Heterozygous loss-of-function variants in DOCK4 cause neurodevelopmental delay and microcephaly.


Journal

Human genetics
ISSN: 1432-1203
Titre abrégé: Hum Genet
Pays: Germany
ID NLM: 7613873

Informations de publication

Date de publication:
25 Mar 2024
Historique:
received: 08 11 2023
accepted: 09 02 2024
medline: 25 3 2024
pubmed: 25 3 2024
entrez: 25 3 2024
Statut: aheadofprint

Résumé

Neurons form the basic anatomical and functional structure of the nervous system, and defects in neuronal differentiation or formation of neurites are associated with various psychiatric and neurodevelopmental disorders. Dynamic changes in the cytoskeleton are essential for this process, which is, inter alia, controlled by the dedicator of cytokinesis 4 (DOCK4) through the activation of RAC1. Here, we clinically describe 7 individuals (6 males and one female) with variants in DOCK4 and overlapping phenotype of mild to severe global developmental delay. Additional symptoms include coordination or gait abnormalities, microcephaly, nonspecific brain malformations, hypotonia and seizures. Four individuals carry missense variants (three of them detected de novo) and three individuals carry null variants (two of them maternally inherited). Molecular modeling of the heterozygous missense variants suggests that the majority of them affect the globular structure of DOCK4. In vitro functional expression studies in transfected Neuro-2A cells showed that all missense variants impaired neurite outgrowth. Furthermore, Dock4 knockout Neuro-2A cells also exhibited defects in promoting neurite outgrowth. Our results, including clinical, molecular and functional data, suggest that loss-of-function variants in DOCK4 probable cause a variable spectrum of a novel neurodevelopmental disorder with microcephaly.

Identifiants

pubmed: 38526744
doi: 10.1007/s00439-024-02655-4
pii: 10.1007/s00439-024-02655-4
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Informations de copyright

© 2024. The Author(s).

