Structural characterization and functional insights into the Type II Secretion System of the poly-extremophile Deinococcus radiodurans.

Deinococcus radiodurans T2SS cell envelope cryo-electron microscopy lipoproteins

Journal

The Journal of biological chemistry
ISSN: 1083-351X
Titre abrégé: J Biol Chem
Pays: United States
ID NLM: 2985121R

Informations de publication

Date de publication:
08 Dec 2023
Historique:
received: 05 10 2023
revised: 07 11 2023
accepted: 28 11 2023
medline: 11 12 2023
pubmed: 11 12 2023
entrez: 10 12 2023
Statut: aheadofprint

Résumé

The extremophile bacterium D. radiodurans boasts a distinctive cell envelope characterized by the regular arrangement of three protein complexes. Among these, the Type II Secretion System (T2SS) stands out as a pivotal structural component. We used cryo-electron microscopy to reveal unique features, such as an unconventional protein belt (DR_1364) around the main secretin (GspD), and a cap (DR_0940) found to be a separated subunit rather than integrated with GspD. Furthermore, a novel region at the N-terminus of the GspD constitutes an additional second gate, supplementing the one typically found in the outer membrane region. This T2SS was found to contribute to envelope integrity, while also playing a role in nucleic acid and nutrient trafficking. Studies on intact cell envelopes show a consistent T2SS structure repetition, highlighting its significance within the cellular framework.

Identifiants

pubmed: 38072042
pii: S0021-9258(23)02565-6
doi: 10.1016/j.jbc.2023.105537
pii:
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

105537

Informations de copyright

Copyright © 2023 The Authors. Published by Elsevier Inc. All rights reserved.

Déclaration de conflit d'intérêts

Competing interests Authors declare that they have no competing interests.

Auteurs

Domenica Farci (D)

Department of Plant Physiology, Warsaw University of Life Sciences - SGGW, Warsaw, 02-776, Poland;; Department of Life and Environmental Sciences, Università degli Studi di Cagliari, Viale S. Ignazio da Laconi 13, 09123 Cagliari, Italy;; ReGenFix Laboratories, R&D Department, Via Carducci 34, Sardara, Italy;. Electronic address: domenica.farci@unica.it.

Stefan Milenkovic (S)

Department of Physics and IOM/CNR, Università degli Studi di Cagliari, 09042 Monserrato, Italy.

Luca Iesu (L)

Department of Life and Environmental Sciences, Università degli Studi di Cagliari, Viale S. Ignazio da Laconi 13, 09123 Cagliari, Italy.

Marta Tanas (M)

Department of Life and Environmental Sciences, Università degli Studi di Cagliari, Viale S. Ignazio da Laconi 13, 09123 Cagliari, Italy.

Matteo Ceccarelli (M)

Department of Physics and IOM/CNR, Università degli Studi di Cagliari, 09042 Monserrato, Italy.

Dario Piano (D)

Department of Plant Physiology, Warsaw University of Life Sciences - SGGW, Warsaw, 02-776, Poland;; Department of Life and Environmental Sciences, Università degli Studi di Cagliari, Viale S. Ignazio da Laconi 13, 09123 Cagliari, Italy;; ReGenFix Laboratories, R&D Department, Via Carducci 34, Sardara, Italy;. Electronic address: dario.piano@unica.it.

Classifications MeSH