Genetic insights into resting heart rate and its role in cardiovascular disease.


Journal

Nature communications
ISSN: 2041-1723
Titre abrégé: Nat Commun
Pays: England
ID NLM: 101528555

Informations de publication

Date de publication:
02 08 2023
Historique:
received: 06 06 2022
accepted: 16 06 2023
medline: 4 8 2023
pubmed: 3 8 2023
entrez: 2 8 2023
Statut: epublish

Résumé

Resting heart rate is associated with cardiovascular diseases and mortality in observational and Mendelian randomization studies. The aims of this study are to extend the number of resting heart rate associated genetic variants and to obtain further insights in resting heart rate biology and its clinical consequences. A genome-wide meta-analysis of 100 studies in up to 835,465 individuals reveals 493 independent genetic variants in 352 loci, including 68 genetic variants outside previously identified resting heart rate associated loci. We prioritize 670 genes and in silico annotations point to their enrichment in cardiomyocytes and provide insights in their ECG signature. Two-sample Mendelian randomization analyses indicate that higher genetically predicted resting heart rate increases risk of dilated cardiomyopathy, but decreases risk of developing atrial fibrillation, ischemic stroke, and cardio-embolic stroke. We do not find evidence for a linear or non-linear genetic association between resting heart rate and all-cause mortality in contrast to our previous Mendelian randomization study. Systematic alteration of key differences between the current and previous Mendelian randomization study indicates that the most likely cause of the discrepancy between these studies arises from false positive findings in previous one-sample MR analyses caused by weak-instrument bias at lower P-value thresholds. The results extend our understanding of resting heart rate biology and give additional insights in its role in cardiovascular disease development.

Identifiants

pubmed: 37532724
doi: 10.1038/s41467-023-39521-2
pii: 10.1038/s41467-023-39521-2
pmc: PMC10397318
doi:

Types de publication

Meta-Analysis Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

4646

Subventions

Organisme : Medical Research Council
ID : MC_UU_00032/1
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_PC_19009
Pays : United Kingdom
Organisme : Wellcome Trust
ID : 202802/Z/16/Z
Pays : United Kingdom
Organisme : Medical Research Council
ID : MR/R010676/1
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_UU_00002/7
Pays : United Kingdom
Organisme : Wellcome Trust
Pays : United Kingdom
Organisme : Wellcome Trust
ID : 100114
Pays : United Kingdom
Organisme : Wellcome Trust
ID : 217065/Z/19/Z
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_UU_00006/1
Pays : United Kingdom
Organisme : Wellcome Trust
ID : 212945/Z/18/Z
Pays : United Kingdom
Organisme : NIDDK NIH HHS
ID : R01 DK072193
Pays : United States
Organisme : Medical Research Council
ID : MC_UU_00011/1
Pays : United Kingdom
Organisme : Cancer Research UK
ID : C18281/A29019
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : R01 HL105756
Pays : United States
Organisme : Medical Research Council
ID : MC_PC_15018
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_PC_U127592696
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_UU_00007/10
Pays : United Kingdom
Organisme : Medical Research Council
ID : G9815508
Pays : United Kingdom
Organisme : Wellcome Trust
ID : 204623/Z/16/Z
Pays : United Kingdom

Investigateurs

Delnaz Roshandel (D)
Andrew D Paterson (AD)

Informations de copyright

© 2023. The Author(s).

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Auteurs

Yordi J van de Vegte (YJ)

Department of Cardiology, University of Groningen, University Medical Center Groningen, Groningen, 9700RB, the Netherlands.

Ruben N Eppinga (RN)

Department of Cardiology, Isala Zwolle ziekenhuis, Zwolle, 8025 AB, the Netherlands.

M Yldau van der Ende (MY)

Department of Cardiology, University medical Center Utrecht, Utrecht, 3584 Cx, the Netherlands.

Yanick P Hagemeijer (YP)

Department of Cardiology, University of Groningen, University Medical Center Groningen, Groningen, 9700RB, the Netherlands.
Analytical Biochemistry, University of Groningen, Groningen, 9713 AV, the Netherlands.

Yuvaraj Mahendran (Y)

The Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medicine Science, University of Copenhagen, Copenhagen Ø, 2100, Denmark.

Elias Salfati (E)

Department of Medicine, Stanford University School of Medicine, Stanford, 94305, USA.
Faculty of Medicine, University of Iceland, Reykjavik, 101, Iceland.

Albert V Smith (AV)

Department of Biostatistics, University of Michigan, Ann Arbor, MI48109, USA.

Vanessa Y Tan (VY)

Bristol Medical School, Population Health Sciences, University of Bristol, Bristol, BS82BN, UK.
MRC Integrative Epidemiology, University of Bristol, Bristol, BS82BN, UK.

Dan E Arking (DE)

McKusick-Nathans Institute, Department of Genetic Medicine, Johns Hopkins University School of Medicine, Baltimore, 21215, USA.

Ioanna Ntalla (I)

Clinical Pharmacology and Precision Medicine, William Harvey Research Institute, Barts and The London Faculty of Medicine and Dentistry, Queen Mary University of London, London, EC1M 6BQ, UK.

Emil V Appel (EV)

The Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medicine Science, University of Copenhagen, Copenhagen Ø, 2100, Denmark.