Références

Abraham S, Scarcia M, Bagshaw RD, McMahon K, Grant G, Harvey T, Yeo M, Esteves FOG, Thygesen HH, Jones PF, Speirs V, Hanby AM, Selby PJ, Lorger M, Dear TN, Pawson T, Marshall CJ, Mavria G (2015) A Rac/Cdc42 exchange factor complex promotes formation of lateral filopodia and blood vessel lumen morphogenesis. Nat Commun 6:7286
pubmed: 26129894 doi: 10.1038/ncomms8286
Cacheiro P, Muñoz-Fuentes V, Murray SA, Dickinson ME, Bucan M, Nutter LMJ, Peterson KA, Haselimashhadi H, Flenniken AM, Morgan H, Westerberg H, Konopka T, Hsu C-W, Christiansen A, Lanza DG, Beaudet AL, Heaney JD, Fuchs H, Gailus-Durner V, Sorg T, Prochazka J, Novosadova V, Lelliott CJ, Wardle-Jones H, Wells S, Teboul L, Cater H, Stewart M, Hough T, Wurst W, Sedlacek R, Adams DJ, Seavitt JR, Tocchini-Valentini G, Mammano F, Braun RE, McKerlie C, Herault Y, de Angelis MH, Mallon A-M, Lloyd KCK, Brown SDM, Parkinson H, Meehan TF, Smedley D (2020) Human and mouse essentiality screens as a resource for disease gene discovery. Nat Commun 11(1):655
pubmed: 32005800 pmcid: 6994715 doi: 10.1038/s41467-020-14284-2
Chen Q, Peto CA, Shelton GD, Mizisin A, Sawchenko PE, Schubert D (2009) Loss of modifier of cell adhesion reveals a pathway leading to axonal degeneration. J Neurosci Official J Soc Neurosci 29(1):118–130
doi: 10.1523/JNEUROSCI.3985-08.2009
Cioclu MC, Mosca I, Ambrosino P, Puzo D, Bayat A, Wortmann SB, Koch J, Strehlow V, Shirai K, Matsumoto N, Sanders SJ, Michaud V, Legendre M, Riva A, Striano P, Muhle H, Pendziwiat M, Lesca G, Mangano GD, Nardello R, Lemke JR, Møller RS, Soldovieri MV, Rubboli G, Taglialatela M (2023) KCNT2-related disorders: phenotypes, functional, and pharmacological properties. Ann Neurol 94(2):332–349
pubmed: 37062836 doi: 10.1002/ana.26662
Ehrhart F, Willighagen EL, Kutmon M, van Hoften M, Curfs LMG, Evelo CT (2021) A resource to explore the discovery of rare diseases and their causative genes. Sci Data 8(1):124
pubmed: 33947870 pmcid: 8096966 doi: 10.1038/s41597-021-00905-y
Guex N, Peitsch MC (1997) SWISS-MODEL and the Swiss-PdbViewer: an environment for comparative protein modeling. Electrophoresis 18(15):2714–2723
pubmed: 9504803 doi: 10.1002/elps.1150181505
Guo D, Peng Y, Wang L, Sun X, Wang X, Liang C, Yang X, Li S, Xu J, Ye W-C, Jiang B, Shi L (2021) Autism-like social deficit generated by Dock4 deficiency is rescued by restoration of Rac1 activity and NMDA receptor function. Mol Psych 26(5):1505–1519
doi: 10.1038/s41380-019-0472-7
Hornbeck PV, Zhang B, Murray B, Kornhauser JM, Latham V, Skrzypek E (2015) PhosphoSitePlus, 2014: mutations, PTMs and recalibrations. Nucleic acids research 43(Database issue):D512–20
Huang M, Liang C, Li S, Zhang J, Guo D, Zhao B, Liu Y, Peng Y, Xu J, Liu W, Guo G, Shi L (2019) Two autism/dyslexia linked variations of DOCK4 disrupt the gene function on Rac1/Rap1 activation, neurite outgrowth, and synapse development. Front Cell Neurosci 13:577
pubmed: 32009906 doi: 10.3389/fncel.2019.00577
Huang X, Henck J, Qiu C, Sreenivasan VKA, Balachandran S, Amarie OV, Hrabě de Angelis M, Behncke RY, Chan W-L, Despang A, Dickel DE, Duran M, Feuchtinger A, Fuchs H, Gailus-Durner V, Haag N, Hägerling R, Hansmeier N, Hennig F, Marshall C, Rajderkar S, Ringel A, Robson M, Saunders LM, Da Silva-Buttkus P, Spielmann N, Srivatsan SR, Ulferts S, Wittler L, Zhu Y, Kalscheuer VM, Ibrahim DM, Kurth I, Kornak U, Visel A, Pennacchio LA, Beier DR, Trapnell C, Cao J, Shendure J, Spielmann M (2023) Single-cell, whole-embryo phenotyping of mammalian developmental disorders. Nature 623(7988):772–781
