Within-sibship genome-wide association analyses decrease bias in estimates of direct genetic effects.


Journal

Nature genetics
ISSN: 1546-1718
Titre abrégé: Nat Genet
Pays: United States
ID NLM: 9216904

Informations de publication

Date de publication:
05 2022
Historique:
received: 05 03 2021
accepted: 25 03 2022
pubmed: 10 5 2022
medline: 20 5 2022
entrez: 9 5 2022
Statut: ppublish

Résumé

Estimates from genome-wide association studies (GWAS) of unrelated individuals capture effects of inherited variation (direct effects), demography (population stratification, assortative mating) and relatives (indirect genetic effects). Family-based GWAS designs can control for demographic and indirect genetic effects, but large-scale family datasets have been lacking. We combined data from 178,086 siblings from 19 cohorts to generate population (between-family) and within-sibship (within-family) GWAS estimates for 25 phenotypes. Within-sibship GWAS estimates were smaller than population estimates for height, educational attainment, age at first birth, number of children, cognitive ability, depressive symptoms and smoking. Some differences were observed in downstream SNP heritability, genetic correlations and Mendelian randomization analyses. For example, the within-sibship genetic correlation between educational attainment and body mass index attenuated towards zero. In contrast, analyses of most molecular phenotypes (for example, low-density lipoprotein-cholesterol) were generally consistent. We also found within-sibship evidence of polygenic adaptation on taller height. Here, we illustrate the importance of family-based GWAS data for phenotypes influenced by demographic and indirect genetic effects.

Identifiants

pubmed: 35534559
doi: 10.1038/s41588-022-01062-7
pii: 10.1038/s41588-022-01062-7
pmc: PMC9110300
doi:

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

581-592

Subventions

Organisme : Medical Research Council
ID : MC_UU_00011/6
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : R01 HL109946
Pays : United States
Organisme : Medical Research Council
ID : MR/T030852/1
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_UU_12026/2
Pays : United Kingdom
Organisme : Wellcome Trust
ID : 208806/Z/17/Z
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_UU_00017/1
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_U137686851
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_UU_00007/10
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_PC_13049
Pays : United Kingdom
Organisme : Wellcome Trust
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_PC_14135
Pays : United Kingdom
Organisme : NIA NIH HHS
ID : R01 AG042568
Pays : United States
Organisme : Medical Research Council
ID : MC_UU_00011/1
Pays : United Kingdom
Organisme : NIA NIH HHS
ID : R24 AG065184
Pays : United States
Organisme : Wellcome Trust
ID : 212946/Z/18/Z
Pays : United Kingdom
Organisme : Medical Research Council
ID : MR/R026408/1
Pays : United Kingdom
Organisme : NICHD NIH HHS
ID : P2C HD042849
Pays : United States
Organisme : NIA NIH HHS
ID : R56 AG042568
Pays : United States

Investigateurs

Hyeokmoon Kweon (H)
Philipp D Koellinger (PD)
Daniel J Benjamin (DJ)
Patrick Turley (P)
Laurence J Howe (LJ)
Michel G Nivard (MG)
Tim T Morris (TT)
Ailin F Hansen (AF)
Humaira Rasheed (H)
Yoonsu Cho (Y)
Geetha Chittoor (G)
Rafael Ahlskog (R)
Penelope A Lind (PA)
Teemu Palviainen (T)
Matthijs D van der Zee (MD)
Rosa Cheesman (R)
Massimo Mangino (M)
Yunzhang Wang (Y)
Shuai Li (S)
Lucija Klaric (L)
Scott M Ratliff (SM)
Lawrence F Bielak (LF)
Marianne Nygaard (M)
Alexandros Giannelis (A)
Emily A Willoughby (EA)
Chandra A Reynolds (CA)
Jared V Balbona (JV)
Ole A Andreassen (OA)
Helga Ask (H)
Dorret I Boomsma (DI)
Archie Campbell (A)
Harry Campbell (H)
Zhengming Chen (Z)
Paraskevi Christofidou (P)
Elizabeth Corfield (E)
Christina C Dahm (CC)
Deepika R Dokuru (DR)
Luke M Evans (LM)
Eco J C de Geus (EJC)
Sudheer Giddaluru (S)
Scott D Gordon (SD)
K Paige Harden (KP)
W David Hill (WD)
Amanda Hughes (A)
Shona M Kerr (SM)
Yongkang Kim (Y)
Antti Latvala (A)
Deborah A Lawlor (DA)
Liming Li (L)
Kuang Lin (K)
Per Magnus (P)
Patrik K E Magnusson (PKE)
Travis T Mallard (TT)
Pekka Martikainen (P)
Melinda C Mills (MC)
Pål Rasmus Njølstad (PR)
Nancy L Pedersen (NL)
David J Porteous (DJ)
Karri Silventoinen (K)
Melissa C Southey (MC)
Camilla Stoltenberg (C)
Elliot M Tucker-Drob (EM)
Margaret J Wright (MJ)
John K Hewitt (JK)
Matthew C Keller (MC)
Michael C Stallings (MC)
James J Lee (JJ)
Kaare Christensen (K)
Sharon L R Kardia (SLR)
Patricia A Peyser (PA)
Jennifer A Smith (JA)
James F Wilson (JF)
John L Hopper (JL)
Sara Hägg (S)
Tim D Spector (TD)
Jean-Baptiste Pingault (JB)
Robert Plomin (R)
Alexandra Havdahl (A)
Meike Bartels (M)
Nicholas G Martin (NG)
Sven Oskarsson (S)
Anne E Justice (AE)
Iona Y Millwood (IY)
Kristian Hveem (K)
Øyvind Naess (Ø)
Cristen J Willer (CJ)
Bjørn Olav Åsvold (BO)
Jaakko Kaprio (J)
Sarah E Medland (SE)
Robin G Walters (RG)
David M Evans (DM)
George Davey Smith (GD)
Caroline Hayward (C)
Ben Brumpton (B)
Gibran Hemani (G)
Neil M Davies (NM)

