Effects of weaning on intestinal longitudinal muscle-myenteric plexus function in piglets.

diarrhea longitudinal muscle-myenteric plexus piglets weaning

Journal

Science China. Life sciences
ISSN: 1869-1889
Titre abrégé: Sci China Life Sci
Pays: China
ID NLM: 101529880

Informations de publication

Date de publication:
09 Oct 2023
Historique:
received: 01 04 2023
accepted: 18 05 2023
medline: 12 10 2023
pubmed: 12 10 2023
entrez: 12 10 2023
Statut: aheadofprint

Résumé

Weaning piglets usually suffer from severe diarrhea (commonly known as postweaning diarrhea, PWD) along with intestinal motility disorder. Intestinal peristalsis is mainly regulated by the longitudinal muscle-myenteric plexus (LM-MP). To understand the relationship between intestinal LM-MP function and the development of PWD, we compared the intestinal electrical activity, and the transcriptional profile of the LM-MP between 21-day-old piglets (just weaned, n=7) and 24-day-old piglets (suffered the most severe weaning stress, n=7). The results showed that 24-day-old piglets exhibited different degrees of diarrhea. A significant increase in the slow-wave frequency in the ileum and colon was observed in 24-day-old piglets, while c-kit expression in the intestinal LM-MPs was significantly decreased, indicating that PWD caused by elevated slow-wave frequency may be associated with loss of c-kit. The real-time quantitative PCR (RT-qPCR) and enzyme-linked immunosorbent assay (ELISA) showed that intestinal LM-MPs in 24-day-old piglets may undergo inflammation and oxidative stress. Significant increases in 8-hydroxy-2'-deoxyguanosine and decreases in thioredoxin suggest that weaning may lead to DNA damage in the LM-MP of 24-day-old piglets. In addition, activating transcription factor 3 was significantly upregulated, indicating nerve damage in the LM-MP of 24-day-old piglets. The transcriptomic results showed that most of the differentially expressed genes in the ileal LM-MP after weaning were downregulated and closely related to the cell cycle process. Subsequent RT-qPCR analysis showed that the relative expression of p21 was upregulated, while the expression of cyclin A2, cyclin B1, and proliferating cell nuclear antigen was downregulated in the ileal and colonic LM-MP of 24-day-old piglets, suggesting that weaning may inhibit cell proliferation and cause G1/S cell cycle arrest in ileal and colonic LM-MP. In conclusion, weaning may lead to cell cycle arrest by causing DNA damage in the LM-MP, impairing intestinal motility regulation, and ultimately leading to diarrhea in piglets.

Identifiants

pubmed: 37824029
doi: 10.1007/s11427-022-2391-x
pii: 10.1007/s11427-022-2391-x
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Informations de copyright

© 2023. Science China Press.