pubmed: 37968388 pmcid: 10665194 doi: 10.1038/s41586-023-06548-w
Humphrey W, Dalke A, Schulten K (1996) VMD: visual molecular dynamics. J Mol Graphs 14(1):33–38
doi: 10.1016/0263-7855(96)00018-5
Ioannidis NM, Rothstein JH, Pejaver V, Middha S, McDonnell SK, Baheti S, Musolf A, Li Q, Holzinger E, Karyadi D, Cannon-Albright LA, Teerlink CC, Stanford JL, Isaacs WB, Xu J, Cooney KA, Lange EM, Schleutker J, Carpten JD, Powell IJ, Cussenot O, Cancel-Tassin G, Giles GG, MacInnis RJ, Maier C, Hsieh C-L, Wiklund F, Catalona WJ, Foulkes WD, Mandal D, Eeles RA, Kote-Jarai Z, Bustamante CD, Schaid DJ, Hastie T, Ostrander EA, Bailey-Wilson JE, Radivojac P, Thibodeau SN, Whittemore AS, Sieh W (2016) REVEL: an ensemble method for predicting the pathogenicity of rare missense variants. Am J Hum Genet 99(4):877–885
pubmed: 27666373 pmcid: 5065685 doi: 10.1016/j.ajhg.2016.08.016
Jónsson H, Sulem P, Kehr B, Kristmundsdottir S, Zink F, Hjartarson E, Hardarson MT, Hjorleifsson KE, Eggertsson HP, Gudjonsson SA, Ward LD, Arnadottir GA, Helgason EA, Helgason H, Gylfason A, Jonasdottir A, Jonasdottir A, Rafnar T, Frigge M, Stacey SN, Th Magnusson O, Thorsteinsdottir U, Masson G, Kong A, Halldorsson BV, Helgason A, Gudbjartsson DF, Stefansson K (2017) Parental influence on human germline de novo mutations in 1,548 trios from Iceland. Nature 549(7673):519–522
pubmed: 28959963 doi: 10.1038/nature24018
Jumper J, Evans R, Pritzel A, Green T, Figurnov M, Ronneberger O, Tunyasuvunakool K, Bates R, Žídek A, Potapenko A, Bridgland A, Meyer C, Kohl SAA, Ballard AJ, Cowie A, Romera-Paredes B, Nikolov S, Jain R, Adler J, Back T, Petersen S, Reiman D, Clancy E, Zielinski M, Steinegger M, Pacholska M, Berghammer T, Bodenstein S, Silver D, Vinyals O, Senior AW, Kavukcuoglu K, Kohli P, Hassabis D (2021) Highly accurate protein structure prediction with AlphaFold. Nature 596(7873):583–589
pubmed: 34265844 pmcid: 8371605 doi: 10.1038/s41586-021-03819-2
Karczewski KJ, Francioli LC, Tiao G, Cummings BB, Alföldi J, Wang Q, Collins RL, Laricchia KM, Ganna A, Birnbaum DP, Gauthier LD, Brand H, Solomonson M, Watts NA, Rhodes D, Singer-Berk M, England EM, Seaby EG, Kosmicki JA, Walters RK, Tashman K, Farjoun Y, Banks E, Poterba T, Wang A, Seed C, Whiffin N, Chong JX, Samocha KE, Pierce-Hoffman E, Zappala Z, O’Donnell-Luria AH, Minikel EV, Weisburd B, Lek M, Ware JS, Vittal C, Armean IM, Bergelson L, Cibulskis K, Connolly KM, Covarrubias M, Donnelly S, Ferriera S, Gabriel S, Gentry J, Gupta N, Jeandet T, Kaplan D, Llanwarne C, Munshi R, Novod S, Petrillo N, Roazen D, Ruano-Rubio V, Saltzman A, Schleicher M, Soto J, Tibbetts K, Tolonen C, Wade G, Talkowski ME, Neale BM, Daly MJ, MacArthur DG (2020) The mutational constraint spectrum quantified from variation in 141,456 humans. Nature 581(7809):434–443
pubmed: 32461654 pmcid: 7334197 doi: 10.1038/s41586-020-2308-7
Klau J, Abou Jamra R, Radtke M, Oppermann H, Lemke JR, Beblo S, Popp B (2022) Exome first approach to reduce diagnostic costs and time - retrospective analysis of 111 individuals with rare neurodevelopmental disorders. Eur J Hum Genet 30(1):117–125
pubmed: 34690354 doi: 10.1038/s41431-021-00981-z
