Polymorphisms in transcription factor binding sites and enhancer regions and pancreatic ductal adenocarcinoma risk.

Association study Enhancer Pancreatic cancer Single nucleotide polymorphism Transcription factor binding site

Journal

Human genomics
ISSN: 1479-7364
Titre abrégé: Hum Genomics
Pays: England
ID NLM: 101202210

Informations de publication

Date de publication:
02 Feb 2024
Historique:
received: 09 11 2023
accepted: 23 01 2024
medline: 3 2 2024
pubmed: 3 2 2024
entrez: 3 2 2024
Statut: epublish

Résumé

Genome-wide association studies (GWAS) are a powerful tool for detecting variants associated with complex traits and can help risk stratification and prevention strategies against pancreatic ductal adenocarcinoma (PDAC). However, the strict significance threshold commonly used makes it likely that many true risk loci are missed. Functional annotation of GWAS polymorphisms is a proven strategy to identify additional risk loci. We aimed to investigate single-nucleotide polymorphisms (SNP) in regulatory regions [transcription factor binding sites (TFBSs) and enhancers] that could change the expression profile of multiple genes they act upon and thereby modify PDAC risk. We analyzed a total of 12,636 PDAC cases and 43,443 controls from PanScan/PanC4 and the East Asian GWAS (discovery populations), and the PANDoRA consortium (replication population). We identified four associations that reached study-wide statistical significance in the overall meta-analysis: rs2472632(A) (enhancer variant, OR 1.10, 95%CI 1.06,1.13, p = 5.5 × 10

Identifiants

pubmed: 38308339
doi: 10.1186/s40246-024-00576-x
pii: 10.1186/s40246-024-00576-x
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

12

Subventions

Organisme : National Institute for Cancer Research - NICR (Programme EXCELES)
ID : LX22NPO5102
Organisme : National Institute for Cancer Research - NICR (Programme EXCELES)
ID : LX22NPO5102
Organisme : National Institute for Cancer Research - NICR (Programme EXCELES)
ID : LX22NPO5102
Organisme : National Institute for Cancer Research - NICR (Programme EXCELES)
ID : LX22NPO5102
Organisme : National Institute for Cancer Research - NICR (Programme EXCELES)
ID : LX22NPO5102

Informations de copyright

© 2024. The Author(s).