Références

Aiello, P., Sharghi, M., Mansourkhani, S.M., Ardekan, A.P., Jouybari, L., Daraei, N., Peiro, K., Mohamadian, S., Rezaei, M., Heidari, M., et al. (2019). Medicinal plants in the prevention and treatment of colon cancer. Oxid Med Cell Longev 2019, 1–51.
doi: 10.1155/2019/2075614
Balk, R.S., Silva, M.H., Bridi, J.C., Carvalho, N.R., Portella, R.L., Dobrachinski, F., Amaral, G.P., Barcelos, R., Dias, G.R.M., Rocha, J.B.T., et al. (2011). Effect of repeated restraint stress and clomipramine on Na
pubmed: 21762772 doi: 10.1016/j.ijdevneu.2011.06.010
Brun, P., Qesari, M., Marconi, P.C., Kotsafti, A., Porzionato, A., Macchi, V., Schwendener, R.A., Scarpa, M., Giron, M.C., Palù, G., et al. (2018). Herpes simplex virus type 1 infects enteric neurons and triggers gut dysfunction via macrophage recruitment. Front Cell Infect Microbiol 8, 74.
pubmed: 29600197 pmcid: 5862801 doi: 10.3389/fcimb.2018.00074
Chen, X., Meng, X., Zhang, H., Feng, C., Wang, B., Li, N., Abdullahi, K.M., Wu, X., Yang, J., Li, Z., et al. (2020). Intestinal proinflammatory macrophages induce a phenotypic switch in interstitial cells of Cajal. J Clin Invest 130, 6443–6456.
pubmed: 32809970 pmcid: 7685750 doi: 10.1172/JCI126584
Corbillé, A.G., Coron, E., Neunlist, M., Derkinderen, P., and Lebouvier, T. (2014). Appraisal of the dopaminergic and noradrenergic innervation of the submucosal plexus in PD. J Parkinsons Dis 4, 571–576.
pubmed: 25055959 doi: 10.3233/JPD-140422
Cortese, K., Daga, A., Monticone, M., Tavella, S., Stefanelli, A., Aiello, C., Bisio, A., Bellese, G., and Castagnola, P. (2016). Carnosic acid induces proteasomal degradation of Cyclin B1, RB and SOX2 along with cell growth arrest and apoptosis in GBM cells. Phytomedicine 23, 679–685.
pubmed: 27235706 doi: 10.1016/j.phymed.2016.03.007
Demedts, I., Masaoka, T., Kindt, S., Hertogh, G.D., Geboes, K., Farré, R., Berghe, P.V., and Tack, J. (2013). Gastrointestinal motility changes and myenteric plexus alterations in spontaneously diabetic biobreeding rats. J Neurogastroenterol Motil 19, 161–170.
pubmed: 23667747 pmcid: 3644652 doi: 10.5056/jnm.2013.19.2.161
Drissi, R., Chauvin, A., McKenna, A., Lévesque, D., Blais-Brochu, S., Jean, D., and Boisvert, F.M. (2018). Destabilization of the MiniChromosome Maintenance (MCM) complex modulates the cellular response to DNA double strand breaks. Cell Cycle 17, 2593–2609.
pubmed: 30516086 pmcid: 6300108 doi: 10.1080/15384101.2018.1553336
Farra, R., Dapas, B., Pozzato, G., Scaggiante, B., Agostini, F., Zennaro, C., Grassi, M., Rosso, N., Giansante, C., Fiotti, N., et al. (2011). Effects of E2F1-cyclin E1-E2 circuit down regulation in hepatocellular carcinoma cells. Dig Liver Dis 43, 1006–1014.
pubmed: 21831731 doi: 10.1016/j.dld.2011.07.007
Faussone-Pellegrini, M.S. (1985). Cytodifferentiation of the interstitial cells of Cajal related to the myenteric plexus of mouse intestinal muscle coat. Anat Embryol 171, 163–169.
doi: 10.1007/BF00341410
Feng, X., Noguchi, Y., Barbon, M., Stillman, B., Speck, C., and Li, H. (2021). The structure of ORC-Cdc6 on an origin DNA reveals the mechanism of ORC activation by the replication initiator Cdc6. Nat Commun 12, 3883.
pubmed: 34162887 pmcid: 8222357 doi: 10.1038/s41467-021-24199-1
Fintl, C., Lindberg, R., and McL. Press, C. (2020). Myenteric networks of interstitial cells of Cajal are reduced in horses with inflammatory bowel disease. Equine Vet J 52, 298–304.
pubmed: 31397916 doi: 10.1111/evj.13160
Foong, D., Zhou, J., Zarrouk, A., Ho, V., and O’Connor, M.D. (2020). Understanding the biology of human interstitial cells of Cajal in gastrointestinal motility. Int J Mol Sci 21, 4540.
pubmed: 32630607 pmcid: 7352366 doi: 10.3390/ijms21124540
Fung, C., and Vanden Berghe, P. (2020). Functional circuits and signal processing in the enteric nervous system. Cell Mol Life Sci 77, 4505–4522.
pubmed: 32424438 pmcid: 7599184 doi: 10.1007/s00018-020-03543-6
Gao, J., Yang, Z., Zhao, C., Tang, X., Jiang, Q., and Yin, Y. (2023). A comprehensive review on natural phenolic compounds as alternatives to in-feed antibiotics. Sci China Life Sci 66, 1518–1534.
