DNA methylation make-up of the genome. However, the importance of DNA methylation in ageing remains unclear. The study of subtelomeric regions could give promising insights into this issue. Previously reported associations between susceptibility to age-related diseases and epigenetic instability at subtelomeres suggest that the DNA methylation profile of subtelomeres undergoes remodelling during ageing. In the present work, this hypothesis has been tested in the context of the European large-scale project MARK-AGE. In this cross-sectional study, we profiled the DNA methylation of chromosomes 5 and 21 subtelomeres, in more than 2000 age-stratified women and men recruited in eight European countries. The study included individuals from the general population as well as the offspring of nonagenarians and Down syndrome subjects, who served as putative models of delayed and accelerated ageing, respectively. Significant linear changes of subtelomeric DNA methylation with increasing age were detected in the general population, indicating that subtelomeric DNA methylation changes are typical signs of ageing. Data also show that, compared to the general population, the dynamics of age-related DNA methylation changes are attenuated in the offspring of centenarian, while they accelerate in Down syndrome individuals. This result suggests that subtelomeric DNA methylation changes reflect the rate of ageing progression. We next attempted to trace the age-related changes of subtelomeric methylation back to the influence of diverse variables associated with methylation variations in the population, including demographics, dietary/health habits and clinical parameters. Results indicate that the effects of age on subtelomeric DNA methylation are mostly independent of all other variables evaluated.

Identifiants

pubmed: 33870444
doi: 10.1007/s11357-021-00347-9
pii: 10.1007/s11357-021-00347-9
pmc: PMC8190237
doi:

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

1283-1302

Références

Linardopoulou EV, Williams EM, Fan Y, Friedman C, Young JM, Trask BJ. Human subtelomeres are hot spots of interchromosomal recombination and segmental duplication. Nature. 2005;437(7055):94–100.
pubmed: 16136133 pmcid: 1368961
Mefford HC, Trask BJ. The complex structure and dynamic evolution of human subtelomeres. Nature reviews. 2002;3(2):91–102.
pubmed: 11836503
Riethman H, Ambrosini A, Paul S. Human subtelomere structure and variation. Chromosom Res. 2005;13(5):505–15.
Blasco MA. The epigenetic regulation of mammalian telomeres. Nature reviews. 2007;8(4):299–309.
pubmed: 17363977
Mikkelsen TS, Ku M, Jaffe DB, Issac B, Lieberman E, Giannoukos G, et al. Genome-wide maps of chromatin state in pluripotent and lineage-committed cells. Nature. 2007;448(7153):553–60.
pubmed: 17603471 pmcid: 2921165
Gonzalo S, Jaco I, Fraga MF, Chen T, Li E, Esteller M, et al. DNA methyltransferases control telomere length and telomere recombination in mammalian cells. Nat Cell Biol. 2006;8(4):416–24.
pubmed: 16565708
Vera E, Canela A, Fraga MF, Esteller M, Blasco MA. Epigenetic regulation of telomeres in human cancer. Oncogene. 2008;27(54):6817–33.
pubmed: 18762811
Feil R, Fraga MF. Epigenetics and the environment: emerging patterns and implications. Nature reviews. 2012;13(2):97–109.
pubmed: 22215131
Schubeler D. Function and information content of DNA methylation. Nature. 2015;517(7534):321–6.
pubmed: 25592537
Lopez-Otin C, Blasco MA, Partridge L, Serrano M, Kroemer G. The hallmarks of aging. Cell. 2013;153(6):1194–217.
pubmed: 23746838 pmcid: 3836174
Ciccarone F, Tagliatesta S, Caiafa P, Zampieri M. DNA methylation dynamics in aging: how far are we from understanding the mechanisms? Mech Ageing Dev. 2018;174:3–17.
pubmed: 29268958
Pal S, Tyler JK. Epigenetics and aging. Sci Adv. 2016;2(7):e1600584.
pubmed: 27482540 pmcid: 4966880
Bell CG, Lowe R, Adams PD, Baccarelli AA, Beck S, Bell JT, et al. DNA methylation aging clocks: challenges and recommendations. Genome Biol. 2019;20(1):249.
pubmed: 31767039 pmcid: 6876109
Horvath S, Raj K. DNA methylation-based biomarkers and the epigenetic clock theory of ageing. Nature reviews. 2018;19(6):371–84.
pubmed: 29643443
Zhang Q, Vallerga CL, Walker RM, Lin T, Henders AK, Montgomery GW, et al. Improved precision of epigenetic clock estimates across tissues and its implication for biological ageing. Genome medicine. 2019;11(1):54.
pubmed: 31443728 pmcid: 6708158
Levine ME, Lu AT, Quach A, Chen BH, Assimes TL, Bandinelli S, et al. An epigenetic biomarker of aging for lifespan and healthspan. Aging. 2018;10(4):573–91.
pubmed: 29676998 pmcid: 5940111
Hu H, Li B, Duan S. The alteration of subtelomeric DNA methylation in aging-related diseases. Front Genet. 2019;9:697.
pubmed: 30687384 pmcid: 6333653
Buxton JL, Suderman M, Pappas JJ, Borghol N, McArdle W, Blakemore AI, et al. Human leukocyte telomere length is associated with DNA methylation levels in multiple subtelomeric and imprinted loci. Sci Rep. 2014;4:4954.
pubmed: 24828261 pmcid: 4344300
Guan JZ, Guan WP, Maeda T, Makino N. The subtelomere of short telomeres is hypermethylated in Alzheimer’s disease. Aging Dis. 2012;3(2):164–70.
pubmed: 22724077
Guan JZ, Guan WP, Maeda T, Makino N. Analysis of telomere length and subtelomeric methylation of circulating leukocytes in women with Alzheimer’s disease. Aging Clin Exp Res. 2013;25(1):17–23.
pubmed: 23740629
Maeda T, Guan JZ, Koyanagi M, Higuchi Y, Makino N. Aging-associated alteration of telomere length and subtelomeric status in female patients with Parkinson’s disease. J Neurogenet. 2012;26(2):245–51.
pubmed: 22364520
Maeda T, Guan JZ, Oyama J, Higuchi Y, Makino N. Aging-associated alteration of subtelomeric methylation in Parkinson’s disease. J Gerontol A Biol Sci Med Sci. 2009;64(9):949–55.
pubmed: 19502593
Makino N, Maeda T, Abe N. Short telomere subtelomeric hypomethylation is associated with telomere attrition in elderly diabetic patients. Can J Physiol Pharmacol. 2019;97(4):335–9.
pubmed: 30785764
Choudhury SR, Cui Y, Milton JR, Li J, Irudayaraj J. Selective increase in subtelomeric DNA methylation: an epigenetic biomarker for malignant glioma. Clin Epigenetics. 2015;7:107.