Références

Siegel RL, Miller KD, Fuchs HE, Jemal A. Cancer statistics, 2022. CA Cancer J Clin. 2022;72(1):7–33. https://doi.org/10.3322/caac.21708 .
doi: 10.3322/caac.21708 pubmed: 35020204
Afghani E, Klein AP. Pancreatic adenocarcinoma: trends in epidemiology, risk factors, and outcomes. Hematol Oncol Clin N Am. 2022;36(5):879–95. https://doi.org/10.1016/j.hoc.2022.07.002 .
doi: 10.1016/j.hoc.2022.07.002
Klein AP. Pancreatic cancer epidemiology: understanding the role of lifestyle and inherited risk factors. Nat Rev Gastroenterol Hepatol. 2021;18(7):493–502. https://doi.org/10.1038/s41575-021-00457-x .
doi: 10.1038/s41575-021-00457-x pubmed: 34002083 pmcid: 9265847
Ushio J, Kanno A, Ikeda E, et al. Pancreatic ductal adenocarcinoma: epidemiology and risk factors. Diagnostics. 2021;11(3):562. https://doi.org/10.3390/diagnostics11030562 .
doi: 10.3390/diagnostics11030562 pubmed: 33804776 pmcid: 8003883
Klein AP. Genetic susceptibility to pancreatic cancer. Mol Carcinog. 2012;51(1):14–24. https://doi.org/10.1002/mc.20855 .
doi: 10.1002/mc.20855 pubmed: 22162228 pmcid: 3570154
Buniello A, MacArthur JAL, Cerezo M, et al. The NHGRI-EBI GWAS Catalog of published genome-wide association studies, targeted arrays and summary statistics 2019. Nucleic Acids Res. 2019;47(D1):D1005–12. https://doi.org/10.1093/nar/gky1120 .
doi: 10.1093/nar/gky1120 pubmed: 30445434
Zhang Y, Qi G, Park JH, Chatterjee N. Estimation of complex effect-size distributions using summary-level statistics from genome-wide association studies across 32 complex traits. Nat Genet. 2018;50(9):1318–26. https://doi.org/10.1038/s41588-018-0193-x .
doi: 10.1038/s41588-018-0193-x pubmed: 30104760
Gentiluomo M, Canzian F, et al. Germline genetic variability in pancreatic cancer risk and prognosis. Semin Cancer Biol. 2022;79:105–31. https://doi.org/10.1016/j.semcancer.2020.08.003 .
doi: 10.1016/j.semcancer.2020.08.003 pubmed: 32818625
Childs EJ, Mocci E, Campa D, et al. Common variation at 2p13.3, 3q29, 7p13 and 17q25.1 associated with susceptibility to pancreatic cancer. Nat Genet. 2015;47(8):911–6. https://doi.org/10.1038/ng.3341 .
doi: 10.1038/ng.3341 pubmed: 26098869 pmcid: 4520746
Zhang YD, Hurson AN, Zhang H, et al. Assessment of polygenic architecture and risk prediction based on common variants across fourteen cancers. Nat Commun. 2020;11(1):3353. https://doi.org/10.1038/s41467-020-16483-3 .
doi: 10.1038/s41467-020-16483-3 pubmed: 32620889 pmcid: 7335068
Corradi C, Gentiluomo M, Gajdán L, et al. Genome-wide scan of long noncoding RNA single nucleotide polymorphisms and pancreatic cancer susceptibility. Int J Cancer. 2021;148(11):2779–88. https://doi.org/10.1002/ijc.33475 .
doi: 10.1002/ijc.33475 pubmed: 33534179
Lu Y, Corradi C, Gentiluomo M, et al. Association of genetic variants affecting microRNAs and pancreatic cancer risk. Front Genet. 2021;12:693933. https://doi.org/10.3389/fgene.2021.693933 .
doi: 10.3389/fgene.2021.693933 pubmed: 34527018 pmcid: 8435735
Pistoni L, Gentiluomo M, Lu Y, et al. Associations between pancreatic expression quantitative traits and risk of pancreatic ductal adenocarcinoma. Carcinogenesis. 2021;42(8):1037–45. https://doi.org/10.1093/carcin/bgab057 .
doi: 10.1093/carcin/bgab057 pubmed: 34216462
Gentiluomo M, Lu Y, Canzian F, Campa D. Genetic variants in taste-related genes and risk of pancreatic cancer. Mutagenesis. 2019;34(5–6):391–4. https://doi.org/10.1093/mutage/gez032 .