Informations de copyright

© 2022. The Author(s).

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Auteurs

Laurence J Howe (LJ)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK. laurence.howe@bristol.ac.uk.
Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK. laurence.howe@bristol.ac.uk.

Michel G Nivard (MG)

Department of Biological Psychology, Netherlands Twin Register, Vrije Universiteit, Amsterdam, the Netherlands.

Tim T Morris (TT)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK.
Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK.

Ailin F Hansen (AF)

K.G. Jebsen Center for Genetic Epidemiology, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Trondheim, Norway.

Humaira Rasheed (H)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK.
K.G. Jebsen Center for Genetic Epidemiology, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Trondheim, Norway.

Yoonsu Cho (Y)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK.
Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK.

Geetha Chittoor (G)

Department of Population Health Sciences, Geisinger Health, Danville, PA, USA.

Rafael Ahlskog (R)

Department of Government, Uppsala University, Uppsala, Sweden.

Penelope A Lind (PA)

Psychiatric Genetics, QIMR Berghofer Medical Research Institute, Brisbane, Australia.
School of Biomedical Sciences, Queensland University of Technology, Brisbane, Australia.
Faculty of Medicine, University of Queensland, Brisbane, Australia.

Teemu Palviainen (T)

Institute for Molecular Medicine FIMM, University of Helsinki, Helsinki, Finland.

Matthijs D van der Zee (MD)

Department of Biological Psychology, Netherlands Twin Register, Vrije Universiteit, Amsterdam, the Netherlands.

Rosa Cheesman (R)

PROMENTA Research Center, Department of Psychology, University of Oslo, Oslo, Norway.
Social Genetic & Developmental Psychiatry Centre, Institute of Psychiatry, Psychology & Neuroscience, King's College London, London, UK.

Massimo Mangino (M)

Department of Twin Research and Genetic Epidemiology, King's College London, London, UK.
NIHR Biomedical Research Centre at Guy's and St Thomas' Foundation Trust, London, UK.

Yunzhang Wang (Y)

Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden.

Shuai Li (S)

Centre for Epidemiology and Biostatistics, Melbourne School of Population and Global Health, The University of Melbourne, Parkville, Victoria, Australia.
Centre for Cancer Genetic Epidemiology, Department of Public Health and Primary Care, University of Cambridge, Cambridge, UK.
Precision Medicine, School of Clinical Sciences at Monash Health, Monash University, Clayton, Victoria, Australia.

Lucija Klaric (L)

MRC Human Genetics Unit, Institute of Genetics and Cancer, University of Edinburgh, Western General Hospital, Edinburgh, UK.

Scott M Ratliff (SM)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.

Lawrence F Bielak (LF)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.

Marianne Nygaard (M)

The Danish Twin Registry, Department of Public Health, University of Southern Denmark, Odense, Denmark.
Department of Clinical Genetics, Odense University Hospital, Odense, Denmark.

Alexandros Giannelis (A)

Department of Psychology, University of Minnesota, Minneapolis, MN, USA.

Emily A Willoughby (EA)

Department of Psychology, University of Minnesota, Minneapolis, MN, USA.

Chandra A Reynolds (CA)

Department of Psychology, University of California, Riverside, Riverside, CA, USA.