Claudia Schurmann (C)

The Charles Bronfman Institute for Personalized Medicine, The Icahn School of Medicine at Mount Sinai, New York, 10029, USA.

Jennifer A Brody (JA)

Medicine, University of Washington, Seattle, 98195, USA.

Rico Rueedi (R)

Department of Computational Biology, University of Lausanne, Lausanne, 1015, Switzerland.
Swiss Institute of Bioinformatics, Lausanne, 1015, Switzerland.

Ozren Polasek (O)

Department of Public Health, University of Split School of Medicine, Split, 21000, Croatia.
Algebra LAB, Algebra University College, Zagreb, 10000, Croatia.

Gardar Sveinbjornsson (G)

deCODE genetics / Amgen Inc., Reykjavik, 102, Iceland.

Cecile Lecoeur (C)

UMR 8199, University of Lille Nord de France, Lille, 59000, France.

Claes Ladenvall (C)

Clinial Genomics Uppsala, Department of Immunology, Genetics and Pathology, Science for Life Laboratory, Uppsala University, Uppsala, 75185, Sweden.
Lund University Diabetes Center, Department of Clinical Sciences, Lund University, Malmö, 20502, Sweden.

Jing Hua Zhao (JH)

BHF Cardiovascular Epidemiology Unit, Department of Public Health and Primary Care, Victor Phillip Dahdaleh Heart & Lung Research Institute, University of Cambridge, Cambridge, CB2 0BB, UK.

Aaron Isaacs (A)

CARIM School for Cardiovascular Diseases, Maastricht Centre for Systems Biology (MaCSBio), Department of Physiology, Maastricht University, Maastricht, 6229ER, Netherlands.

Lihua Wang (L)

Division of Statistical Genomics, Department of Genetics, Washington University School of Medicine, St. Louis, MO, 63108-2212, Campus Box 8506, USA.

Jian'an Luan (J)

MRC Epidemiology Unit, University of Cambridge School of Clinical Medicine, Cambridge, CB2 0QQ, UK.

Shih-Jen Hwang (SJ)

Division of Intramural Research, National Heart Lung and Blood Institute, NIH, USA, Framingham, 1702, USA.

Nina Mononen (N)

Department of Clinical Chemistry, Fimlab Laboratories, Tampere, FI-33014, Finland.
Department of Clinical Chemistry, Finnish Cardiovascular Research Center - Tampere, Faculty of Medicine and Health Technology, Tampere University, Tampere, FI-33014, Finland.

Kirsi Auro (K)

Department of Health, unit of genetics and biomarkers, , National Institute for Health and Welfare, Finland, Helsinki, FI-00290, Finland.
Department of molecular medicine, University of Helsinki, Helsinki, FI-00290, Finland.

Anne U Jackson (AU)

Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, MI, 48109, USA.

Lawrence F Bielak (LF)

Department of Epidemiology, University of Michigan, Ann Arbor, MI, 48109, USA.

Linyao Zeng (L)

Department of Cardiology, Deutsches Herzzentrum München, Technische Universität München, Munich, 80636, Germany.

Nabi Shah (N)

Division of Population Health and Genomics, School of Medicine, University of Dundee, Dundee, DD1 9SY, UK.
Pharmacogenetics Research Lab, Department of Pharmacy, COMSATS University Islamabad, Abbottabad, 22060, Pakistan.

Maria Nethander (M)

Sahlgrenska Osteoporosis Centre, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, 41345, Sweden.
Bioinformatics Core Facility, Sahlgrenska Academy, University of Gothenburg, Gothenburg, 40530, Sweden.

Archie Campbell (A)

Centre for Genomic and Experimental Medicine, Institute of Genetics & Cancer, University of Edinburgh, Edinburgh, EH4 2XU, UK.
Usher Institute for Population Health Sciences and Informatics, The University of Edinburgh, Edinburgh, EH16 4UX, UK.

Tuomo Rankinen (T)

Human Genomics Laboratory, Pennington Biomedical Research Center, Baton Rouge, LA, 70808, USA.

Sonali Pechlivanis (S)

Institute for Medical Informatics, Biometry and Epidemiology, University Hospital of Essen, University Duisburg-Essen, Essen, 45122, Germany.

Lu Qi (L)

Department of Epidemiology, Tulane University, New Orleans, LA, 70112, USA.

Wei Zhao (W)

Department of Epidemiology, University of Michigan, Ann Arbor, MI, 48109, USA.
Survey Research Center, Institute for Social Research, University of Michigan, Ann Arbor, MI, 48104, USA.

Federica Rizzi (F)

Unit of Biomedicine, Bio4Dreams-Business Nursery for Life Sciences, Milano, 20121, Italy.

Toshiko Tanaka (T)

Longitudinal Study Section, National Institute on Aging, Baltimore, 21224, USA.

Antonietta Robino (A)

Institute for Maternal and Child Health - IRCCS "Burlo Garofolo", Trieste, 34137, Italy.

Massimiliano Cocca (M)

Institute for Maternal and Child Health - IRCCS "Burlo Garofolo", Trieste, 34137, Italy.

Leslie Lange (L)

Medicine, University of Colorado Anschutz Medical Campus, Aurora, 80045, USA.

