Enhanced nucleosome assembly at CpG sites containing an extended 5-methylcytosine analogue.


Journal

Nucleic acids research
ISSN: 1362-4962
Titre abrégé: Nucleic Acids Res
Pays: England
ID NLM: 0411011

Informations de publication

Date de publication:
24 06 2022
Historique:
accepted: 16 05 2022
revised: 05 05 2022
received: 15 10 2021
pubmed: 2 6 2022
medline: 28 6 2022
entrez: 1 6 2022
Statut: ppublish

Résumé

Methylation of cytosine to 5-methylcytosine (mC) at CpG sites is a prevalent reversible epigenetic mark in vertebrates established by DNA methyltransferases (MTases); the attached methyl groups can alter local structure of DNA and chromatin as well as binding of dedicated proteins. Nucleosome assembly on methylated DNA has been studied extensively, however little is known how the chromatin structure is affected by larger chemical variations in the major groove of DNA. Here, we studied the nucleosome formation in vitro on DNA containing an extended 5mC analog, 5-(6-azidohex-2-ynyl)cytosine (ahyC) installed at biological relevant CpG sites. We found that multiple ahyC residues on 80-Widom and Hsp70 promoter DNA fragments proved compatible with nucleosome assembly. Moreover, unlike mC, ahyC increases the affinity of histones to the DNA, partially altering nucleosome positioning, stability, and the action of chromatin remodelers. Based on molecular dynamics calculations, we suggest that these new features are due to increased DNA flexibility at ahyC-modified sites. Our findings provide new insights into the biophysical behavior of modified DNA and open new ways for directed design of synthetic nucleosomes.

Identifiants

pubmed: 35648439
pii: 6596862
doi: 10.1093/nar/gkac444
pmc: PMC9226530
doi:

Substances chimiques

Chromatin 0
Nucleosomes 0
5-Methylcytosine 6R795CQT4H
Cytosine 8J337D1HZY
DNA 9007-49-2

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

6549-6561

Informations de copyright

© The Author(s) 2022. Published by Oxford University Press on behalf of Nucleic Acids Research.

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Auteurs

Miglė Tomkuvienė (M)

Institute of Biotechnology, Life Sciences Center, Vilnius University, Vilnius LT-10257, Lithuania.

Markus Meier (M)

Biochemistry III, University of Regensburg, Regensburg, Bavaria, DE-93053, Germany.

Diana Ikasalaitė (D)

Institute of Biotechnology, Life Sciences Center, Vilnius University, Vilnius LT-10257, Lithuania.

Julia Wildenauer (J)

Biochemistry III, University of Regensburg, Regensburg, Bavaria, DE-93053, Germany.

Visvaldas Kairys (V)

Institute of Biotechnology, Life Sciences Center, Vilnius University, Vilnius LT-10257, Lithuania.

Saulius Klimašauskas (S)

Institute of Biotechnology, Life Sciences Center, Vilnius University, Vilnius LT-10257, Lithuania.

Laura Manelytė (L)

Biochemistry III, University of Regensburg, Regensburg, Bavaria, DE-93053, Germany.

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Classifications MeSH