Jared V Balbona (JV)

Department of Psychology & Neuroscience, University of Colorado at Boulder, Boulder, CO, USA.
Institute for Behavioral Genetics, University of Colorado at Boulder, Boulder, CO, USA.

Ole A Andreassen (OA)

NORMENT Centre, University of Oslo, Oslo, Norway.
Division of Mental Health and Addiction, Oslo University Hospital, Oslo, Norway.

Helga Ask (H)

Department of Mental Disorders, Norwegian Institute of Public Health, Oslo, Norway.

Aris Baras (A)

Regeneron Genetics Center, Tarrytown, NY, USA.

Christopher R Bauer (CR)

BioMarin Pharmaceutical Inc., Novato, CA, USA.
Biomedical and Translational Informatics, Geisinger Health, Danville, PA, USA.

Dorret I Boomsma (DI)

Department of Biological Psychology, Netherlands Twin Register, Vrije Universiteit, Amsterdam, the Netherlands.
Amsterdam Public Health (APH) and Amsterdam Reproduction and Development (AR&D), Amsterdam, the Netherlands.

Archie Campbell (A)

Centre for Genomic and Experimental Medicine, Institute of Genetics & Cancer, University of Edinburgh, Western General Hospital, Edinburgh, UK.

Harry Campbell (H)

Centre for Global Health, Usher Institute, University of Edinburgh, Edinburgh, UK.

Zhengming Chen (Z)

Nuffield Department of Population Health, University of Oxford, Oxford, UK.
MRC Population Health Research Unit, University of Oxford, Oxford, UK.

Paraskevi Christofidou (P)

Department of Twin Research and Genetic Epidemiology, King's College London, London, UK.

Elizabeth Corfield (E)

Department of Mental Disorders, Norwegian Institute of Public Health, Oslo, Norway.
Nic Waals Institute, Lovisenberg Diaconal Hospital, Oslo, Norway.

Christina C Dahm (CC)

Department of Public Health, Aarhus University, Aarhus, Denmark.

Deepika R Dokuru (DR)

Department of Psychology & Neuroscience, University of Colorado at Boulder, Boulder, CO, USA.
Institute for Behavioral Genetics, University of Colorado at Boulder, Boulder, CO, USA.

Luke M Evans (LM)

Institute for Behavioral Genetics, University of Colorado at Boulder, Boulder, CO, USA.
Department of Ecology & Evolutionary Biology, University of Colorado at Boulder, Boulder, CO, USA.

Eco J C de Geus (EJC)

Department of Biological Psychology, Netherlands Twin Register, Vrije Universiteit, Amsterdam, the Netherlands.
Amsterdam Public Health Research Institute, Amsterdam UMC, Amsterdam, the Netherlands.

Sudheer Giddaluru (S)

Institute of Health and Society, University of Oslo, Oslo, Norway.
Norwegian Institute of Public Health, Oslo, Norway.

Scott D Gordon (SD)

Department of Genetics and Computational Biology, QIMR Berghofer Medical Research Institute, Brisbane, Queensland, Australia.

K Paige Harden (KP)

Department of Psychology and Population Research Center, University of Texas at Austin, Austin, TX, USA.

W David Hill (WD)

Lothian Birth Cohorts Group, Department of Psychology, University of Edinburgh, Edinburgh, UK.
Department of Psychology, University of Edinburgh, Edinburgh, UK.

Amanda Hughes (A)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK.
Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK.

Shona M Kerr (SM)

MRC Human Genetics Unit, Institute of Genetics and Cancer, University of Edinburgh, Western General Hospital, Edinburgh, UK.

Yongkang Kim (Y)

Institute for Behavioral Genetics, University of Colorado at Boulder, Boulder, CO, USA.

Hyeokmoon Kweon (H)

Department of Economics, School of Business and Economics, Vrije Universiteit Amsterdam, Amsterdam, the Netherlands.

Antti Latvala (A)

Institute for Molecular Medicine FIMM, University of Helsinki, Helsinki, Finland.
Institute of Criminology and Legal Policy, Faculty of Social Sciences, University of Helsinki, Helsinki, Finland.

Deborah A Lawlor (DA)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK.
Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK.
Bristol NIHR Biomedical Research Centre, Bristol, UK.

Liming Li (L)

Department of Epidemiology and Biostatistics, School of Public Health, Peking University Health Science Center, Beijing, China.

Kuang Lin (K)

Nuffield Department of Population Health, University of Oxford, Oxford, UK.

Per Magnus (P)

Centre for Fertility and Health, Norwegian Institute of Public Health, Skøyen, Oslo, Norway.

