Fully activated structure of the sterol-bound Smoothened GPCR-Gi protein complex.


Journal

Proceedings of the National Academy of Sciences of the United States of America
ISSN: 1091-6490
Titre abrégé: Proc Natl Acad Sci U S A
Pays: United States
ID NLM: 7505876

Informations de publication

Date de publication:
05 Dec 2023
Historique:
pmc-release: 28 05 2024
medline: 30 11 2023
pubmed: 28 11 2023
entrez: 28 11 2023
Statut: ppublish

Résumé

Smoothened (SMO) is an oncoprotein and signal transducer in the Hedgehog signaling pathway that regulates cellular differentiation and embryogenesis. As a member of the Frizzled (Class F) family of G protein-coupled receptors (GPCRs), SMO biochemically and functionally interacts with Gi family proteins. However, key molecular features of fully activated, G protein-coupled SMO remain elusive. We present the atomistic structure of activated human SMO complexed with the heterotrimeric Gi protein and two sterol ligands, equilibrated at 310 K in a full lipid bilayer at physiological salt concentration and pH. In contrast to previous experimental structures, our equilibrated SMO complex exhibits complete breaking of the pi-cation interaction between R451

Identifiants

pubmed: 38015850
doi: 10.1073/pnas.2300919120
pmc: PMC10710022
doi:

Substances chimiques

Sterols 0
Ligands 0
Hedgehog Proteins 0
Receptors, G-Protein-Coupled 0
Smoothened Receptor 0

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

e2300919120

Subventions

Organisme : NHLBI NIH HHS
ID : R01 HL155532
Pays : United States
Organisme : NHLBI NIH HHS
ID : R35 HL150807
Pays : United States

Déclaration de conflit d'intérêts

Competing interests statement:The authors declare no competing interest.

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Auteurs

Amy-Doan P Vo (AP)

Materials and Process Simulation Center, Division of Chemistry & Chemical Engineering, California Institute of Technology, Pasadena, CA 91125.

Soo-Kyung Kim (SK)

Materials and Process Simulation Center, Division of Chemistry & Chemical Engineering, California Institute of Technology, Pasadena, CA 91125.

Moon Young Yang (MY)

Materials and Process Simulation Center, Division of Chemistry & Chemical Engineering, California Institute of Technology, Pasadena, CA 91125.

Alison E Ondrus (AE)

Department of Chemistry, University of Illinois Chicago, Chicago, IL 60607.
Department of Pharmaceutical Sciences, University of Illinois Chicago, Chicago, IL 60607.

William A Goddard (WA)

Materials and Process Simulation Center, Division of Chemistry & Chemical Engineering, California Institute of Technology, Pasadena, CA 91125.

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Classifications MeSH