Kochinke K, Zweier C, Nijhof B, Fenckova M, Cizek P, Honti F, Keerthikumar S, Oortveld MAW, Kleefstra T, Kramer JM, Webber C, Huynen MA, Schenck A (2016) Systematic phenomics analysis deconvolutes genes mutated in intellectual disability into biologically coherent modules. Am J Hum Genet 98(1):149–164
pubmed: 26748517 pmcid: 4716705 doi: 10.1016/j.ajhg.2015.11.024
Kukimoto-Niino M, Katsura K, Kaushik R, Ehara H, Yokoyama T, Uchikubo-Kamo T, Nakagawa R, Mishima-Tsumagari C, Yonemochi M, Ikeda M, Hanada K, Zhang KYJ, Shirouzu M (2021) Cryo-EM structure of the human ELMO1-DOCK5-Rac1 complex. Sci Adv 7(30)
Kumar M, Michael S, Alvarado-Valverde J, Mészáros B, Sámano-Sánchez H, Zeke A, Dobson L, Lazar T, Örd M, Nagpal A, Farahi N, Käser M, Kraleti R, Davey NE, Pancsa R, Chemes LB, Gibson TJ (2022) The eukaryotic linear motif resource: 2022 release. Nucleic Acids Res 50(D1):D497–D508
pubmed: 34718738 doi: 10.1093/nar/gkab975
Kurishev AO, Karpov DS, Nadolinskaia NI, Goncharenko AV, Golimbet VE (2022) CRISPR/Cas-based approaches to study schizophrenia and other neurodevelopmental disorders. Int J Mol Sci 24(1)
Larsen R, Fink T, Müller-Wolff T (2016) Nervensystem. Anästhesie und Intensivmedizin für die Fachpflege:13–25
Leitão E, Schröder C, Parenti I, Dalle C, Rastetter A, Kühnel T, Kuechler A, Kaya S, Gérard B, Schaefer E, Nava C, Drouot N, Engel C, Piard J, Duban-Bedu B, Villard L, Stegmann APA, Vanhoutte EK, Verdonschot JAJ, Kaiser FJ, Tran Mau-Them F, Scala M, Striano P, Frints SGM, Argilli E, Sherr EH, Elder F, Buratti J, Keren B, Mignot C, Héron D, Mandel J-L, Gecz J, Kalscheuer VM, Horsthemke B, Piton A, Depienne C (2022) Systematic analysis and prediction of genes associated with monogenic disorders on human chromosome X. Nat Commun 13(1):6570
pubmed: 36323681 pmcid: 9630267 doi: 10.1038/s41467-022-34264-y
Lindeboom RG, Supek F, Lehner B (2016) The rules and impact of nonsense-mediated mRNA decay in human cancers. Nat Genet 48(10):1112–1118
pubmed: 27618451 pmcid: 5045715 doi: 10.1038/ng.3664
Meijering E, Jacob M, Sarria J-CF, Steiner P, Hirling H, Unser M (2004) Design and validation of a tool for neurite tracing and analysis in fluorescence microscopy images. Cytom Part a: J Int Soc Anal Cytol 58(2):167–176
doi: 10.1002/cyto.a.20022
Nimchinsky EA, Sabatini BL, Svoboda K (2002) Structure and function of dendritic spines. Annu Rev Physiol 64:313–353
pubmed: 11826272 doi: 10.1146/annurev.physiol.64.081501.160008
Oppermann H, Faust H, Yamanishi U, Meixensberger J, Gaunitz F (2019) Carnosine inhibits glioblastoma growth independent from PI3K/Akt/mTOR signaling. PLoS ONE 14(6):e0218972
pubmed: 31247000 pmcid: 6597087 doi: 10.1371/journal.pone.0218972
Oppermann H, Marcos-Grañeda E, Weiss LA, Gurnett CA, Jelsig AM, Vineke SH, Isidor B, Mercier S, Magnussen K, Zacher P, Hashim M, Pagnamenta AT, Race S, Srivastava S, Frazier Z, Maiwald R, Pergande M, Milani D, Rinelli M, Levy J, Krey I, Fontana P, Lonardo F, Riley S, Kretzer J, Rankin J, Reis LM, Semina EV, Reuter MS, Scherer SW, Iascone M, Weis D, Fagerberg CR, Brasch-Andersen C, Hansen LK, Kuechler A, Noble N, Gardham A, Tenney J, Rathore G, Beck-Woedl S, Haack TB, Pavlidou DC, Atallah I, Vodopiutz J, Janecke AR, Hsieh T-C, Lesmann H, Klinkhammer H, Krawitz PM, Lemke JR, Jamra RA, Nieto M, Tümer Z, Platzer K (2023) CUX1-related neurodevelopmental disorder: deep insights into phenotype-genotype spectrum and underlying pathology. Eur J Hum Genet 31(11):1251–1260