Patrik K E Magnusson (PKE)

Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden.

Travis T Mallard (TT)

Department of Psychology and Population Research Center, University of Texas at Austin, Austin, TX, USA.

Pekka Martikainen (P)

Population Research Unit, Faculty of Social Sciences, University of Helsinki, Helsinki, Finland.
The Max Planck Institute for Demographic Research, Rostock, Germany.
Department of Public Health Sciences, Stockholm University, Stockholm, Sweden.

Melinda C Mills (MC)

Leverhulme Centre for Demographic Science, University of Oxford, Oxford, UK.

Pål Rasmus Njølstad (PR)

Department of Clinical Science, University of Bergen, Bergen, Norway.
Children and Youth Clinic, Haukeland University Hospital, Bergen, Norway.

John D Overton (JD)

Regeneron Genetics Center, Tarrytown, NY, USA.

Nancy L Pedersen (NL)

Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden.

David J Porteous (DJ)

Centre for Genomic and Experimental Medicine, Institute of Genetics & Cancer, University of Edinburgh, Western General Hospital, Edinburgh, UK.

Jeffrey Reid (J)

Regeneron Genetics Center, Tarrytown, NY, USA.

Karri Silventoinen (K)

Population Research Unit, Faculty of Social Sciences, University of Helsinki, Helsinki, Finland.

Melissa C Southey (MC)

Precision Medicine, School of Clinical Sciences at Monash Health, Monash University, Clayton, Victoria, Australia.
Department of Clinical Pathology, Melbourne Medical School, The University of Melbourne, Melbourne, Victoria, Australia.
Cancer Epidemiology Division, Cancer Council Victoria, Melbourne, Victoria, Australia.

Camilla Stoltenberg (C)

Norwegian Institute of Public Health, Oslo, Norway.
Department of Global Public Health and Primary Care, University of Bergen, Bergen, Norway.

Elliot M Tucker-Drob (EM)

Department of Psychology and Population Research Center, University of Texas at Austin, Austin, TX, USA.

Margaret J Wright (MJ)

Queensland Brain Institute, The University of Queensland, Brisbane, Queensland, Australia.

John K Hewitt (JK)

Department of Psychology & Neuroscience, University of Colorado at Boulder, Boulder, CO, USA.
Institute for Behavioral Genetics, University of Colorado at Boulder, Boulder, CO, USA.

Matthew C Keller (MC)

Department of Psychology & Neuroscience, University of Colorado at Boulder, Boulder, CO, USA.
Institute for Behavioral Genetics, University of Colorado at Boulder, Boulder, CO, USA.

Michael C Stallings (MC)

Department of Psychology & Neuroscience, University of Colorado at Boulder, Boulder, CO, USA.
Institute for Behavioral Genetics, University of Colorado at Boulder, Boulder, CO, USA.

James J Lee (JJ)

Department of Psychology, University of Minnesota, Minneapolis, MN, USA.

Kaare Christensen (K)

The Danish Twin Registry, Department of Public Health, University of Southern Denmark, Odense, Denmark.
Department of Clinical Genetics, Odense University Hospital, Odense, Denmark.
Department of Clinical Biochemistry and Pharmacology, Odense University Hospital, Odense, Denmark.

Sharon L R Kardia (SLR)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.

Patricia A Peyser (PA)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.

Jennifer A Smith (JA)

Department of Epidemiology, School of Public Health, University of Michigan, Ann Arbor, MI, USA.
Survey Research Center, Institute for Social Research, University of Michigan, Ann Arbor, MI, USA.

James F Wilson (JF)

MRC Human Genetics Unit, Institute of Genetics and Cancer, University of Edinburgh, Western General Hospital, Edinburgh, UK.
Centre for Global Health, Usher Institute, University of Edinburgh, Edinburgh, UK.

John L Hopper (JL)

Centre for Epidemiology and Biostatistics, Melbourne School of Population and Global Health, The University of Melbourne, Parkville, Victoria, Australia.

Sara Hägg (S)

Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden.

Tim D Spector (TD)

Department of Twin Research and Genetic Epidemiology, King's College London, London, UK.

Jean-Baptiste Pingault (JB)

Social Genetic & Developmental Psychiatry Centre, Institute of Psychiatry, Psychology & Neuroscience, King's College London, London, UK.
Department of Clinical, Educational and Health Psychology, University College London, London, UK.

Robert Plomin (R)

Social Genetic & Developmental Psychiatry Centre, Institute of Psychiatry, Psychology & Neuroscience, King's College London, London, UK.