pubmed: 26451167 pmcid: 4597615
Han Y, Xu J, Kim J, Wu X, Gu J. Methylation of subtelomeric repeat D4Z4 in peripheral blood leukocytes is associated with biochemical recurrence in localized prostate cancer patients. Carcinogenesis. 2017;38(8):821–6.
pubmed: 28854562 pmcid: 6248474
Lee ME, Rha SY, Jeung HC, Chung HC, Oh BK. Subtelomeric DNA methylation and telomere length in human cancer cells. Cancer Lett. 2009;281(1):82–91.
pubmed: 19375218
Oh BK, Um TH, Choi GH, Park YN. Frequent changes in subtelomeric DNA methylation patterns and its relevance to telomere regulation during human hepatocarcinogenesis. Int J Cancer. 2011;128(4):857–68.
pubmed: 20473888
Xu J, Tsai CW, Chang WS, Han Y, Bau DT, Pettaway CA, et al. Methylation of global DNA repeat LINE-1 and subtelomeric DNA repeats D4Z4 in leukocytes is associated with biochemical recurrence in African American prostate cancer patients. Carcinogenesis. 2019;40:1055–60.
Maeda T, Guan JZ, Oyama J, Higuchi Y, Makino N. Age-related changes in subtelomeric methylation in the normal Japanese population. J Gerontol A Biol Sci Med Sci. 2009;64(4):426–34.
pubmed: 19223605
Burkle A, Moreno-Villanueva M, Bernhard J, Blasco M, Zondag G, Hoeijmakers JH, et al. MARK-AGE biomarkers of ageing. Mech Ageing Dev. 2015;151:2–12.
pubmed: 25818235
Capri M, Moreno-Villanueva M, Cevenini E, Pini E, Scurti M, Borelli V, et al. MARK-AGE population: from the human model to new insights. Mech Ageing Dev. 2015;151:13–7.
pubmed: 25843237
Moreno-Villanueva M, Kotter T, Sindlinger T, Baur J, Oehlke S, Burkle A, et al. The MARK-AGE phenotypic database: structure and strategy. Mech Ageing Dev. 2015;151:26–30.
pubmed: 25817205
Moreno-Villanueva M, Capri M, Breusing N, Siepelmeyer A, Sevini F, Ghezzo A, et al. MARK-AGE standard operating procedures (SOPs): a successful effort. Mech Ageing Dev. 2015;151:18–25.
pubmed: 25817206
Baur J, Kotter T, Moreno-Villanueva M, Sindlinger T, Berthold MR, Burkle A, et al. The MARK-AGE extended database: data integration and pre-processing. Mech Ageing Dev. 2015;151:31–7.
pubmed: 26004672
Baur J, Moreno-Villanueva M, Kotter T, Sindlinger T, Burkle A, Berthold MR, et al. MARK-AGE data management: cleaning, exploration and visualization of data. Mech Ageing Dev. 2015;151:38–44.
pubmed: 26004801
Ciccarone F, Malavolta M, Calabrese R, Guastafierro T, Bacalini MG, Reale A, et al. Age-dependent expression of DNMT1 and DNMT3B in PBMCs from a large European population enrolled in the MARK-AGE study. Aging Cell. 2016;15(4):755–65.
pubmed: 27169697 pmcid: 4933658
Zampieri M, Ciccarone F, Palermo R, Cialfi S, Passananti C, Chiaretti S, et al. The epigenetic factor BORIS/CTCFL regulates the NOTCH3 gene expression in cancer cells. Biochim Biophys Acta. 2014;1839(9):813–25.
pubmed: 24984200
Suchiman HE, Slieker RC, Kremer D, Slagboom PE, Heijmans BT, Tobi EW. Design, measurement and processing of region-specific DNA methylation assays: the mass spectrometry-based method EpiTYPER. Front Genet. 2015;6:287.
pubmed: 26442105 pmcid: 4585020
Franceschi C, Bezrukov V, Blanche H, Bolund L, Christensen K, de Benedictis G, et al. Genetics of healthy aging in Europe: the EU-integrated project GEHA (GEnetics of Healthy Aging). Ann N Y Acad Sci. 2007;1100:21–45.
pubmed: 17460163
Ambrosini A, Paul S, Hu S, Riethman H. Human subtelomeric duplicon structure and organization. Genome Biol. 2007;8(7):R151.
pubmed: 17663781 pmcid: 2323237
Young E, Abid HZ, Kwok PY, Riethman H, Xiao M. Comprehensive analysis of human subtelomeres by whole genome mapping. PLoS Genet. 2020;16(1):e1008347.
pubmed: 31986135 pmcid: 7004388
Maeda T, Guan JZ, Higuchi Y, Oyama J, Makino N. Aging-related alterations of subtelomeric methylation in sarcoidosis patients. J Gerontol A Biol Sci Med Sci. 2009;64(7):752–60.
pubmed: 19414507
Atzmon G, Rincon M, Rabizadeh P, Barzilai N. Biological evidence for inheritance of exceptional longevity. Mech Ageing Dev. 2005;126(2):341–5.
pubmed: 15621216
Patterson D, Cabelof DC. Down syndrome as a model of DNA polymerase beta haploinsufficiency and accelerated aging. Mech Ageing Dev. 2012;133(4):133–7.