pubmed: 37644171 pmcid: 10620399 doi: 10.1038/s41431-023-01445-2
Pagnamenta AT, Bacchelli E, de Jonge MV, Mirza G, Scerri TS, Minopoli F, Chiocchetti A, Ludwig KU, Hoffmann P, Paracchini S, Lowy E, Harold DH, Chapman JA, Klauck SM, Poustka F, Houben RH, Staal WG, Ophoff RA, O’Donovan MC, Williams J, Nöthen MM, Schulte-Körne G, Deloukas P, Ragoussis J, Bailey AJ, Maestrini E, Monaco AP (2010) Characterization of a family with rare deletions in CNTNAP5 and DOCK4 suggests novel risk loci for autism and dyslexia. Biol Psychiat 68(4):320–328
pubmed: 20346443 doi: 10.1016/j.biopsych.2010.02.002
Pejaver V, Urresti J, Lugo-Martinez J, Pagel KA, Lin GN, Nam H-J, Mort M, Cooper DN, Sebat J, Iakoucheva LM, Mooney SD, Radivojac P (2020) Inferring the molecular and phenotypic impact of amino acid variants with MutPred2. Nat Commun 11(1):5918
pubmed: 33219223 pmcid: 7680112 doi: 10.1038/s41467-020-19669-x
Perrault I, Hamdan FF, Rio M, Capo-Chichi J-M, Boddaert N, Décarie J-C, Maranda B, Nabbout R, Sylvain M, Lortie A, Roux PP, Rossignol E, Gérard X, Barcia G, Berquin P, Munnich A, Rouleau GA, Kaplan J, Rozet J-M, Michaud JL (2014) Mutations in DOCK7 in individuals with epileptic encephalopathy and cortical blindness. Am J Hum Genet 94(6):891–897
pubmed: 24814191 pmcid: 4121477 doi: 10.1016/j.ajhg.2014.04.012
Rahimi MJ, Urban N, Wegler M, Sticht H, Schaefer M, Popp B, Gaunitz F, Morleo M, Nigro V, Maitz S, Mancini GMS, Ruivenkamp C, Suk E-K, Bartolomaeus T, Merkenschlager A, Koboldt D, Bartholomew D, Stegmann APA, Sinnema M, Duynisveld I, Salvarinova R, Race S, de Vries BBA, Trimouille A, Naudion S, Marom D, Hamiel U, Henig N, Demurger F, Rahner N, Bartels E, Hamm JA, Putnam AM, Person R, Abou Jamra R, Oppermann H (2022) De novo variants in ATP2B1 lead to neurodevelopmental delay. Am J Hum Genet 109(5):944–952
pubmed: 35358416 pmcid: 9118097 doi: 10.1016/j.ajhg.2022.03.009
Rentzsch P, Schubach M, Shendure J, Kircher M (2021) CADD-Splice-improving genome-wide variant effect prediction using deep learning-derived splice scores. Genome Med 13(1):31
pubmed: 33618777 pmcid: 7901104 doi: 10.1186/s13073-021-00835-9
Richards S, Aziz N, Bale S, Bick D, Das S, Gastier-Foster J, Grody WW, Hegde M, Lyon E, Spector E, Voelkerding K, Rehm HL (2015) Standards and guidelines for the interpretation of sequence variants: a joint consensus recommendation of the American College of Medical Genetics and Genomics and the Association for Molecular Pathology. Genet Med off J Am Coll Med Gen 17(5):405–424
Sheridan E, Wright J, Small N, Corry PC, Oddie S, Whibley C, Petherick ES, Malik T, Pawson N, McKinney PA, Parslow RC (2013) Risk factors for congenital anomaly in a multiethnic birth cohort: an analysis of the Born in Bradford study. Lancet (london, England) 382(9901):1350–1359
pubmed: 23830354 doi: 10.1016/S0140-6736(13)61132-0
Shi L (2013) Dock protein family in brain development and neurological disease. Commun Integr Biol 6(6):e26839
pubmed: 24563715 pmcid: 3922786 doi: 10.4161/cib.26839
Sobreira N, Schiettecatte F, Valle D, Hamosh A (2015) GeneMatcher: a matching tool for connecting investigators with an interest in the same gene. Hum Mutat 36(10):928–930
pubmed: 26220891 pmcid: 4833888 doi: 10.1002/humu.22844