doi: 10.1093/mutage/gez032 pubmed: 31606007
Gentiluomo M, García PP, Galeotti AA, et al. Genetic variability of the ABCC2 gene and clinical outcomes in pancreatic cancer patients. Carcinogenesis. 2019;40(4):544–50. https://doi.org/10.1093/carcin/bgz006 .
doi: 10.1093/carcin/bgz006 pubmed: 30629142
Walsh N, Zhang H, Hyland PL, et al. Agnostic pathway/gene set analysis of genome-wide association data identifies associations for pancreatic cancer. J Natl Cancer Inst. 2019;111(6):557–67. https://doi.org/10.1093/jnci/djy155 .
doi: 10.1093/jnci/djy155 pubmed: 30541042
Pennacchio LA, Bickmore W, Dean A, Nobrega MA, Bejerano G. Enhancers: five essential questions. Nat Rev Genet. 2013;14(4):288–95. https://doi.org/10.1038/nrg3458 .
doi: 10.1038/nrg3458 pubmed: 23503198 pmcid: 4445073
Wray GA, Hahn MW, Abouheif E, et al. The evolution of transcriptional regulation in eukaryotes. Mol Biol Evol. 2003;20(9):1377–419. https://doi.org/10.1093/molbev/msg140 .
doi: 10.1093/molbev/msg140 pubmed: 12777501
Johnston AD, Simões-Pires CA, Thompson TV, Suzuki M, Greally JM. Functional genetic variants can mediate their regulatory effects through alteration of transcription factor binding. Nat Commun. 2019;10(1):3472. https://doi.org/10.1038/s41467-019-11412-5 .
doi: 10.1038/s41467-019-11412-5 pubmed: 31375681 pmcid: 6677801
Lin Y, Nakatochi M, Hosono Y, et al. Genome-wide association meta-analysis identifies GP2 gene risk variants for pancreatic cancer. Nat Commun. 2020;11(1):3175. https://doi.org/10.1038/s41467-020-16711-w .
doi: 10.1038/s41467-020-16711-w pubmed: 32581250 pmcid: 7314803
Campa D, Rizzato C, Capurso G, et al. Genetic susceptibility to pancreatic cancer and its functional characterisation: the PANcreatic Disease ReseArch (PANDoRA) consortium. Dig Liver Dis Off J Ital Soc Gastroenterol Ital Assoc Study Liver. 2013;45(2):95–9. https://doi.org/10.1016/j.dld.2012.09.014 .
doi: 10.1016/j.dld.2012.09.014
Riboli E, Hunt KJ, Slimani N, et al. European prospective investigation into cancer and nutrition (EPIC): study populations and data collection. Public Health Nutr. 2002;5(6B):1113–24. https://doi.org/10.1079/PHN2002394 .
doi: 10.1079/PHN2002394 pubmed: 12639222
Löw M, Stegmaier C, Ziegler H, Rothenbacher D, Brenner H, ESTHER Study. Epidemiological investigations of the chances of preventing, recognizing early and optimally treating chronic diseases in an elderly population (ESTHER study). Dtsch Med Wochenschr. 2004;129(49):2643–7. https://doi.org/10.1055/s-2004-836089 .
doi: 10.1055/s-2004-836089 pubmed: 15578318
Kumar S, Ambrosini G, Bucher P. SNP2TFBS—a database of regulatory SNPs affecting predicted transcription factor binding site affinity. Nucleic Acids Res. 2017;45(D1):D139–44. https://doi.org/10.1093/nar/gkw1064 .
doi: 10.1093/nar/gkw1064 pubmed: 27899579
Nasser J, Bergman DT, Fulco CP, et al. Genome-wide enhancer maps link risk variants to disease genes. Nature. 2021;593(7858):238–43. https://doi.org/10.1038/s41586-021-03446-x21 .
doi: 10.1038/s41586-021-03446-x21 pubmed: 33828297 pmcid: 9153265
Kent WJ, Sugnet CW, Furey TS, et al. The human genome browser at UCSC. Genome Res. 2002;12(6):996–1006. https://doi.org/10.1101/gr.229102 .
doi: 10.1101/gr.229102 pubmed: 12045153 pmcid: 186604
French JD, Edwards SL. The role of noncoding variants in heritable disease. Trends Genet. 2020;36(11):880–91. https://doi.org/10.1016/j.tig.2020.07.004 .