pubmed: 22019846
Yehezkel S, Rebibo-Sabbah A, Segev Y, Tzukerman M, Shaked R, Huber I, et al. Reprogramming of telomeric regions during the generation of human induced pluripotent stem cells and subsequent differentiation into fibroblast-like derivatives. Epigenetics. 2011;6(1):63–75.
pubmed: 20861676 pmcid: 3052915
Elliott G, Hong C, Xing X, Zhou X, Li D, Coarfa C, et al. Intermediate DNA methylation is a conserved signature of genome regulation. Nat Commun. 2015;6:6363.
pubmed: 25691127 pmcid: 4333717
Slieker RC, van Iterson M, Luijk R, Beekman M, Zhernakova DV, Moed MH, et al. Age-related accrual of methylomic variability is linked to fundamental ageing mechanisms. Genome Biol. 2016;17(1):191.
pubmed: 27654999 pmcid: 5032245
Robin JD, Ludlow AT, Batten K, Magdinier F, Stadler G, Wagner KR, et al. Telomere position effect: regulation of gene expression with progressive telomere shortening over long distances. Genes Dev. 2014;28(22):2464–76.
pubmed: 25403178 pmcid: 4233240
Chu HP, Cifuentes-Rojas C, Kesner B, Aeby E, Lee HG, Wei C, et al. TERRA RNA antagonizes ATRX and protects telomeres. Cell. 2017;170(1):86–101 e16.
pubmed: 28666128 pmcid: 5552367
Brando B, Longo A, Beltrami B, Passoni D, Verna R, Licastro F, et al. Determination of telomere length by flow-fluorescence in situ hybridization in Down’s syndrome patients. Int J Tissue React. 2004;26(1–2):61–4.
pubmed: 15573694
Vaziri H, Schachter F, Uchida I, Wei L, Zhu X, Effros R, et al. Loss of telomeric DNA during aging of normal and trisomy 21 human lymphocytes. Am J Hum Genet. 1993;52(4):661–7.
pubmed: 8460632 pmcid: 1682068
Horvath S, Pirazzini C, Bacalini MG, Gentilini D, Di Blasio AM, Delledonne M, et al. Decreased epigenetic age of PBMCs from Italian semi-supercentenarians and their offspring. Aging. 2015;7(12):1159–70.
pubmed: 26678252 pmcid: 4712339
Arai Y, Martin-Ruiz CM, Takayama M, Abe Y, Takebayashi T, Koyasu S, et al. Inflammation, but not telomere length, predicts successful ageing at extreme old age: a longitudinal study of semi-supercentenarians. EBioMedicine. 2015;2(10):1549–58.
pubmed: 26629551 pmcid: 4634197
Atzmon G, Cho M, Cawthon RM, Budagov T, Katz M, Yang X, et al. Evolution in health and medicine Sackler colloquium: genetic variation in human telomerase is associated with telomere length in Ashkenazi centenarians. Proc Natl Acad Sci U S A. 2010;107(Suppl 1):1710–7.
pubmed: 19915151
Tedone E, Arosio B, Gussago C, Casati M, Ferri E, Ogliari G, et al. Leukocyte telomere length and prevalence of age-related diseases in semisupercentenarians, centenarians and centenarians’ offspring. Exp Gerontol. 2014;58:90–5.
pubmed: 24975295
Colacino JA, Arthur AE, Dolinoy DC, Sartor MA, Duffy SA, Chepeha DB, et al. Pretreatment dietary intake is associated with tumor suppressor DNA methylation in head and neck squamous cell carcinomas. Epigenetics. 2012;7(8):883–91.
pubmed: 22722388 pmcid: 3427284
Garcia-Calzon S, Moleres A, Martinez-Gonzalez MA, Martinez JA, Zalba G, Marti A. Dietary total antioxidant capacity is associated with leukocyte telomere length in a children and adolescent population. Clin Nutr. 2015;34(4):694–9.
pubmed: 25131600
Liu C, Marioni RE, Hedman AK, Pfeiffer L, Tsai PC, Reynolds LM, et al. A DNA methylation biomarker of alcohol consumption. Mol Psychiatry. 2018;23(2):422–33.
pubmed: 27843151
Wilson LE, Xu Z, Harlid S, White AJ, Troester MA, Sandler DP, et al. Alcohol and DNA methylation: an epigenome-wide association study in blood and normal breast tissue. Am J Epidemiol. 2019;188(6):1055–65.
pubmed: 30938765 pmcid: 6545285
Harpaz T, Abumock H, Beery E, Edel Y, Lahav M, Rozovski U, et al. The effect of ethanol on telomere dynamics and regulation in human cells. Cells. 2018;7(10).
Kirchner H, Shaheen F, Kalscheuer H, Schmid SM, Oster H, Lehnert H. The telomeric complex and metabolic disease. Genes (Basel). 2017;8(7).
Revesz D, Milaneschi Y, Verhoeven JE, Penninx BW. Telomere length as a marker of cellular aging is associated with prevalence and progression of metabolic syndrome. J Clin Endocrinol Metab. 2014;99(12):4607–15.
pubmed: 25188715