Martina Müller-Nurasyid (M)

IBE, Ludwig-Maximilians-University Munich, LMU Munich, Munich, 81377, Germany.
Institute of Medical Biostatistics, Epidemiology and Informatics (IMBEI), University Medical Center, Johannes Gutenberg University, Mainz, 55101, Germany.
Institute of Genetic Epidemiology, Helmholtz Zentrum München - German Research Center for Environmental Health, Neuherberg, 85764, Germany.

Carolina Roselli (C)

Institute of Genetic Epidemiology, Helmholtz Zentrum München - German Research Center for Environmental Health, Neuherberg, 85764, Germany.
Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, 02142, USA.

Weihua Zhang (W)

Department of Epidemiology and Biostatistics, Imperial College London, London, W2 1PG, UK.
Department of Cardiology, Ealing Hospital, London North West University Healthcare NHS Trust, Middlesex, UB1 3HW, UK.

Marcus E Kleber (ME)

Vth Department of Medicine (Nephrology, Hypertensiology, Rheumatology, Endocrinology, Diabetology), Medical Faculty Mannheim, University of Heidelberg, Mannheim, 68167, Germany.
SYNLAB MVZ Humangenetik Mannheim, Mannheim, 68163, Germany.

Xiuqing Guo (X)

Pediatrics, The Institute for Translational Genomics and Population Sciences, The Lundquist Institute for Biomedical Innovation at Harbor-UCLA, Torrance, 90502, USA.
Department of Pediatrics, Harbor-UCLA Medical Center, Torrance, 90502, USA.

Henry J Lin (HJ)

Pediatrics, The Institute for Translational Genomics and Population Sciences, The Lundquist Institute for Biomedical Innovation at Harbor-UCLA, Torrance, 90502, USA.
Department of Pediatrics, Harbor-UCLA Medical Center, Torrance, 90502, USA.

Francesca Pavani (F)

Institute for Biomedicine, Eurac Research, Bolzano, 39100, Italy.

Tessel E Galesloot (TE)

Radboud university medical center, Nijmegen, 6500 HB, the Netherlands.

Raymond Noordam (R)

Department of Internal Medicine, section Gerontology and Geriatrics, Leiden University Medical Center, Leiden, 2300 RC, the Netherlands.

Yuri Milaneschi (Y)

Department of Psychiatry, Amsterdam Public Health, Amsterdam UMC, Amsterdam UMC, Vrije Universiteit, Amsterdam, Amsterdam, 1081 HL, the Netherlands.

Katharina E Schraut (KE)

Centre for Cardiovascular Science, Queen's Medical Research Institute, University of Edinburgh, Edinburgh, EH16 4TJ, Scotland, UK.

Marcel den Hoed (M)

The Beijer laboratory and Department of Immunology, Genetics and Pathology, Uppsala University and Science for Life Laboratory, Uppsala, 75237, Sweden.

Frauke Degenhardt (F)

Institute of Clinical Molecular Biology, Christian-Albrechts-University of Kiel, Kiel, 24105, Germany.

Stella Trompet (S)

Department of Internal Medicine, section Gerontology and Geriatrics, Leiden University Medical Center, Leiden, 2300 RC, the Netherlands.
Department of Cardiology, Leiden University Medical Center, Leiden, ZA, 2333, the Netherlands.

Marten E van den Berg (ME)

Department of Epidemiology, Erasmus Medical Center, Rotterdam, 3015GD, the Netherlands.

Giorgio Pistis (G)

Institute of Genetics and Biomedic Research (IRGB), Italian National Research Council (CNR), Monserrato, (CA), 9042, Italy.
Center for Statistical Genetics, University of Michigan, Ann Arbor, 48109, USA.

Yih-Chung Tham (YC)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, 169856, Singapore.

Stefan Weiss (S)

Interfaculty Institute for Genetics and Functional Genomics, University Medicine Greifswald, Greifswald, 17475, Germany.
DZHK (German Center for Cardiovascular Research), Partner Site Greifswald, Greifswald, 17475, Germany.

Xueling S Sim (XS)

Saw Swee Hock School of Public Health, National University Health System and National University of Singapore, Singapore, 117549, Singapore.

Hengtong L Li (HL)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, 169856, Singapore.

Peter J van der Most (PJ)

Department of Epidemiology, University of Groningen, University Medical Center Groningen, Groningen, 9700 RB, The Netherlands.

Ilja M Nolte (IM)

Department of Epidemiology, University of Groningen, University Medical Center Groningen, Groningen, 9700 RB, The Netherlands.

Leo-Pekka Lyytikäinen (LP)

Department of Clinical Chemistry, Fimlab Laboratories, Tampere, FI-33014, Finland.
Department of Clinical Chemistry, Finnish Cardiovascular Research Center - Tampere, Faculty of Medicine and Health Technology, Tampere University, Tampere, FI-33014, Finland.
Cardiovascular Epidemiology Unit, Department of Public Health and Primary Care, University of Cambridge, Cambridge, CB2 0SL, UK.

M Abdullah Said (MA)

Department of Cardiology, University of Groningen, University Medical Center Groningen, Groningen, 9700RB, the Netherlands.