pubmed: 36586071 doi: 10.1007/s11427-022-2246-4
Gao, Y., Tang, Y., Zhang, H., Chu, X., Yan, B., Li, J., and Liu, C. (2021). Vincristine leads to colonic myenteric neurons injury via pro-inflammatory macrophages activation. Biochem Pharmacol 186, 114479.
pubmed: 33617842 doi: 10.1016/j.bcp.2021.114479
Gehen, S.C., Vitiello, P.F., Bambara, R.A., Keng, P.C., and O’Reilly, M.A. (2007). Downregulation of PCNA potentiates p21-mediated growth inhibition in response to hyperoxia. Am J Physiol Lung Cell Mol Physiol 292, L716–L724.
pubmed: 17085526 doi: 10.1152/ajplung.00135.2006
Gfroerer, S., and Rolle, U. (2013). Interstitial cells of Cajal in the normal human gut and in Hirschsprung disease. Pediatr Surg Int 29, 889–897.
pubmed: 23917331 doi: 10.1007/s00383-013-3364-y
Gilchrist, M., Henderson Jr, W.R., Morotti, A., Johnson, C.D., Nachman, A., Schmitz, F., Smith, K.D., and Aderem, A. (2010). A key role for ATF3 in regulating mast cell survival and mediator release. Blood 115, 4734–4741.
pubmed: 20203264 pmcid: 2890168 doi: 10.1182/blood-2009-03-213512
Gresse, R., Chaucheyras-Durand, F., Fleury, M.A., Van de Wiele, T., Forano, E., and Blanquet-Diot, S. (2017). Gut microbiota dysbiosis in postweaning piglets: understanding the keys to health. Trends Microbiol 25, 851–873.
pubmed: 28602521 doi: 10.1016/j.tim.2017.05.004
Hedemann, M.S., Hojsgaard, S., and Jensen, B.B. (2003). Small intestinal morphology and activity of intestinal peptidases in piglets around weaning. J Anim Physiol Anim Nutr 87, 32–41.
doi: 10.1046/j.1439-0396.2003.00405.x
Higgens, C.S., and Allan, R.N. (1980). Crohn’s disease of the distal ileum. Gut 21, 933–940.
pubmed: 7450558 pmcid: 1419288 doi: 10.1136/gut.21.11.933
Holland, A.M., Bon-Frauches, A.C., Keszthelyi, D., Melotte, V., and Boesmans, W. (2021). The enteric nervous system in gastrointestinal disease etiology. Cell Mol Life Sci 78, 4713–4733.
pubmed: 33770200 pmcid: 8195951 doi: 10.1007/s00018-021-03812-y
Huang, Z., Liao, L., Wang, Z., Lu, Y., Yan, W., Cao, H., and Tan, B. (2021). An efficient approach for wholemount preparation of the myenteric plexus of rat colon. J Neurosci Methods 348, 109012.
pubmed: 33249181 doi: 10.1016/j.jneumeth.2020.109012
Huizinga, J.D., and Lammers, W.J.E.P. (2009). Gut peristalsis is governed by a multitude of cooperating mechanisms. Am J Physiol Gastrointest Liver Physiol 296, G1–G8.
pubmed: 18988693 doi: 10.1152/ajpgi.90380.2008
Iino, S., Horiguchi, K., and Horiguchi, S. (2020). c-Kit-stem cell factor signal-independent development of interstitial cells of Cajal in murine small intestine. Cell Tissue Res 379, 121–129.
pubmed: 31741038 doi: 10.1007/s00441-019-03120-9
Islam, O., Gong, X., Rose-John, S., and Heese, K. (2009). Interleukin-6 and neural stem cells: more than gliogenesis. Mol Biol Cell 20, 188–199.
pubmed: 18971377 pmcid: 2613125 doi: 10.1091/mbc.e08-05-0463
James, C.G., Woods, A., Underhill, T.M., and Beier, F. (2006). The transcription factor ATF3 is upregulated during chondrocyte differentiation and represses cyclin D1 and A gene transcription. BMC Mol Biol 7, 30.
pubmed: 16984628 pmcid: 1584246 doi: 10.1186/1471-2199-7-30
Jin, Q.H., Shen, H.X., Wang, H., Shou, Q.Y., and Liu, Q. (2013). Curcumin improves expression of SCF/c-kit through attenuating oxidative stress and NF-κB activation in gastric tissues of diabetic gastroparesis rats. Diabetol Metab Syndr 5, 12.
pubmed: 23448582 pmcid: 3630009 doi: 10.1186/1758-5996-5-12
Lalles, J.P., Bosi, P., Smidt, H., and Stokes, C.R. (2007). Nutritional management of gut health in pigs around weaning. Proc Nutr Soc 66, 260–268.
pubmed: 17466106 doi: 10.1017/S0029665107005484
Lin, J.H., Yu, Y.W., Chuang, Y.C., Lee, C.H., and Chen, C.C. (2021). ATF3-expressing large-diameter sensory afferents at acute stage as bio-signatures of persistent pain associated with lumbar radiculopathy. Cells 10, 992.
pubmed: 33922541 pmcid: 8145235 doi: 10.3390/cells10050992
Liu, J., Zhang, Y., Li, Y., Yan, H., and Zhang, H. (2019). L-tryptophan enhances intestinal integrity in diquat-challenged piglets associated with improvement of redox status and mitochondrial function. Animals 9, 266.