Harte AL, da Silva NF, Miller MA, Cappuccio FP, Kelly A, O’Hare JP, et al. Telomere length attrition, a marker of biological senescence, is inversely correlated with triglycerides and cholesterol in South Asian males with type 2 diabetes mellitus. Exp Diabetes Res. 2012;2012:895185.
pubmed: 22474429 pmcid: 3303685
Weischer M, Bojesen SE, Cawthon RM, Freiberg JJ, Tybjaerg-Hansen A, Nordestgaard BG. Short telomere length, myocardial infarction, ischemic heart disease, and early death. Arterioscler Thromb Vasc Biol. 2012;32(3):822–9.
pubmed: 22199369
Masi S, Nightingale CM, Day IN, Guthrie P, Rumley A, Lowe GD, et al. Inflammation and not cardiovascular risk factors is associated with short leukocyte telomere length in 13- to 16-year-old adolescents. Arterioscler Thromb Vasc Biol. 2012;32(8):2029–34.
pubmed: 22679311
von Kanel R, Malan NT, Hamer M, van der Westhuizen FH, Malan L. Leukocyte telomere length and hemostatic factors in a South African cohort: the SABPA Study. J Thromb Haemost. 2014;12(12):1975–85.
Valentini E, Zampieri M, Malavolta M, Bacalini MG, Calabrese R, Guastafierro T, et al. Analysis of the machinery and intermediates of the 5hmC-mediated DNA demethylation pathway in aging on samples from the MARK-AGE Study. Aging. 2016;8(9):1896–922.
pubmed: 27587280 pmcid: 5076444
El-Maarri O, Kareta MS, Mikeska T, Becker T, Diaz-Lacava A, Junen J, et al. A systematic search for DNA methyltransferase polymorphisms reveals a rare DNMT3L variant associated with subtelomeric hypomethylation. Hum Mol Genet. 2009;18(10):1755–68.
pubmed: 19246518
Yang J, Guo R, Wang H, Ye X, Zhou Z, Dan J, et al. Tet enzymes regulate telomere maintenance and chromosomal stability of mouse ESCs. Cell Rep. 2016;15(8):1809–21.
pubmed: 27184841
Yehezkel S, Segev Y, Viegas-Pequignot E, Skorecki K, Selig S. Hypomethylation of subtelomeric regions in ICF syndrome is associated with abnormally short telomeres and enhanced transcription from telomeric regions. Hum Mol Genet. 2008;17(18):2776–89.
pubmed: 18558631
Lin Y, Damjanovic A, Metter EJ, Nguyen H, Truong T, Najarro K, et al. Age-associated telomere attrition of lymphocytes in vivo is co-ordinated with changes in telomerase activity, composition of lymphocyte subsets and health conditions. Clin Sci (Lond). 2015;128(6):367–77.
Valiathan R, Ashman M, Asthana D. Effects of ageing on the immune system: infants to elderly. Scand J Immunol. 2016;83(4):255–66.
pubmed: 26808160
Horvath S. DNA methylation age of human tissues and cell types. Genome Biol. 2013;14(10):R115.
pubmed: 24138928 pmcid: 4015143
Matsuyama M, WuWong DJ, Horvath S, Matsuyama S. Epigenetic clock analysis of human fibroblasts in vitro: effects of hypoxia, donor age, and expression of hTERT and SV40 largeT. Aging. 2019;11(10):3012–22.
pubmed: 31113906 pmcid: 6555444
Soraas A, Matsuyama M, de Lima M, Wald D, Buechner J, Gedde-Dahl T, et al. Epigenetic age is a cell-intrinsic property in transplanted human hematopoietic cells. Aging Cell. 2019;18(2):e12897.
pubmed: 30712319 pmcid: 6413751
Quach A, Levine ME, Tanaka T, Lu AT, Chen BH, Ferrucci L, et al. Epigenetic clock analysis of diet, exercise, education, and lifestyle factors. Aging. 2017;9(2):419–46.
pubmed: 28198702 pmcid: 5361673
Marioni RE, Harris SE, Shah S, McRae AF, von Zglinicki T, Martin-Ruiz C, et al. The epigenetic clock and telomere length are independently associated with chronological age and mortality. Int J Epidemiol. 2018;45(2):424–32.
pubmed: 27075770
Bucci L, Ostan R, Cevenini E, Pini E, Scurti M, Vitale G, et al. Centenarians’ offspring as a model of healthy aging: a reappraisal of the data on Italian subjects and a comprehensive overview. Aging. 2016;8(3):510–9.
pubmed: 26979133 pmcid: 4833142
Rozing MP, Westendorp RG, de Craen AJ, Frolich M, de Goeij MC, Heijmans BT, et al. Favorable glucose tolerance and lower prevalence of metabolic syndrome in offspring without diabetes mellitus of nonagenarian siblings: the Leiden longevity study. J Am Geriatr Soc. 2010;58(3):564–9.
pubmed: 20398121
Wijsman CA, Rozing MP, Streefland TC, le Cessie S, Mooijaart SP, Slagboom PE, et al. Familial longevity is marked by enhanced insulin sensitivity. Aging Cell. 2011;10(1):114–21.