Daniel R Witte (DR)

Department of Public Health, Aarhus University, Aarhus C, 8000, Denmark.

Carlos Iribarren (C)

Division of Research, Kaiser Permenente of Northern California, Oakland, 94612, USA.
The Scripps Research Institute, La Jolla, 10550, USA.

Lenore Launer (L)

National Institute on Aging, Bethesda, MSC2292, USA.

Susan M Ring (SM)

Bristol Medical School, Population Health Sciences, University of Bristol, Bristol, BS82BN, UK.
MRC Integrative Epidemiology, University of Bristol, Bristol, BS82BN, UK.

Paul S de Vries (PS)

Department of Epidemiology, Human Genetics, and Environmental Sciences, University of Texas Health Science Center at Houston, School of Public Health, Houston, 77030, USA.

Peter Sever (P)

National Heart and Lung Institute, Imperial College London, London, W12 0NN, UK.

Allan Linneberg (A)

Center for Clinical Research and Prevention, Bispebjerg and Frederiksberg Hospital, Copenhagen, 2400, Denmark.
Department of Clinical Medicine, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, 2200, Denmark.

Erwin P Bottinger (EP)

The Charles Bronfman Institute for Personalized Medicine, The Icahn School of Medicine at Mount Sinai, New York, 10029, USA.
Department of Preventive Medicine, The Icahn School of Medicine at Mount Sinai, New York, 10029, USA.

Sandosh Padmanabhan (S)

Institute of Cardiovascular and Medical Sciences, University of Glasgow, Glasgow, G12 8TA, UK.

Bruce M Psaty (BM)

Departments of Medicine, Epidemiology and Health Systems and Population Health, University of Washington, Seattle, 98195, USA.

Nona Sotoodehnia (N)

Medicine and Epidemiology, University of Washington, Seattle, 98195, USA.

Ivana Kolcic (I)

Department of Public Health, University of Split School of Medicine, Split, 21000, Croatia.
Algebra LAB, Algebra University College, Zagreb, 10000, Croatia.

David O Arnar (DO)

deCODE genetics / Amgen Inc., Reykjavik, 102, Iceland.
Faculty of Medicine, School of Health Sciences, University of Iceland, Reykjavik, 101, Iceland.
Department of Medicine, Landspitali-The National University Hospital of Iceland, Reykjavik, 101, Iceland.

Daniel F Gudbjartsson (DF)

deCODE genetics / Amgen Inc., Reykjavik, 102, Iceland.
School of Engineering and Natural Sciences, University of Iceland, Reykjavik, 101, Iceland.

Hilma Holm (H)

deCODE genetics / Amgen Inc., Reykjavik, 102, Iceland.

Beverley Balkau (B)

Centre for Research in Epidemiology and Population Health, Institut national de la santé et de la recherche médicale, Villejuif, 94800, France.
UMRS 1018, University Versailles Saint-Quentin-en-Yvelines, Versailles, 78035, France.
UMRS 1018, University Paris Sud, Villejuif, 94807, France.

Claudia T Silva (CT)

Genetic Epidemiology Unit, Dept. of Epidemiology, Erasmus University Medical Center, Rotterdam, 3000CA, Netherlands.

Christopher H Newton-Cheh (CH)

Cardiology Section, Massachusetts general hospital,, Boston, 02130, USA.

Kjell Nikus (K)

Department of Cardiology, Heart Center, Tampere University Hospital, Tampere, FI-33521, Finland.
Department of Cardiology, Finnish Cardiovascular Research Center - Tampere, Faculty of Medicine and Health Technology, Tampere University, Tampere, FI-33014, Finland.

Perttu Salo (P)

Department of Health, unit of genetics and biomarkers, , National Institute for Health and Welfare, Finland, Helsinki, FI-00290, Finland.
Department of molecular medicine, University of Helsinki, Helsinki, FI-00290, Finland.

Karen L Mohlke (KL)

Department of Genetics, University of North Carolina, Chapel Hill, NC, 27599, USA.

Patricia A Peyser (PA)

Department of Epidemiology, University of Michigan, Ann Arbor, MI, 48109, USA.

Heribert Schunkert (H)

Department of Cardiology, Deutsches Herzzentrum München, Technische Universität München, Munich, 80636, Germany.
Deutsches Zentrum für Herz- und Kreislauferkrankungen (DZHK), Partner Site Munich Heart Alliance, Munich, 80636, Germany.

Mattias Lorentzon (M)

Sahlgrenska Osteoporosis Centre, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, 41345, Sweden.
Region Västra Götaland, Geriatric Medicine, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Mölndal, 43180, Sweden.
Mary McKillop Institute for Health Research, Australian Catholic University, Melbourne, 3000, Australia.

Jari Lahti (J)

Department of Psychology and Logopedics, University of Helsinki, Helsinki, 00014, Finland.

Dabeeru C Rao (DC)

Division of Biostatistics, Washington University, St. Louis, MO, 63110, USA.

Marilyn C Cornelis (MC)

Preventive Medicine, Northwestern University, Chicago, IL, 60611, USA.

Jessica D Faul (JD)

Survey Research Center, Institute for Social Research, University of Michigan, Ann Arbor, MI, 48104, USA.