Strande NT, Riggs ER, Buchanan AH, Ceyhan-Birsoy O, DiStefano M, Dwight SS, Goldstein J, Ghosh R, Seifert BA, Sneddon TP, Wright MW, Milko LV, Cherry JM, Giovanni MA, Murray MF, O’Daniel JM, Ramos EM, Santani AB, Scott AF, Plon SE, Rehm HL, Martin CL, Berg JS (2017) Evaluating the clinical validity of gene-disease associations: an evidence-based framework developed by the clinical genome resource. Am J Hum Genet 100(6):895–906
pubmed: 28552198 pmcid: 5473734 doi: 10.1016/j.ajhg.2017.04.015
Sukalo M, Tilsen F, Kayserili H, Müller D, Tüysüz B, Ruddy DM, Wakeling E, Ørstavik KH, Snape KM, Trembath R, de Smedt M, van der Aa N, Skalej M, Mundlos S, Wuyts W, Southgate L, Zenker M (2015) DOCK6 mutations are responsible for a distinct autosomal-recessive variant of Adams-Oliver syndrome associated with brain and eye anomalies. Hum Mutat 36(6):593–598
pubmed: 25824905 doi: 10.1002/humu.22795
Tonikian R, Zhang Y, Sazinsky SL, Currell B, Yeh J-H, Reva B, Held HA, Appleton BA, Evangelista M, Wu Y, Xin X, Chan AC, Seshagiri S, Lasky LA, Sander C, Boone C, Bader GD, Sidhu SS (2008) A specificity map for the PDZ domain family. PLoS Biol 6(9):e239
pubmed: 18828675 pmcid: 2553845 doi: 10.1371/journal.pbio.0060239
Ueda J, Ozawa T, Miyazaki M, Fujiwara Y (1993) SOD-like activity of complexes of nickel(II) ion with some biologically important peptides and their novel reactions with hydrogen peroxide. Inorg Chim Acta 214(1–2):29–32
doi: 10.1016/S0020-1693(00)87522-2
Ueda S, Negishi M, Katoh H (2013) Rac GEF Dock4 interacts with cortactin to regulate dendritic spine formation. Mol Biol Cell 24(10):1602–1613
pubmed: 23536706 pmcid: 3655820 doi: 10.1091/mbc.e12-11-0782
Varadi M, Anyango S, Deshpande M, Nair S, Natassia C, Yordanova G, Yuan D, Stroe O, Wood G, Laydon A, Žídek A, Green T, Tunyasuvunakool K, Petersen S, Jumper J, Clancy E, Green R, Vora A, Lutfi M, Figurnov M, Cowie A, Hobbs N, Kohli P, Kleywegt G, Birney E, Hassabis D, Velankar S (2022) AlphaFold protein structure database: massively expanding the structural coverage of protein-sequence space with high-accuracy models. Nucleic Acids Res 50(D1):D439–D444
pubmed: 34791371 doi: 10.1093/nar/gkab1061
Wiel L, Baakman C, Gilissen D, Veltman JA, Vriend G, Gilissen C (2019) MetaDome: Pathogenicity analysis of genetic variants through aggregation of homologous human protein domains. Hum Mutat 40(8):1030–1038
pubmed: 31116477 pmcid: 6772141
Wiltrout K, Ferrer A, van de Laar I, Namekata K, Harada T, Klee EW, Zimmerman MT, Cousin MA, Kempainen JL, Babovic-Vuksanovic D, van Slegtenhorst MA, Aarts-Tesselaar CD, Schnur RE, Andrews M, Shinawi M (2019) Variants in DOCK3 cause developmental delay and hypotonia. Eur J Hum Genet 27(8):1225–1234
pubmed: 30976111 pmcid: 6777627 doi: 10.1038/s41431-019-0397-2
Xiao Y, Peng Y, Wan J, Tang G, Chen Y, Tang J, Ye W-C, Ip NY, Shi L (2013) The atypical guanine nucleotide exchange factor Dock4 regulates neurite differentiation through modulation of Rac1 GTPase and actin dynamics. J Biol Chem 288(27):20034–20045
pubmed: 23720743 pmcid: 3707701 doi: 10.1074/jbc.M113.458612
Yajnik V, Paulding C, Sordella R, McClatchey AI, Saito M, Wahrer DCR, Reynolds P, Bell DW, Lake R, van den Heuvel S, Settleman J, Haber DA (2003) DOCK4, a GTPase activator, is disrupted during tumorigenesis. Cell 112(5):673–684
pubmed: 12628187 doi: 10.1016/S0092-8674(03)00155-7

