Diverging co-translational protein complex assembly pathways are governed by interface energy distribution.


Journal

Nature communications
ISSN: 2041-1723
Titre abrégé: Nat Commun
Pays: England
ID NLM: 101528555

Informations de publication

Date de publication:
25 Mar 2024
Historique:
received: 09 10 2023
accepted: 12 03 2024
medline: 26 3 2024
pubmed: 26 3 2024
entrez: 26 3 2024
Statut: epublish

Résumé

Protein-protein interactions are at the heart of all cellular processes, with the ribosome emerging as a platform, orchestrating the nascent-chain interplay dynamics. Here, to study the characteristics governing co-translational protein folding and complex assembly, we combine selective ribosome profiling, imaging, and N-terminomics with all-atoms molecular dynamics. Focusing on conserved N-terminal acetyltransferases (NATs), we uncover diverging co-translational assembly pathways, where highly homologous subunits serve opposite functions. We find that only a few residues serve as "hotspots," initiating co-translational assembly interactions upon exposure at the ribosome exit tunnel. These hotspots are characterized by high binding energy, anchoring the entire interface assembly. Alpha-helices harboring hotspots are highly thermolabile, folding and unfolding during simulations, depending on their partner subunit to avoid misfolding. In vivo hotspot mutations disrupted co-translational complexation, leading to aggregation. Accordingly, conservation analysis reveals that missense NATs variants, causing neurodevelopmental and neurodegenerative diseases, disrupt putative hotspot clusters. Expanding our study to include phosphofructokinase, anthranilate synthase, and nucleoporin subcomplex, we employ AlphaFold-Multimer to model the complexes' complete structures. Computing MD-derived interface energy profiles, we find similar trends. Here, we propose a model based on the distribution of interface energy as a strong predictor of co-translational assembly.

Identifiants

pubmed: 38528060
doi: 10.1038/s41467-024-46881-w
pii: 10.1038/s41467-024-46881-w
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

2638

Subventions

Organisme : Israel Science Foundation (ISF)
ID : 2106/20
Organisme : Israel Science Foundation (ISF)
ID : 1515/23
Organisme : EC | EU Framework Programme for Research and Innovation H2020 | H2020 Priority Excellent Science | H2020 European Research Council (H2020 Excellent Science - European Research Council)
ID : 2031817

Informations de copyright

© 2024. The Author(s).

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Auteurs

Johannes Venezian (J)

Faculty of Biology, Technion Israel institute of Technology, Haifa, Israel.

Hagit Bar-Yosef (H)

Faculty of Biology, Technion Israel institute of Technology, Haifa, Israel.

Hila Ben-Arie Zilberman (H)

Faculty of Biology, Technion Israel institute of Technology, Haifa, Israel.

Noam Cohen (N)

Faculty of Biology, Technion Israel institute of Technology, Haifa, Israel.

Oded Kleifeld (O)

Faculty of Biology, Technion Israel institute of Technology, Haifa, Israel.

Juan Fernandez-Recio (J)

Instituto de Ciencias de la Vid y del Vino (ICVV), CSIC-Universidad de La Rioja-Gobierno de La Rioja, Logroño, Spain.

Fabian Glaser (F)

Lorry I. Lokey Interdisciplinary Center for Life Sciences & Engineering, Haifa, Israel.

Ayala Shiber (A)

Faculty of Biology, Technion Israel institute of Technology, Haifa, Israel. ayalashiber@technion.ac.il.

Classifications MeSH