Jennifer A Smith (JA)

Department of Epidemiology, University of Michigan, Ann Arbor, MI, 48109, USA.
Survey Research Center, Institute for Social Research, University of Michigan, Ann Arbor, MI, 48104, USA.

Katarzyna Stolarz-Skrzypek (K)

Department of Cardiology, Interventional Electrocardiology and Hypertension, Jagiellonian University Medical College, Kraków, 31-008, Poland.

Stefania Bandinelli (S)

Geriatric Unit, Unità sanitaria locale Toscana Centro, Florence, 50142, Italy.

Maria Pina Concas (MP)

Institute for Maternal and Child Health - IRCCS "Burlo Garofolo", Trieste, 34137, Italy.

Gianfranco Sinagra (G)

Cardiovascular Department, "Ospedali Riuniti and University of Trieste", Trieste, 34149, Italy.

Thomas Meitinger (T)

Institute of Human Genetics, Klinikum rechts der Isar, Technische Universität München, München, 81675, Germany.
Institute of Human Genetics, Helmholtz Zentrum München - German Research Center for Environmental Health, Neuherberg, 85764, Germany.

Melanie Waldenberger (M)

Research Unit Molecular Epidemiology, Helmholtz Zentrum München - German Research Center for Environmental Health, Neuherberg, 85764, Germany.
Institute of Epidemiology, Helmholtz Zentrum München - German Research Center for Environmental Health, Neuherberg, 85764, Germany.
German Centre for Cardiovascular Research (DZHK), partner site: Munich Heart Alliance, Munich, 80802, Germany.

Moritz F Sinner (MF)

German Centre for Cardiovascular Research (DZHK), partner site: Munich Heart Alliance, Munich, 80802, Germany.
Department of Cardiology, University Hospital, LMU Munich, Munich, 81377, Germany.

Konstantin Strauch (K)

Institute of Medical Biostatistics, Epidemiology and Informatics (IMBEI), University Medical Center, Johannes Gutenberg University, Mainz, 55101, Germany.
Institute of Genetic Epidemiology, Helmholtz Zentrum München - German Research Center for Environmental Health, Neuherberg, 85764, Germany.
Chair of Genetic Epidemiology, IBE, Faculty of Medicine, LMU Munich, Munich, 81377, Germany.

Graciela E Delgado (GE)

Vth Department of Medicine (Nephrology, Hypertensiology, Rheumatology, Endocrinology, Diabetology), Medical Faculty Mannheim, University of Heidelberg, Mannheim, 68167, Germany.

Kent D Taylor (KD)

Pediatrics, The Institute for Translational Genomics and Population Sciences, The Lundquist Institute for Biomedical Innovation at Harbor-UCLA, Torrance, 90502, USA.
Department of Pediatrics, Harbor-UCLA Medical Center, Torrance, 90502, USA.

Jie Yao (J)

Pediatrics, The Institute for Translational Genomics and Population Sciences, The Lundquist Institute for Biomedical Innovation at Harbor-UCLA, Torrance, 90502, USA.
Department of Pediatrics, Harbor-UCLA Medical Center, Torrance, 90502, USA.

Luisa Foco (L)

Institute for Biomedicine, Eurac Research, Bolzano, 39100, Italy.

Olle Melander (O)

Department of Internal Medicine, Clinical Sciences, Lund University and Skåne University Hospital, Malmo, 221 85, Sweden.
Lund University Diabetes Center, Lund University, Malmö, 221 85, Sweden.

Jacqueline de Graaf (J)

Radboud university medical center, Nijmegen, 6500 HB, the Netherlands.

Renée de Mutsert (R)

Department of Clinical Epidemiology, Leiden University Medical Center, Leiden, 2300 RC, the Netherlands.

Eco J C de Geus (EJC)

Biological Psychology, EMGO+ Institute for Health and Care Research and Neuroscience Campus Amsterdam, VU University, Amsterdam, 1081 BT, the Netherlands.

Åsa Johansson (Å)

Department of Immunology, Genetics and Pathology, Uppsala University, Uppsala, 75108, Sweden.

Peter K Joshi (PK)

Centre for Global Health Research, Usher Institute, University of Edinburgh, Edinburgh, EH8 9AG, Scotland, UK.

Lars Lind (L)

Department of Medical Sciences, Cardiovascular Epidemiology, Uppsala University Hospital, Uppsala, 75237, Sweden.

Andre Franke (A)

Institute of Clinical Molecular Biology, Christian-Albrechts-University of Kiel, Kiel, 24105, Germany.

Peter W Macfarlane (PW)

Institute of Health and Wellbeing, Faculty of Medicine, University of Glasgow, Glasgow, G12 0XH, UK.

Kirill V Tarasov (KV)

Laboratory of Cardiovascular Sciences, Intramural Research Program, National Institute on Aging, National Institutes of Health, Baltimore, MD, 21224, USA.

Nicholas Tan (N)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, 169856, Singapore.

Stephan B Felix (SB)

DZHK (German Center for Cardiovascular Research), Partner Site Greifswald, Greifswald, 17475, Germany.
Department of Internal Medicine B, University Medicine Greifswald, Greifswald, 17475, Germany.

E-Shyong Tai (ES)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, 169856, Singapore.
Department of Medicine, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, 119228, Singapore.