Auteurs

Charlotte Herbst (C)

Institute of Human Genetics, University of Leipzig Medical Center, 04103, Leipzig, Germany.

Viktoria Bothe (V)

Institute of Human Genetics, University of Leipzig Medical Center, 04103, Leipzig, Germany.

Meret Wegler (M)

Institute of Human Genetics, University of Leipzig Medical Center, 04103, Leipzig, Germany.

Susanne Axer-Schaefer (S)

Department of Epileptology, Krankenhaus Mara Bethel Epilepsy Center Medical School OWL, Bielefeld University, Campus Bethel, Bielefeld, Germany.

Séverine Audebert-Bellanger (S)

Department of Genetics, CHU Brest, 29000, Brest, France.

Jozef Gecz (J)

Adelaide Medical School and Robinson Research Institute, The University of Adelaide, Adelaide, SA, Australia.

Benjamin Cogne (B)

Service de Génétique Médicale, CHU Nantes, 44000, Nantes, France.
l'institut du Thorax, Nantes Université, CHU Nantes, CNRS, INSERM, 44000, Nantes, France.

Hagit Baris Feldman (HB)

The Genetics Institute and Genomics Center, Tel Aviv Sourasky Medical Center, Tel Aviv, Israel.
Sackler Faculty of Medicine, Tel Aviv University, Tel Aviv, Israel.

Anselm H C Horn (AHC)

Institute of Biochemistry, Friedrich-Alexander-Universität Erlangen-Nürnberg, Erlangen, Germany.
Erlangen National High Performance Computing Center, Friedrich-Alexander-Universität Erlangen-Nürnberg, Erlangen, Germany.

Anna C E Hurst (ACE)

Department of Genetics, University of Alabama at Birmingham, Birmingham, AL, USA.

Melissa A Kelly (MA)

HudsonAlpha Clinical Services Lab, HudsonAlpha Institute for Biotechnology, Huntsville, AL, USA.

Michael C Kruer (MC)

Barrow Neurological Institute, Phoenix Children's Hospital University of Arizona College of Medicine, Phoenix, USA.

Alina Kurolap (A)

The Genetics Institute and Genomics Center, Tel Aviv Sourasky Medical Center, Tel Aviv, Israel.

Annie Laquerriere (A)

Department of Anatomy, Inserm U1245 and CHU Rouen, Univ Rouen Normandie, 76000, Rouen, France.

Megan Li (M)

Invitae Corp, San Francisco, CA, USA.

Paul R Mark (PR)

Division of Medical Genetics, Helen DeVos Children's Hospital, Corewell Health, Grand Rapids, MI, USA.

Markus Morawski (M)

Center of Neuropathology and Brain Research, Medical Faculty, Paul Flechsig Institute, University of Leipzig, Leipzig, Germany.

Mathilde Nizon (M)

Service de Génétique Médicale, CHU Nantes, 44000, Nantes, France.
l'institut du Thorax, Nantes Université, CHU Nantes, CNRS, INSERM, 44000, Nantes, France.

Tomi Pastinen (T)

Genomic Medicine Center, Children's Mercy Hospital, Kansas City, USA.
University of Missouri Kansas City School of Medicine, Kansas City, USA.

Tilman Polster (T)

Department of Epileptology, Krankenhaus Mara Bethel Epilepsy Center Medical School OWL, Bielefeld University, Campus Bethel, Bielefeld, Germany.

Pascale Saugier-Veber (P)

Department of Genetics and Reference Center for Developmental Disorders, Inserm U1245 and CHU Rouen, Univ Rouen Normandie, 76000, Rouen, France.

Jang SeSong (J)

Genomic Medicine Institute, Seoul National University, Seoul, Republic of Korea.

Heinrich Sticht (H)

Institute of Biochemistry, Friedrich-Alexander-Universität Erlangen-Nürnberg, Erlangen, Germany.

Jens T Stieler (JT)

Center of Neuropathology and Brain Research, Medical Faculty, Paul Flechsig Institute, University of Leipzig, Leipzig, Germany.

Isabelle Thifffault (I)

Genomic Medicine Center, Children's Mercy Hospital, Kansas City, USA.
University of Missouri Kansas City School of Medicine, Kansas City, USA.

Clare L van Eyk (CL)

Adelaide Medical School and Robinson Research Institute, The University of Adelaide, Adelaide, SA, Australia.

Pascale Marcorelles (P)

Department of Anatomy, CHU Brest, 29000, Brest, France.

Myriam Vezain-Mouchard (M)

Department of Genetics and Reference Center for Developmental Disorders, Inserm U1245 and CHU Rouen, Univ Rouen Normandie, 76000, Rouen, France.

Rami Abou Jamra (R)

Institute of Human Genetics, University of Leipzig Medical Center, 04103, Leipzig, Germany.

Henry Oppermann (H)

Institute of Human Genetics, University of Leipzig Medical Center, 04103, Leipzig, Germany. henry.oppermann@medizin.uni-leipzig.de.

Classifications MeSH