doi: 10.1016/j.tig.2020.07.004 pubmed: 32741549
Farh KKH, Marson A, Zhu J, et al. Genetic and epigenetic fine mapping of causal autoimmune disease variants. Nature. 2015;518(7539):337–43. https://doi.org/10.1038/nature1383 .
doi: 10.1038/nature1383 pubmed: 25363779
Gong Y, Qiu W, Ning X, et al. CCDC34 is up-regulated in bladder cancer and regulates bladder cancer cell proliferation, apoptosis and migration. Oncotarget. 2015;6(28):25856–67. https://doi.org/10.18632/oncotarget.4624 .
doi: 10.18632/oncotarget.4624 pubmed: 26312564 pmcid: 4694871
Liu LB, Huang J, Zhong JP, et al. High expression of CCDC34 Is associated with poor survival in cervical cancer patients. Med Sci Monit Int Med J Exp Clin Res. 2018;24:8383–90. https://doi.org/10.12659/MSM.913346 .
doi: 10.12659/MSM.913346
Geng W, Liang W, Fan Y, Ye Z, Zhang L. Overexpression of CCDC34 in colorectal cancer and its involvement in tumor growth, apoptosis and invasion. Mol Med Rep. 2018;17(1):465–73. https://doi.org/10.3892/mmr.2017.7860 .
doi: 10.3892/mmr.2017.7860 pubmed: 29115580
Qi W, Shao F, Huang Q. Expression of coiled-coil domain containing 34 (CCDC34) and its prognostic significance in pancreatic adenocarcinoma. Med Sci Monit Int Med J Exp Clin Res. 2017;23:6012–8. https://doi.org/10.12659/msm.907951 .
doi: 10.12659/msm.907951
Glinka A, Dolde C, Kirsch N, et al. LGR4 and LGR5 are R-spondin receptors mediating Wnt/β-catenin and Wnt/PCP signaling. EMBO Rep. 2011;12(10):1055–61. https://doi.org/10.1038/embor.2011.175 .
doi: 10.1038/embor.2011.175 pubmed: 21909076 pmcid: 3185347
Fan G, Ye D, Zhu S, et al. RTL1 promotes melanoma proliferation by regulating Wnt/β-catenin signaling. Oncotarget. 2017;8(62):106026–37. https://doi.org/10.18632/oncotarget.22523 .
doi: 10.18632/oncotarget.22523 pubmed: 29285312 pmcid: 5739699
van Andel H, Ren Z, Koopmans I, et al. Aberrantly expressed LGR4 empowers Wnt signaling in multiple myeloma by hijacking osteoblast-derived R-spondins. Proc Natl Acad Sci. 2017;114(2):376–81. https://doi.org/10.1073/pnas.1618650114 .
doi: 10.1073/pnas.1618650114 pubmed: 28028233
Wang Z, Yin P, Sun Y, et al. LGR4 maintains HGSOC cell epithelial phenotype and stem-like traits. Gynecol Oncol. 2020;159(3):839–49. https://doi.org/10.1016/j.ygyno.2020.09.020 .
doi: 10.1016/j.ygyno.2020.09.020 pubmed: 32980127
Kang YE, Kim JM, Kim KS, et al. Upregulation of RSPO2-GPR48/LGR4 signaling in papillary thyroid carcinoma contributes to tumor progression. Oncotarget. 2017;8(70):114980–94. https://doi.org/10.18632/oncotarget.22692 .
doi: 10.18632/oncotarget.22692 pubmed: 29383135 pmcid: 5777747
Modi S, Kir D, Banerjee S, Saluja A. Control of apoptosis in treatment and biology of pancreatic cancer. J Cell Biochem. 2016;117(2):279–88. https://doi.org/10.1002/jcb.25284 .
doi: 10.1002/jcb.25284 pubmed: 26206252 pmcid: 5724757
Mazerbourg S, Bouley DM, Sudo S, et al. Leucine-rich repeat-containing, G protein-coupled receptor 4 null mice exhibit intrauterine growth retardation associated with embryonic and perinatal lethality. Mol Endocrinol Baltim Md. 2004;18(9):2241–54. https://doi.org/10.1210/me.2004-0133 .
doi: 10.1210/me.2004-0133
Onda T, Uzawa K, Nakashima D, et al. Lin-7C/VELI3/MALS-3: an essential component in metastasis of human squamous cell carcinoma. Cancer Res. 2007;67(20):9643–8. https://doi.org/10.1158/0008-5472.CAN-07-1911 .