Debra Q Quek (DQ)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, 169856, Singapore.

Harold Snieder (H)

Department of Epidemiology, University of Groningen, University Medical Center Groningen, Groningen, 9700 RB, The Netherlands.

Johan Ormel (J)

Department of Psychiatry, University of Groningen, University Medical Center Groningen, Groningen, 9700 RB, The Netherlands.

Martin Ingelsson (M)

Department of Public Health and Caring Sciences, Molecular Geriatrics, Uppsala University, Uppsala, 75237, Sweden.

Cecilia Lindgren (C)

Genetic and Genomic Epidemiology Unit, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, OX3 7BN, UK.

Andrew P Morris (AP)

Genetic and Genomic Epidemiology Unit, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, OX3 7BN, UK.

Olli T Raitakari (OT)

Centre for Population Health Research, University of Turku and Turku University Hospital, Turku, FI-20521, Finland.
Research Centre of Applied and Preventive Cardiovascular Medicine, University of Turku, Turku, FI-20521, Finland.
Department of Clinical Physiology and Nuclear Medicine, Turku University Hospital, Turku, FI-20521, Finland.

Torben Hansen (T)

The Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medicine Science, University of Copenhagen, Copenhagen Ø, 2100, Denmark.

Themistocles Assimes (T)

Department of Medicine, Stanford University School of Medicine, Stanford, 94305, USA.

Vilmundur Gudnason (V)

Faculty of Medicine, University of Iceland, Reykjavik, 101, Iceland.
Icelandic Heart Association, Kopavogur, 201, Iceland.

Nicholas J Timpson (NJ)

MRC Integrative Epidemiology Unit, University of Bristol, Bristol, BS8 2BN, UK.
Population Health Sciences, Bristol Medical School,, University of Bristol, Bristol, BS8 2BN, UK.

Alanna C Morrison (AC)

Department of Epidemiology, Human Genetics, and Environmental Sciences, University of Texas Health Science Center at Houston, School of Public Health, Houston, 77030, USA.

Patricia B Munroe (PB)

Clinical Pharmacology and Precision Medicine, William Harvey Research Institute, Barts and The London Faculty of Medicine and Dentistry, Queen Mary University of London, London, EC1M 6BQ, UK.
NIHR Barts Biomedical Research Centre, Barts and The London Faculty of Medicine and Dentistry, Queen Mary University of London, London, EC1M 6BQ, UK.

David P Strachan (DP)

Population Health Research Institute, St George's, University of London, London, SW17 0RE, UK.

Niels Grarup (N)

The Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medicine Science, University of Copenhagen, Copenhagen Ø, 2100, Denmark.

Ruth J F Loos (RJF)

The Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medicine Science, University of Copenhagen, Copenhagen Ø, 2100, Denmark.
The Charles Bronfman Institute for Personalized Medicine, The Icahn School of Medicine at Mount Sinai, New York, 10029, USA.
The Mindich Child Health and Development Institute, The Icahn School of Medicine at Mount Sinai, New York, 10029, USA.

Susan R Heckbert (SR)

Department of Epidemiology, University of Washington, Seattle, 98195, USA.

Peter Vollenweider (P)

Department of Medicine, Internal Medicine, Lausanne University hospital, Lausanne, 1015, Switzerland.

Caroline Hayward (C)

MRC Human Genetics Unit, Institute of Genetics and Cancer, University of Edinburgh, Edinburgh, EH4 2XU, Scotland, UK.

Kari Stefansson (K)

deCODE genetics / Amgen Inc., Reykjavik, 102, Iceland.
Faculty of Medicine, School of Health Sciences, University of Iceland, Reykjavik, 101, Iceland.

Philippe Froguel (P)

Department of Metabolism, Imperial College London, London, W12 0HS, UK.
Inserm/CNRS UMR 1283/8199, Pasteur Institute of Lille, Lille University Hospital, EGID, Lille, 59000, France.
University of Lille, Lille, 59000, France.

Leif Groop (L)

Lund University Diabetes Center, Department of Clinical Sciences, Lund University, Malmö, 20502, Sweden.
Institute for Molecular Medicine Finland (FIMM), University of Helsinki, Helsinki, 00290, Finland.

Nicholas J Wareham (NJ)

MRC Epidemiology Unit, University of Cambridge School of Clinical Medicine, Cambridge, CB2 0QQ, UK.

Cornelia M van Duijn (CM)

Genetic Epidemiology Unit, Dept. of Epidemiology, Erasmus University Medical Center, Rotterdam, 3000CA, Netherlands.

Mary F Feitosa (MF)

Division of Statistical Genomics, Department of Genetics, Washington University School of Medicine, St. Louis, MO, 63108-2212, Campus Box 8506, USA.

Christopher J O'Donnell (CJ)

Cardiology Section, VA Boston Healthcare System, Harvard Medical School, Boston, MA, 02132, USA.

Mika Kähönen (M)

Department of Clinical Physiology, Tampere University Hospital, Tampere, FI-33521, Finland.
Department of Clinical Physiology, Finnish Cardiovascular Research Center - Tampere, Faculty of Medicine and Health Technology, Tampere University, Tampere, FI-33521, Finland.