pubmed: 31121956 pmcid: 6562546 doi: 10.3390/ani9050266
Lohberger, B., Leithner, A., Stuendl, N., Kaltenegger, H., Kullich, W., and Steinecker-Frohnwieser, B. (2015). Diacerein retards cell growth of chondrosarcoma cells at the G2/M cell cycle checkpoint via cyclin B1/CDK1 and CDK2 downregulation. BMC Cancer 15, 1.
doi: 10.1186/s12885-015-1915-4
Love, M.I., Huber, W., and Anders, S. (2014). Moderated estimation of fold change and dispersion for RNA-seq data with DESeq2. Genome Biol 15, 550.
pubmed: 25516281 pmcid: 4302049 doi: 10.1186/s13059-014-0550-8
Lu, J., and Holmgren, A. (2014). The thioredoxin antioxidant system. Free Radic Biol Med 66, 75–87.
pubmed: 23899494 doi: 10.1016/j.freeradbiomed.2013.07.036
Machado, C.C.A., Watanabe, P.S., Mendes, J.D.L., Pupim, A.C.E., Ortigoza, S.M., Bergoc, H.G., Nino, B.S.L., Gois, M.B., Garcia, J.L., Blackshaw, L.A., et al. (2021). Toxoplasma gondii infection impairs the colonic motility of rats due to loss of myenteric neurons. Neurogastroenterol Motil 33.
Mansilla, S.F., De La Vega, M.B., Calzetta, N.L., Siri, S.O., and Gottifredi, V. (2020). CDK-independent and PCNA-dependent functions of p21 in DNA replication. Genes 11, 593.
pubmed: 32481484 pmcid: 7349641 doi: 10.3390/genes11060593
Medland, J.E., Pohl, C.S., Edwards, L.L., Frandsen, S., Bagley, K., Li, Y., and Moeser, A. J. (2016). Early life adversity in piglets induces long-term upregulation of the enteric cholinergic nervous system and heightened, sex-specific secretomotor neuron responses. Neurogastroenterol Motil 28, 1317–1329.
pubmed: 27134125 pmcid: 5002263 doi: 10.1111/nmo.12828
Nag, S., Qin, J., Srivenugopal, K.S., Wang, M., and Zhang, R. (2013). The MDM2-p53 pathway revisited. J Biomed Res 27, 254.
pubmed: 23885265 pmcid: 3721034 doi: 10.7555/JBR.27.20130030
Nordberg, J., and Arner, E.S.J. (2001). Reactive oxygen species, antioxidants, and the mammalian thioredoxin system. Free Radic Biol Med 31, 1287–1312.
pubmed: 11728801 doi: 10.1016/S0891-5849(01)00724-9
Pai, J.T., Hsu, M.W., Leu, Y.L., Chang, K.T., and Weng, M.S. (2021). Induction of G2/ M cell cycle arrest via p38/p21Waf1/Cip1-dependent signaling pathway activation by bavachinin in non-small-cell lung cancer cells. Molecules 26, 5161.
pubmed: 34500594 pmcid: 8434044 doi: 10.3390/molecules26175161
Park, J., Jang, H., Hwang, S.S., Kim, E.J., Kim, D.E., Oh, K.B., Kwon, D.J., Koh, J.T., Kimura, K., Inoue, H., et al. (2014). ER stress-inducible ATF3 suppresses BMP2- induced ALP expression and activation in MC3T3-E1 cells. Biochem Biophys Res Commun 443, 333–338.
pubmed: 24315873 doi: 10.1016/j.bbrc.2013.11.121
Parkhomchuk, D., Borodina, T., Amstislavskiy, V., Banaru, M., Hallen, L., Krobitsch, S., Lehrach, H., and Soldatov, A. (2009). Transcriptome analysis by strand-specific sequencing of complementary DNA. Nucleic Acids Res 37, e123.
pubmed: 19620212 pmcid: 2764448 doi: 10.1093/nar/gkp596
Pasternak, A., Szura, M., Gil, K., and Matyja, A. (2016). Interstitial cells of Cajal — systematic review. Folia Morphol 75, 281–286.
doi: 10.5603/FM.a2016.0002
Qureshi, W.A. (2002). Gastrointestinal uses of botulinum toxin. J Clin Gastroenterol 34, 126–128.
pubmed: 11782604 doi: 10.1097/00004836-200202000-00004
Ran, C., Li, Y., Ma, X., Xie, Y., Xie, M., Zhang, Y., Zhou, W., Yang, Y., Zhang, Z., Zhou, L., et al. (2021). Interactions between commensal bacteria and viral infection: insights for viral disease control in farmed animals. Sci China Life Sci 64, 1437–1448.
pubmed: 33420920 doi: 10.1007/s11427-020-1721-5
Ren, W., Yu, B., Yu, J., Zheng, P., Huang, Z., Luo, J., Mao, X., He, J., Yan, H., Wu, J., et al. (2022). Lower abundance of Bacteroides and metabolic dysfunction are highly associated with the post-weaning diarrhea in piglets. Sci China Life Sci 65, 2062–2075.
pubmed: 35467318 doi: 10.1007/s11427-021-2068-6
Song, L., Martinez, L., Zigmond, Z.M., Hernandez, D.R., Lassance-Soares, R.M., Selman, G., and Vazquez-Padron, R.I. (2017). c-Kit modifies the inflammatory status of smooth muscle cells. PeerJ 5, e3418.
pubmed: 28626608 pmcid: 5472039 doi: 10.7717/peerj.3418