Alexandra Havdahl (A)

PROMENTA Research Center, Department of Psychology, University of Oslo, Oslo, Norway.
Department of Mental Disorders, Norwegian Institute of Public Health, Oslo, Norway.
Nic Waals Institute, Lovisenberg Diaconal Hospital, Oslo, Norway.

Meike Bartels (M)

Department of Biological Psychology, Netherlands Twin Register, Vrije Universiteit, Amsterdam, the Netherlands.

Nicholas G Martin (NG)

Department of Genetics and Computational Biology, QIMR Berghofer Medical Research Institute, Brisbane, Queensland, Australia.

Sven Oskarsson (S)

Department of Government, Uppsala University, Uppsala, Sweden.

Anne E Justice (AE)

Department of Population Health Sciences, Geisinger Health, Danville, PA, USA.

Iona Y Millwood (IY)

Nuffield Department of Population Health, University of Oxford, Oxford, UK.
MRC Population Health Research Unit, University of Oxford, Oxford, UK.

Kristian Hveem (K)

K.G. Jebsen Center for Genetic Epidemiology, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Trondheim, Norway.
HUNT Research Center, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Levanger, Norway.

Øyvind Naess (Ø)

Institute of Health and Society, University of Oslo, Oslo, Norway.
Norwegian Institute of Public Health, Oslo, Norway.

Cristen J Willer (CJ)

K.G. Jebsen Center for Genetic Epidemiology, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Trondheim, Norway.
Department of Internal Medicine: Cardiology, University of Michigan, Ann Arbor, MI, USA.
Department of Computational Medicine and Bioinformatics, University of Michigan, Ann Arbor, MI, USA.

Bjørn Olav Åsvold (BO)

K.G. Jebsen Center for Genetic Epidemiology, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Trondheim, Norway.
HUNT Research Center, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Levanger, Norway.
Department of Endocrinology, Clinic of Medicine, St. Olavs Hospital, Trondheim University Hospital, Trondheim, Norway.

Philipp D Koellinger (PD)

Department of Economics, School of Business and Economics, Vrije Universiteit Amsterdam, Amsterdam, the Netherlands.
La Follette School of Public Affairs, University of Wisconsin-Madison, Madison, WI, USA.

Jaakko Kaprio (J)

Institute for Molecular Medicine FIMM, University of Helsinki, Helsinki, Finland.

Sarah E Medland (SE)

Psychiatric Genetics, QIMR Berghofer Medical Research Institute, Brisbane, Australia.
Faculty of Medicine, University of Queensland, Brisbane, Australia.
School of Psychology, University of Queensland, Brisbane, Queensland, Australia.

Robin G Walters (RG)

Nuffield Department of Population Health, University of Oxford, Oxford, UK.
MRC Population Health Research Unit, University of Oxford, Oxford, UK.

Daniel J Benjamin (DJ)

UCLA Anderson School of Management, Los Angeles, CA, USA.
Human Genetics Department, UCLA David Geffen School of Medicine, Gonda (Goldschmied) Neuroscience and Genetics Research Center, Los Angeles, CA, USA.
National Bureau of Economic Research, Cambridge, MA, USA.

Patrick Turley (P)

Center for Economic and Social Research, University of Southern California, Los Angeles, CA, USA.
Department of Economics, University of Southern California, Los Angeles, CA, USA.

David M Evans (DM)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK.
University of Queensland Diamantina Institute, University of Queensland, Brisbane, Queensland, Australia.
Institute for Molecular Bioscience, University of Queensland, Brisbane, Queensland, Australia.

George Davey Smith (G)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK.
Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK.

Caroline Hayward (C)

MRC Human Genetics Unit, Institute of Genetics and Cancer, University of Edinburgh, Western General Hospital, Edinburgh, UK.

Ben Brumpton (B)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK. ben.brumpton@ntnu.no.
K.G. Jebsen Center for Genetic Epidemiology, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Trondheim, Norway. ben.brumpton@ntnu.no.
HUNT Research Center, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Levanger, Norway. ben.brumpton@ntnu.no.

Gibran Hemani (G)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK. g.hemani@bristol.ac.uk.
Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK. g.hemani@bristol.ac.uk.

Neil M Davies (NM)

Medical Research Council Integrative Epidemiology Unit at the University of Bristol, Bristol, UK. neil.davies@bristol.ac.uk.
Population Health Sciences, Bristol Medical School, University of Bristol, Bristol, UK. neil.davies@bristol.ac.uk.
K.G. Jebsen Center for Genetic Epidemiology, Department of Public Health and Nursing, NTNU, Norwegian University of Science and Technology, Trondheim, Norway. neil.davies@bristol.ac.uk.

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