Auteurs

Maria Giulia Bacalini (MG)

IRCCS Istituto delle Scienze Neurologiche di Bologna, Bologna, Italy.

Anna Reale (A)

Department of Experimental Medicine, Sapienza University of Rome, Viale Regina Elena 324, 00161, Rome, Italy.

Marco Malavolta (M)

Advanced Technology Center for Aging Research, IRCCS INRCA, 60121, Ancona, Italy.

Fabio Ciccarone (F)

IRCCS San Raffaele Pisana, Department of Human Sciences and Promotion of the Quality of Life, San Raffaele Roma Open University, 00166, Rome, Italy.

María Moreno-Villanueva (M)

Molecular Toxicology Group, Department of Biology, University of Konstanz, 78457, Konstanz, Germany.
Human Performance Research Centre, Department of Sport Science, University of Konstanz, 78457, Konstanz, Germany.

Martijn E T Dollé (MET)

Centre for Health Protection, National Institute for Public Health and the Environment, PO Box 1, 3720 BA, Bilthoven, The Netherlands.

Eugène Jansen (E)

Centre for Health Protection, National Institute for Public Health and the Environment, PO Box 1, 3720 BA, Bilthoven, The Netherlands.

Tilman Grune (T)

Department of Molecular Toxicology, German Institute of Human Nutrition Potsdam-Rehbruecke (DIfE), 14558, Nuthetal, Germany.
NutriAct-Competence Cluster Nutrition Research Berlin-Potsdam, 14458, Nuthetal, Germany.

Efstathios S Gonos (ES)

Institute of Biology, Medicinal Chemistry and Biotechnology, National Hellenic Research Foundation, Athens, Greece.

Christiane Schön (C)

BioTeSys GmbH, Schelztorstr. 54-56, 73728, Esslingen, Germany.

Jürgen Bernhardt (J)

BioTeSys GmbH, Schelztorstr. 54-56, 73728, Esslingen, Germany.

Beatrix Grubeck-Loebenstein (B)

Research Institute for Biomedical Aging Research, University of Innsbruck, Rennweg, 10, 6020, Innsbruck, Austria.

Ewa Sikora (E)

Laboratory of the Molecular Bases of Ageing, Nencki Institute of Experimental Biology, Polish Academy of Sciences, 3 Pasteur street, 02-093, Warsaw, Poland.

Olivier Toussaint (O)

URBC-NARILIS, University of Namur, Rue de Bruxelles, 61, Namur, Belgium.

Florence Debacq-Chainiaux (F)

URBC-NARILIS, University of Namur, Rue de Bruxelles, 61, Namur, Belgium.

Miriam Capri (M)

DIMES- Department of Experimental, Diagnostic and Specialty Medicine, Alma Mater Studiorum- University of Bologna, 40126, Bologna, Italy.

Antti Hervonen (A)

Faculty of Medicine and Health Technology, Tampere University, FIN-33014, Tampere, Finland.

Mikko Hurme (M)

Faculty of Medicine and Health Technology, Tampere University, FIN-33014, Tampere, Finland.

P Eline Slagboom (PE)

Department of Molecular Epidemiology, Leiden University Medical Centre, Leiden, The Netherlands.

Nicolle Breusing (N)

Department of Applied Nutritional Science/Dietetics, Institute of Nutritional Medicine, University of Hohenheim, 70599, Stuttgart, Germany.

Valentina Aversano (V)

Department of Experimental Medicine, Sapienza University of Rome, Viale Regina Elena 324, 00161, Rome, Italy.

Stefano Tagliatesta (S)

Department of Experimental Medicine, Sapienza University of Rome, Viale Regina Elena 324, 00161, Rome, Italy.

Claudio Franceschi (C)

Laboratory of Systems Medicine of Healthy Aging and Department of Applied Mathematics, Lobachevsky University, Nizhny Novgorod, Russia.

Maria A Blasco (MA)

Telomeres and Telomerase Group, Spanish National Cancer Research Centre (CNIO), Madrid, Spain.

Alexander Bürkle (A)

Molecular Toxicology Group, Department of Biology, University of Konstanz, 78457, Konstanz, Germany.

Paola Caiafa (P)

Department of Cellular Biotechnologies and Haematology, Sapienza University of Rome, Viale, Regina Elena 324, 00161, Rome, Italy. paola.caiafa44@gmail.com.

Michele Zampieri (M)

Department of Experimental Medicine, Sapienza University of Rome, Viale Regina Elena 324, 00161, Rome, Italy. michele.zampieri@uniroma1.it.

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