Spencer, N.J., and Hu, H. (2020). Enteric nervous system: sensory transduction, neural circuits and gastrointestinal motility. Nat Rev Gastroenterol Hepatol 17, 338–351.
pubmed: 32152479 pmcid: 7474470 doi: 10.1038/s41575-020-0271-2
Sun, X., Cui, Y., Su, Y., Gao, Z., Diao, X., Li, J., Zhu, X., Li, D., Li, Z., Wang, C., et al. (2021). Dietary fiber ameliorates lipopolysaccharide-induced intestinal barrier function damage in piglets by modulation of intestinal microbiome. mSystems 6, e01374–20.
pubmed: 33824201 pmcid: 8547013 doi: 10.1128/mSystems.01374-20
Tilg, H., Trehu, E., Atkins, M.B., Dinarello, C.A., and Mier, J.W. (1994). Interleukin-6 (IL-6) as an anti-inflammatory cytokine: induction of circulating IL-1 receptor antagonist and soluble tumor necrosis factor receptor p55. Blood 83, 113–118.
pubmed: 8274730 doi: 10.1182/blood.V83.1.113.113
Valavanidis, A., Vlachogianni, T., and Fiotakis, C. (2009). 8-Hydroxy-2’-deoxyguanosine (8-OHdG): a critical biomarker of oxidative stress and carcinogenesis. J Environ Sci Health Part C 27, 120–139.
doi: 10.1080/10590500902885684
Vicentini, G.E., Martins, H.A., Fracaro, L., de Souza, S.R.G., da Silva Zanoni, K.P., Silva, T.N.X., Blegniski, F.P., Guarnier, F.A., and Zanoni, J.N. (2017). Does l-glutamine-supplemented diet extenuate NO-mediated damage on myenteric plexus of Walker 256 tumor-bearing rats? Food Res Int 101, 24–34.
pubmed: 28941690 doi: 10.1016/j.foodres.2017.08.054
Vitek, L. (2015). Bile acid malabsorption in inflammatory bowel disease. Inflammatory Bowel Dis 21, 476–483.
doi: 10.1097/MIB.0000000000000193
Wei, J., Li, N., Xia, X., Chen, X., Peng, F., Besner, G.E., and Feng, J. (2014). Effects of lipopolysaccharide-induced inflammation on the interstitial cells of Cajal. Cell Tissue Res 356, 29–37.
pubmed: 24435644 doi: 10.1007/s00441-013-1775-7
Wu, C.M., Yang, C.W., Lee, Y.Z., Chuang, T.H., Wu, P.L., Chao, Y.S., and Lee, S.J. (2009). Tylophorine arrests carcinoma cells at G1 phase by downregulating cyclin A2 expression. Biochem Biophys Res Commun 386, 140–145.
pubmed: 19501048 doi: 10.1016/j.bbrc.2009.05.138
Yan, H., Wei, W., Hu, L., Zhang, Y., Zhang, H., and Liu, J. (2021). Reduced feeding frequency improves feed efficiency associated with altered fecal microbiota and bile acid composition in pigs. Front Microbiol 12, 761210.
pubmed: 34712219 pmcid: 8546368 doi: 10.3389/fmicb.2021.761210
Yan, H.L., Cao, S.C., Hu, Y.D., Zhang, H.F., and Liu, J.B. (2020). Effects of methylsulfonylmethane on growth performance, immunity, antioxidant capacity, and meat quality in Pekin ducks. Poultry Sci 99, 1069–1074.
doi: 10.1016/j.psj.2019.10.002
Yuan, L., Kang, S.Y., Ward, L.A., To, T.L., and Saif, L.J. (1998). Antibody-secreting cell responses and protective immunity assessed in gnotobiotic pigs inoculated orally or intramuscularly with inactivated human rotavirus. J Virol 72, 330–338.
pubmed: 9420231 pmcid: 109380 doi: 10.1128/JVI.72.1.330-338.1998
Yue, B., Yang, H., Wu, J., Wang, J., Ru, W., Cheng, J., Huang, Y., Lan, X., Lei, C., and Chen, H. (2022). circSVIL regulates bovine myoblast development by inhibiting STAT1 phosphorylation. Sci China Life Sci 65, 376–386.
pubmed: 34024027 doi: 10.1007/s11427-020-1908-2
Zhang, S., Chang, M., Zhou, Z., Dai, X., and Xu, Z. (2018). pDHS-ELM: computational predictor for plant DNase I hypersensitive sites based on extreme learning machines. Mol Genet Genomics 293, 1035–1049.
pubmed: 29594496 doi: 10.1007/s00438-018-1436-3
Zhang, Y., Sun, Y., Wu, Z., Xiong, X., Zhang, J., Ma, J., Xiao, S., Huang, L., and Yang, B. (2021). Subcutaneous and intramuscular fat transcriptomes show large differences in network organization and associations with adipose traits in pigs. Sci China Life Sci 64, 1732–1746.
pubmed: 33527326 doi: 10.1007/s11427-020-1824-7
Zhao, S., Zhang, Y., Lu, X., Ding, H., Han, B., Song, X., Miao, H., Cui, X., Wei, S., Liu, W., et al. (2021). CDC20 regulates the cell proliferation and radiosensitivity of P53 mutant HCC cells through the Bcl-2/Bax pathway. Int J Biol Sci 17, 3608–3621.
pubmed: 34512169 pmcid: 8416732 doi: 10.7150/ijbs.64003