doi: 10.1158/0008-5472.CAN-07-1911 pubmed: 17942893
Shinawi M, Sahoo T, Maranda B, et al. 11p14.1 microdeletions associated with ADHD, autism, developmental delay, and obesity. Am J Med Genet A. 2011;155A(6):1272–80. https://doi.org/10.1002/ajmg.a.33878 .
doi: 10.1002/ajmg.a.33878 pubmed: 21567907
Jung SW, Lee J, Cho AE. Elucidating the bacterial membrane disruption mechanism of human α-defensin 5: a theoretical study. J Phys Chem B. 2017;121(4):741–8. https://doi.org/10.1021/acs.jpcb.6b11806 .
doi: 10.1021/acs.jpcb.6b11806 pubmed: 28067516
Yang E, Shen J. The roles and functions of Paneth cells in Crohn’s disease: a critical review. Cell Prolif. 2021;54(1):e12958. https://doi.org/10.1111/cpr.12958 .
doi: 10.1111/cpr.12958 pubmed: 33174662
Selsted ME, Ouellette AJ. Mammalian defensins in the antimicrobial immune response. Nat Immunol. 2005;6:551–7. https://doi.org/10.1038/ni1206 .
doi: 10.1038/ni1206 pubmed: 15908936
Tobi M, Kim M, Weinstein DH, et al. Prospective markers for early diagnosis and prognosis of sporadic pancreatic ductal adenocarcinoma. Dig Dis Sci. 2013;58:744–50. https://doi.org/10.1007/s10620-012-2387-x .
doi: 10.1007/s10620-012-2387-x pubmed: 23001406
Cunha DM, Koike MK, Barbeiro DF, et al. Increased intestinal production of α-defensins in aged rats with acute pancreatic injury. Exp Gerontol. 2014;60:215–9. https://doi.org/10.1016/j.exger.2014.11.008 .
doi: 10.1016/j.exger.2014.11.008 pubmed: 25449854
Shimizu Y, Nakamura K, Kikuchi M, et al. Lower human defensin 5 in elderly people compared to middle-aged is associated with differences in the intestinal microbiota composition: the DOSANCO health study. Geroscience. 2022;44(2):997–1009. https://doi.org/10.1007/s11357-021-00398-y .
doi: 10.1007/s11357-021-00398-y pubmed: 34105106
Zhang JJ, Zhu Y, Xie KL, et al. Yin Yang-1 suppresses invasion and metastasis of pancreatic ductal adenocarcinoma by downregulating MMP10 in a MUC4/ErbB2/p38/MEF2C-dependent mechanism. Mol Cancer. 2014;13(1):130. https://doi.org/10.1186/1476-4598-13-130 .
doi: 10.1186/1476-4598-13-130 pubmed: 24884523 pmcid: 4047260
Mahawithitwong P, Ohuchida K, Ikenaga N, et al. Kindlin-1 expression is involved in migration and invasion of pancreatic cancer. Int J Oncol. 2013;42(4):1360–6. https://doi.org/10.3892/ijo .
doi: 10.3892/ijo pubmed: 23440354
Wang L, Ai M, Nie M, et al. EHF promotes colorectal carcinoma progression by activating TGF-β1 transcription and canonical TGF-β signaling. Cancer Sci. 2020;111(7):2310–24. https://doi.org/10.1111/cas.14444 .
doi: 10.1111/cas.14444 pubmed: 32372436 pmcid: 7385339
Zhou T, Liu J, Xie Y, et al. ESE3/EHF, a promising target of rosiglitazone, suppresses pancreatic cancer stemness by downregulating CXCR4. Gut. 2022;71(2):357–71. https://doi.org/10.1136/gutjnl-2020-321952 .
doi: 10.1136/gutjnl-2020-321952 pubmed: 33674341
Liu J, Jiang W, Zhao K, et al. Tumoral EHF predicts the efficacy of anti-PD1 therapy in pancreatic ductal adenocarcinoma. J Exp Med. 2019;216(3):656–73. https://doi.org/10.1084/jem.2018074920 .
doi: 10.1084/jem.2018074920 pubmed: 30733283 pmcid: 6400540
Merz S, Breunig M, Melzer MK, et al. Single-cell profiling of GP2-enriched pancreatic progenitors to simultaneously create acinar, ductal, and endocrine organoids. Theranostics. 2023;13(6):1949–73. https://doi.org/10.7150/thno.78323 .
doi: 10.7150/thno.78323 pubmed: 37064874 pmcid: 10091881