Markus Perola (M)

Department of Health, unit of genetics and biomarkers, , National Institute for Health and Welfare, Finland, Helsinki, FI-00290, Finland.
Department of molecular medicine, University of Helsinki, Helsinki, FI-00290, Finland.

Michael Boehnke (M)

Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, MI, 48109, USA.

Sharon L R Kardia (SLR)

Department of Epidemiology, University of Michigan, Ann Arbor, MI, 48109, USA.

Jeanette Erdmann (J)

Institute for Cardiogenetics, University of Lübeck, Lübeck, 23562, Germany.

Colin N A Palmer (CNA)

Division of Population Health and Genomics, School of Medicine, University of Dundee, Dundee, DD1 9SY, UK.

Claes Ohlsson (C)

Sahlgrenska Osteoporosis Centre, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, 41345, Sweden.
Department of Drug Treatment, Sahlgrenska University Hospital, Gothenburg, 41345, Sweden.

David J Porteous (DJ)

Centre for Genomic and Experimental Medicine, Institute of Genetics & Molecular Medicine, University of Edinburgh, Edinburgh, EH4 2XU, UK.

Johan G Eriksson (JG)

Department of General practice and primary care, University of Helsinki, Helsinki, 00014, Finland.
Department of Obstetrics and Gynecology, National University of Singapore, Singapore, 119228, Singapore.
Public health Research Program, Folkhalsan Research Center, Helsinki, 000250, Finland.

Claude Bouchard (C)

Human Genomics Laboratory, Pennington Biomedical Research Center, Baton Rouge, LA, 70808, USA.

Susanne Moebus (S)

Institute for Medical Informatics, Biometry and Epidemiology, University Hospital of Essen, University Duisburg-Essen, Essen, 45122, Germany.
Centre for Urban Epidemiology, University Hospital of Essen, University Duisburg-Essen, Essen, 45122, Germany.

Peter Kraft (P)

Epidemiology, Harvard T.H. Chan School of Public Health, Boston, MA, 02112, USA.

David R Weir (DR)

Survey Research Center, Institute for Social Research, University of Michigan, Ann Arbor, MI, 48104, USA.

Daniele Cusi (D)

Unit of Biomedicine, Bio4Dreams-Business Nursery for Life Sciences, Milano, 20121, Italy.
Institute of Biomedical Technologies, National Research Council of Italy, Segrate, (MI), 20090, Italy.

Luigi Ferrucci (L)

Longitudinal Study Section, National Institute on Aging, Baltimore, 21224, USA.

Sheila Ulivi (S)

Institute for Maternal and Child Health - IRCCS "Burlo Garofolo", Trieste, 34137, Italy.

Giorgia Girotto (G)

Institute for Maternal and Child Health - IRCCS "Burlo Garofolo", Trieste, 34137, Italy.
Department of Medicine, Surgery and Health Sciences, University of Trieste, Trieste, 34149, Italy.

Adolfo Correa (A)

Jackson Heart Study, University of Mississippi Medical Center, Jackson, 39216, USA.

Stefan Kääb (S)

German Centre for Cardiovascular Research (DZHK), partner site: Munich Heart Alliance, Munich, 80802, Germany.
Department of Cardiology, University Hospital, LMU Munich, Munich, 81377, Germany.

Annette Peters (A)

Institute of Epidemiology, Helmholtz Zentrum München - German Research Center for Environmental Health, Neuherberg, 85764, Germany.
German Centre for Cardiovascular Research (DZHK), partner site: Munich Heart Alliance, Munich, 80802, Germany.
Chair of Epidemiology, Institute for Medical Information Processing, Biometry and Epidemiology, Ludwig-Maximilians-Universität München, Munich, 81377, Germany.

John C Chambers (JC)

Department of Epidemiology and Biostatistics, Imperial College London, London, W2 1PG, UK.
Department of Cardiology, Ealing Hospital, London North West University Healthcare NHS Trust, Middlesex, UB1 3HW, UK.
Lee Kong Chian School of Medicine, Nanyang Technological University, Singapore, 308232, Singapore.

Jaspal S Kooner (JS)

Department of Cardiology, Ealing Hospital, London North West University Healthcare NHS Trust, Middlesex, UB1 3HW, UK.
National Heart and Lung Institute, Imperial College London, London, W12 0NN, UK.
Imperial College Healthcare NHS Trust, Imperial College London, London, W12 0HS, UK.

Winfried März (W)

Vth Department of Medicine (Nephrology, Hypertensiology, Rheumatology, Endocrinology, Diabetology), Medical Faculty Mannheim, University of Heidelberg, Mannheim, 68167, Germany.
Synlab Academy, Synlab Holding Deutschland GmbH, Mannheim, 68161, Germany.

Jerome I Rotter (JI)

Pediatrics, The Institute for Translational Genomics and Population Sciences, The Lundquist Institute for Biomedical Innovation at Harbor-UCLA, Torrance, 90502, USA.
Department of Pediatrics, Harbor-UCLA Medical Center, Torrance, 90502, USA.

Andrew A Hicks (AA)

Institute for Biomedicine, Eurac Research, Bolzano, 39100, Italy.