Auteurs

Jing Li (J)

Key Laboratory of Agro-Ecological Processes in Subtropical Region, Hunan Provincial Key Laboratory of Animal Nutritional Physiology and Metabolic Process, Hunan Research Center of Livestock and Poultry Sciences, South Central Experimental Station of Animal Nutrition and Feed Science in the Ministry of Agriculture, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, 410125, China.

Fenfen Liu (F)

Key Laboratory of Agro-Ecological Processes in Subtropical Region, Hunan Provincial Key Laboratory of Animal Nutritional Physiology and Metabolic Process, Hunan Research Center of Livestock and Poultry Sciences, South Central Experimental Station of Animal Nutrition and Feed Science in the Ministry of Agriculture, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, 410125, China.
University of Chinese Academy of Sciences, Beijing, 100008, China.

Kaibin Mo (K)

Key Laboratory of Agro-Ecological Processes in Subtropical Region, Hunan Provincial Key Laboratory of Animal Nutritional Physiology and Metabolic Process, Hunan Research Center of Livestock and Poultry Sciences, South Central Experimental Station of Animal Nutrition and Feed Science in the Ministry of Agriculture, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, 410125, China.
Guangdong Key Laboratory for Veterinary Drug Development and Safety Evaluation, College of Veterinary Medicine, South China Agricultural University, Guangzhou, 510642, China.

Hengjia Ni (H)

Key Laboratory of Agro-Ecological Processes in Subtropical Region, Hunan Provincial Key Laboratory of Animal Nutritional Physiology and Metabolic Process, Hunan Research Center of Livestock and Poultry Sciences, South Central Experimental Station of Animal Nutrition and Feed Science in the Ministry of Agriculture, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, 410125, China. nihengjia@isa.ac.cn.
University of Chinese Academy of Sciences, Beijing, 100008, China. nihengjia@isa.ac.cn.

Yulong Yin (Y)

Key Laboratory of Agro-Ecological Processes in Subtropical Region, Hunan Provincial Key Laboratory of Animal Nutritional Physiology and Metabolic Process, Hunan Research Center of Livestock and Poultry Sciences, South Central Experimental Station of Animal Nutrition and Feed Science in the Ministry of Agriculture, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, 410125, China. yinyulong@isa.ac.cn.
University of Chinese Academy of Sciences, Beijing, 100008, China. yinyulong@isa.ac.cn.

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