Auteurs

Pelin Ünal (P)

Genomic Epidemiology Group, German Cancer Research Center, In Neuenheimer Feld 280, 69120, Heidelberg, Germany.

Ye Lu (Y)

Genomic Epidemiology Group, German Cancer Research Center, In Neuenheimer Feld 280, 69120, Heidelberg, Germany.

Bas Bueno-de-Mesquita (B)

Department for Determinants of Chronic Diseases, National Institute for Public Health and the Environment, Bilthoven, The Netherlands.

Casper H J van Eijck (CHJ)

Department of Surgery, Erasmus MC University Medical Center, Rotterdam, The Netherlands.

Renata Talar-Wojnarowska (R)

Department of Digestive Tract Diseases, Medical University of Lodz, Lodz, Poland.

Andrea Szentesi (A)

Institute for Translational Medicine, Medical School, University of Pécs, Pécs, Hungary.

Maria Gazouli (M)

Laboratory of Biology, Medical School, National and Kapodistrian University of Athens, Athens, Greece.

Edita Kreivenaite (E)

Gastroenterology Department and Institute for Digestive Research, Lithuanian University of Health Sciences, Kaunas, Lithuania.

Francesca Tavano (F)

Division of Gastroenterology and Research Laboratory, Fondazione IRCCS "Casa Sollievo della Sofferenza" Hospital, San Giovanni Rotondo, FG, Italy.

Ewa Małecka-Wojciesko (E)

Department of Digestive Tract Diseases, Medical University of Lodz, Lodz, Poland.

Bálint Erőss (B)

Institute for Translational Medicine, Medical School, University of Pécs, Pécs, Hungary.
Center for Translational Medicine, Semmelweis University, Budapest, Hungary.
Division of Pancreatic Diseases, Heart and Vascular Center, Semmelweis University, Budapest, Hungary.

Martin Oliverius (M)

Department of Surgery, University Hospital Kralovske Vinohrady, Third Faculty of Medicine, Charles University, Prague, Czech Republic.

Stefania Bunduc (S)

Center for Translational Medicine, Semmelweis University, Budapest, Hungary.
Division of Pancreatic Diseases, Heart and Vascular Center, Semmelweis University, Budapest, Hungary.
Carol Davila University of Medicine and Pharmacy, Bucharest, Romania.

Mateus Nóbrega Aoki (M)

Laboratory for Applied Science and Technology in Health, Carlos Chagas Institute, Curitiba, PR, Brazil.

Ludmila Vodickova (L)

Biomedical Center, Faculty of Medicine in Pilsen, Charles University, Plzeň, Czech Republic.
Department of Molecular Biology of Cancer, Institute of Experimental Medicine of the Czech Academy of Sciences, Prague, Czech Republic.
Institute of Biology and Medical Genetics, Institute of Physiology, 1st Faculty of Medicine, Charles University, Prague, Czech Republic.

Ugo Boggi (U)

Division of General and Transplant Surgery, Pisa University Hospital, Pisa, Italy.

Matteo Giaccherini (M)

Department of Biology, University of Pisa, Pisa, Italy.

Jurate Kondrackiene (J)

Gastroenterology Department and Institute for Digestive Research, Lithuanian University of Health Sciences, Kaunas, Lithuania.

Roger Chammas (R)

Department of Radiology and Oncology, Institute of Cancer of São Paulo, São Paulo, Brazil.

Orazio Palmieri (O)

Division of Gastroenterology and Research Laboratory, Fondazione IRCCS "Casa Sollievo della Sofferenza" Hospital, San Giovanni Rotondo, FG, Italy.

George E Theodoropoulos (GE)

First Propaedeutic University Surgery Clinic, Hippocratio General Hospital, Medical School, National and Kapodistrian University of Athens, Athens, Greece.