J Gustav Smith (JG)

Department of Cardiology, Clinical Sciences, Lund University and Skåne University Hospital, Lund, 221 85, Sweden.
Wallenberg Center for Molecular Medicine and Lund University Diabetes Center, Lund University, Lund, 221 84, Sweden.
The Wallenberg Laboratory/Department of Molecular and Clinical Medicine, Institute of Medicine, Gothenburg University and the Department of Cardiology, Sahlgrenska University Hospital, Gothenburg, 413 45, Sweden.

Lambertus A L M Kiemeney (LALM)

Radboud university medical center, Nijmegen, 6500 HB, the Netherlands.

Dennis O Mook-Kanamori (DO)

Department of Clinical Epidemiology, Leiden University Medical Center, Leiden, 2300 RC, the Netherlands.
Department of Public Health and Primary Care, Leiden University Medical Center, Leiden, 2300 RC, the Netherlands.

Brenda W J H Penninx (BWJH)

Department of Psychiatry, Amsterdam Public Health, Amsterdam UMC, Amsterdam UMC, Vrije Universiteit, Amsterdam, Amsterdam, 1081 HL, the Netherlands.

Ulf Gyllensten (U)

Department of Immunology, Genetics and Pathology, Uppsala University, Uppsala, 75108, Sweden.

James F Wilson (JF)

Centre for Global Health Research, Usher Institute, University of Edinburgh, Edinburgh, EH8 9AG, Scotland, UK.
MRC Human Genetics Unit, Institute of Genetics and Cancer, University of Edinburgh, Edinburgh, EH4 2XU, Scotland, UK.

Stephen Burgess (S)

MRC Biostatistics Unit, University of Cambridge, Cambridge, CB2 0SR, UK.

Johan Sundström (J)

Department of Medical Sciences, Cardiovascular Epidemiology, Uppsala University Hospital, Uppsala, 75237, Sweden.

Wolfgang Lieb (W)

Institute of Epidemiology and Biobank PopGen, Kiel University, Kiel, 24105, Germany.

J Wouter Jukema (JW)

Department of Cardiology, Leiden University Medical Center, Leiden, ZA, 2333, the Netherlands.
Einthoven Laboratory for Experimental Vascular Medicine, Leiden University Medical Center, Leiden, ZA, 2333, the Netherlands.
Netherlands Heart Institute, Utrecht, 3511 EP, the Netherlands.

Mark Eijgelsheim (M)

Department of Epidemiology, Erasmus Medical Center, Rotterdam, 3015GD, the Netherlands.
Department of Nephrology, University Medical Center Groningen, Groningen, 9700RB, the Netherlands.

Edward L M Lakatta (ELM)

Laboratory of Cardiovascular Sciences, Intramural Research Program, National Institute on Aging, National Institutes of Health, Baltimore, MD, 21224, USA.

Ching-Yu Cheng (CY)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, 169856, Singapore.
Ophthalmology & Visual Sciences Academic Clinical Program (Eye ACP), Duke-NUS Medical School, Singapore, 169857, Singapore.
Department of Ophthalmology, Yong Loo Lin School of Medicine, National University of Singapore, Singapore, 119228, Singapore.

Marcus Dörr (M)

DZHK (German Center for Cardiovascular Research), Partner Site Greifswald, Greifswald, 17475, Germany.
Department of Internal Medicine B, University Medicine Greifswald, Greifswald, 17475, Germany.

Tien-Yin Wong (TY)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, 169856, Singapore.
Ophthalmology & Visual Sciences Academic Clinical Program (Eye ACP), Duke-NUS Medical School, Singapore, 169857, Singapore.
Tsinghua Medicine, Tsinghua University, Beijing, 100084, China.

Charumathi Sabanayagam (C)

Singapore Eye Research Institute, Singapore National Eye Centre, Singapore, 169856, Singapore.
Ophthalmology & Visual Sciences Academic Clinical Program (Eye ACP), Duke-NUS Medical School, Singapore, 169857, Singapore.

Albertine J Oldehinkel (AJ)

Interdisciplinary Center Psychopathology and Emotion Regulation, University of Groningen, University Medical Center Groningen, Groningen, 9700 RB, The Netherlands.

Harriette Riese (H)

Department of Psychiatry, University of Groningen, University Medical Center Groningen, Groningen, 9700 RB, The Netherlands.

Terho Lehtimäki (T)

Department of Clinical Chemistry, Fimlab Laboratories, Tampere, FI-33014, Finland.
Department of Clinical Chemistry, Finnish Cardiovascular Research Center - Tampere, Faculty of Medicine and Health Technology, Tampere University, Tampere, FI-33014, Finland.

Niek Verweij (N)

Department of Cardiology, University of Groningen, University Medical Center Groningen, Groningen, 9700RB, the Netherlands.

Pim van der Harst (P)

Department of Cardiology, University of Groningen, University Medical Center Groningen, Groningen, 9700RB, the Netherlands. P.vanderHarst@umcutrecht.nl.
Department of Cardiology, University medical Center Utrecht, Utrecht, 3584 Cx, the Netherlands. P.vanderHarst@umcutrecht.nl.
Department of Genetics, University of Groningen, University Medical Center Groningen, Groningen, 9700RB, the Netherlands. P.vanderHarst@umcutrecht.nl.

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