Maarten F Bijlsma (MF)

Laboratory for Experimental Oncology and Radiobiology, Center of Experimental Molecular Medicine, Amsterdam UMC Location University of Amsterdam, Amsterdam, the Netherlands.
Cancer Center Amsterdam, Imaging and Biomarkers, Amsterdam, the Netherlands.

Daniela Basso (D)

Department of Medicine, Laboratory Medicine, University of Padova, Padua, Italy.

Beatrice Mohelnikova-Duchonova (B)

Department of Oncology, Faculty of Medicine and Dentistry, Palacky University, Olomouc, Czech Republic.

Pavel Soucek (P)

Biomedical Center, Faculty of Medicine in Pilsen, Charles University, Plzeň, Czech Republic.

Jakob R Izbicki (JR)

Department of General Visceral and Thoracic Surgery, University of Hamburg Medical Institutions, Hamburg, Germany.

Vytautas Kiudelis (V)

Gastroenterology Department and Institute for Digestive Research, Lithuanian University of Health Sciences, Kaunas, Lithuania.

Giuseppe Vanella (G)

PancreatoBiliary Endoscopy and Endosonography Division, Pancreas Translational and Clinical Research Center, San Raffaele Scientific Institute, Milan, Italy.
Digestive and Liver Disease Unit, S. Andrea Hospital, Rome, Italy.

Paolo Giorgio Arcidiacono (PG)

PancreatoBiliary Endoscopy and Endosonography Division, Pancreas Translational and Clinical Research Center, San Raffaele Scientific Institute, Milan, Italy.

Barbara Włodarczyk (B)

Department of Digestive Tract Diseases, Medical University of Lodz, Lodz, Poland.

Thilo Hackert (T)

Department of General, Visceral and Transplant Surgery, Heidelberg University Hospital, Heidelberg, Germany.

Ben Schöttker (B)

Division of Clinical Epidemiology and Aging Research, German Cancer Research Center, Heidelberg, Germany.
Network Aging Research (NAR), Heidelberg University, Heidelberg, Germany.

Faik G Uzunoglu (FG)

Department of General Visceral and Thoracic Surgery, University of Hamburg Medical Institutions, Hamburg, Germany.

Franco Bambi (F)

Blood Transfusion Service, Meyer Children's Hospital, Florence, Italy.

Mara Goetz (M)

Department of General Visceral and Thoracic Surgery, University of Hamburg Medical Institutions, Hamburg, Germany.

Viktor Hlavac (V)

Biomedical Center, Faculty of Medicine in Pilsen, Charles University, Plzeň, Czech Republic.

Hermann Brenner (H)

Division of Clinical Epidemiology and Aging Research, German Cancer Research Center, Heidelberg, Germany.
Network Aging Research (NAR), Heidelberg University, Heidelberg, Germany.
Division of Preventive Oncology, German Cancer Research Center and National Center for Tumor Diseases, Heidelberg, Germany.
German Cancer Consortium (DKTK), German Cancer Research Center, Heidelberg, Germany.

Francesco Perri (F)

Division of Gastroenterology and Research Laboratory, Fondazione IRCCS "Casa Sollievo della Sofferenza" Hospital, San Giovanni Rotondo, FG, Italy.

Silvia Carrara (S)

Endoscopic Unit, Department of Gastroenterology, IRCCS Humanitas Research Hospital, Milan, Italy.

Stefano Landi (S)

Department of Biology, University of Pisa, Pisa, Italy.

Péter Hegyi (P)

Institute for Translational Medicine, Medical School, University of Pécs, Pécs, Hungary.
János Szentágothai Research Center, University of Pécs, Pécs, Hungary.
Center for Translational Medicine, Semmelweis University, Budapest, Hungary.
Division of Pancreatic Diseases, Heart and Vascular Center, Semmelweis University, Budapest, Hungary.

Frederike Dijk (F)

Department of Pathology, Cancer Center Amsterdam, Amsterdam UMC, University of Amsterdam, Amsterdam, the Netherlands.

Evaristo Maiello (E)

Department of Oncology, Fondazione IRCCS "Casa Sollievo della Sofferenza" Hospital, San Giovanni Rotondo, FG, Italy.

Giovanni Capretti (G)

Pancreatic Unit, IRCCS Humanitas Research Hospital, Milan, Italy.
Department of Biomedical Sciences, Humanitas University, Milan, Italy.

Sabrina Gloria Giulia Testoni (SGG)

PancreatoBiliary Endoscopy and Endosonography Division, Pancreas Translational and Clinical Research Center, San Raffaele Scientific Institute, Milan, Italy.

Maria Chiara Petrone (MC)

PancreatoBiliary Endoscopy and Endosonography Division, Pancreas Translational and Clinical Research Center, San Raffaele Scientific Institute, Milan, Italy.

Hannah Stocker (H)

Division of Clinical Epidemiology and Aging Research, German Cancer Research Center, Heidelberg, Germany.
Network Aging Research (NAR), Heidelberg University, Heidelberg, Germany.

Stefano Ermini (S)

Blood Transfusion Service, Meyer Children's Hospital, Florence, Italy.

Livia Archibugi (L)

PancreatoBiliary Endoscopy and Endosonography Division, Pancreas Translational and Clinical Research Center, San Raffaele Scientific Institute, Milan, Italy.
Digestive and Liver Disease Unit, S. Andrea Hospital, Rome, Italy.

Manuel Gentiluomo (M)

Department of Biology, University of Pisa, Pisa, Italy.

Giulia Martina Cavestro (GM)

Gastroenterology and Gastrointestinal Endoscopy Unit, IRCCS San Raffaele Scientific Institute, Vita-Salute San Raffaele University, Milan, Italy.

Raffaele Pezzilli (R)

Department of Gastroenterology, San Carlo Hospital, Potenza, Italy.

Gregorio Di Franco (G)

General Surgery Unit, Department of Translational Research and New Technologies in Medicine and Surgery, University of Pisa, Pisa, Italy.

Anna Caterina Milanetto (AC)

Department of Surgery, Oncology and Gastroenterology, University of Padova, Padua, Italy.

Cosimo Sperti (C)

Department of Surgery, Oncology and Gastroenterology, University of Padova, Padua, Italy.

John P Neoptolemos (JP)

Department of General, Visceral and Transplant Surgery, Heidelberg University Hospital, Heidelberg, Germany.

Luca Morelli (L)

General Surgery Unit, Department of Translational Research and New Technologies in Medicine and Surgery, University of Pisa, Pisa, Italy.

Klara Vokacova (K)

Department of Molecular Biology of Cancer, Institute of Experimental Medicine of the Czech Academy of Sciences, Prague, Czech Republic.
Institute of Biology and Medical Genetics, Institute of Physiology, 1st Faculty of Medicine, Charles University, Prague, Czech Republic.

Claudio Pasquali (C)

Department of Surgery, Oncology and Gastroenterology, University of Padova, Padua, Italy.

Rita T Lawlor (RT)

Department of Diagnostics and Public Health, ARC-Net Centre for Applied Research on Cancer, University of Verona, Verona, Italy.

Francesca Bazzocchi (F)

Department of Surgery, Fondazione IRCCS "Casa Sollievo della Sofferenza" Hospital, San Giovanni Rotondo, FG, Italy.

Juozas Kupcinskas (J)

Gastroenterology Department and Institute for Digestive Research, Lithuanian University of Health Sciences, Kaunas, Lithuania.

Gabriele Capurso (G)

PancreatoBiliary Endoscopy and Endosonography Division, Pancreas Translational and Clinical Research Center, San Raffaele Scientific Institute, Milan, Italy.
Digestive and Liver Disease Unit, S. Andrea Hospital, Rome, Italy.

Daniele Campa (D)

Department of Biology, University of Pisa, Pisa, Italy.

Federico Canzian (F)

Genomic Epidemiology Group, German Cancer Research Center, In Neuenheimer Feld 280, 69120, Heidelberg, Germany. f.canzian@dkfz.de.

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