A systematic review of progranulin concentrations in biofluids in over 7,000 people-assessing the pathogenicity of GRN mutations and other influencing factors.

Frontotemporal dementia Progranulin

Journal

Alzheimer's research & therapy
ISSN: 1758-9193
Titre abrégé: Alzheimers Res Ther
Pays: England
ID NLM: 101511643

Informations de publication

Date de publication:
28 Mar 2024
Historique:
received: 27 10 2023
accepted: 25 02 2024
medline: 28 3 2024
pubmed: 28 3 2024
entrez: 28 3 2024
Statut: epublish

Résumé

Pathogenic heterozygous mutations in the progranulin gene (GRN) are a key cause of frontotemporal dementia (FTD), leading to significantly reduced biofluid concentrations of the progranulin protein (PGRN). This has led to a number of ongoing therapeutic trials aiming to treat this form of FTD by increasing PGRN levels in mutation carriers. However, we currently lack a complete understanding of factors that affect PGRN levels and potential variation in measurement methods. Here, we aimed to address this gap in knowledge by systematically reviewing published literature on biofluid PGRN concentrations. Published data including biofluid PGRN concentration, age, sex, diagnosis and GRN mutation were collected for 7071 individuals from 75 publications. The majority of analyses (72%) had focused on plasma PGRN concentrations, with many of these (56%) measured with a single assay type (Adipogen) and so the influence of mutation type, age at onset, sex, and diagnosis were investigated in this subset of the data. We established a plasma PGRN concentration cut-off between pathogenic mutation carriers and non-carriers of 74.8 ng/mL using the Adipogen assay based on 3301 individuals, with a CSF concentration cut-off of 3.43 ng/mL. Plasma PGRN concentration varied by GRN mutation type as well as by clinical diagnosis in those without a GRN mutation. Plasma PGRN concentration was significantly higher in women than men in GRN mutation carriers (p = 0.007) with a trend in non-carriers (p = 0.062), and there was a significant but weak positive correlation with age in both GRN mutation carriers and non-carriers. No significant association was seen with weight or with TMEM106B rs1990622 genotype. However, higher plasma PGRN levels were seen in those with the GRN rs5848 CC genotype in both GRN mutation carriers and non-carriers. These results further support the usefulness of PGRN concentration for the identification of the large majority of pathogenic mutations in the GRN gene. Furthermore, these results highlight the importance of considering additional factors, such as mutation type, sex and age when interpreting PGRN concentrations. This will be particularly important as we enter the era of trials for progranulin-associated FTD.

Sections du résumé

BACKGROUND BACKGROUND
Pathogenic heterozygous mutations in the progranulin gene (GRN) are a key cause of frontotemporal dementia (FTD), leading to significantly reduced biofluid concentrations of the progranulin protein (PGRN). This has led to a number of ongoing therapeutic trials aiming to treat this form of FTD by increasing PGRN levels in mutation carriers. However, we currently lack a complete understanding of factors that affect PGRN levels and potential variation in measurement methods. Here, we aimed to address this gap in knowledge by systematically reviewing published literature on biofluid PGRN concentrations.
METHODS METHODS
Published data including biofluid PGRN concentration, age, sex, diagnosis and GRN mutation were collected for 7071 individuals from 75 publications. The majority of analyses (72%) had focused on plasma PGRN concentrations, with many of these (56%) measured with a single assay type (Adipogen) and so the influence of mutation type, age at onset, sex, and diagnosis were investigated in this subset of the data.
RESULTS RESULTS
We established a plasma PGRN concentration cut-off between pathogenic mutation carriers and non-carriers of 74.8 ng/mL using the Adipogen assay based on 3301 individuals, with a CSF concentration cut-off of 3.43 ng/mL. Plasma PGRN concentration varied by GRN mutation type as well as by clinical diagnosis in those without a GRN mutation. Plasma PGRN concentration was significantly higher in women than men in GRN mutation carriers (p = 0.007) with a trend in non-carriers (p = 0.062), and there was a significant but weak positive correlation with age in both GRN mutation carriers and non-carriers. No significant association was seen with weight or with TMEM106B rs1990622 genotype. However, higher plasma PGRN levels were seen in those with the GRN rs5848 CC genotype in both GRN mutation carriers and non-carriers.
CONCLUSIONS CONCLUSIONS
These results further support the usefulness of PGRN concentration for the identification of the large majority of pathogenic mutations in the GRN gene. Furthermore, these results highlight the importance of considering additional factors, such as mutation type, sex and age when interpreting PGRN concentrations. This will be particularly important as we enter the era of trials for progranulin-associated FTD.

Identifiants

pubmed: 38539243
doi: 10.1186/s13195-024-01420-z
pii: 10.1186/s13195-024-01420-z
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

66

Subventions

Organisme : JPND GENFI-PROX grant
ID : 2019-02248
Organisme : JPND GENFI-PROX grant
ID : 2019-02248
Organisme : JPND GENFI-PROX grant
ID : 2019-02248
Organisme : JPND GENFI-PROX grant
ID : 2019-02248
Organisme : JPND GENFI-PROX grant
ID : 2019-02248
Organisme : JPND GENFI-PROX grant
ID : 2019-02248
Organisme : MRC Clinician Scientist Fellowship
ID : MR/M008525/1
Organisme : MRC Clinician Scientist Fellowship
ID : MR/M008525/1
Organisme : FAPESP grant number
ID : 2013/017584
Organisme : EU Joint Programme-Neurodegenerative Diseases networks Genfi-Prox
ID : 01ED2008A
Organisme : bPride
ID : 01ED2001
Organisme : German Federal Ministry of Education and Research
ID : FTLDc 01GI1007A, Moodmarker 01EW200
Organisme : the EU
ID : MIRIADE 860197, FAIR-PARK II 633190
Organisme : German Research Foundation/DFG
ID : SFB1279
Organisme : The foundation of the state Baden-Württemberg
ID : D.3830
Organisme : Boehringer Ingelheim Ulm University BioCenter and the Thierry Latran Foundation
ID : D.5009
Organisme : Swedish State Support for Clinical Research
ID : #ALFGBG-71320
Organisme : Alzheimer Drug Discovery Foundation (ADDF), USA
ID : #201809-2016862
Organisme : AD Strategic Fund and the Alzheimer's Association
ID : #ADSF-21-831376-C, #ADSF-21-831381-C, #ADSF-21-831377-C
Organisme : Swedish Research Council
ID : #2019-02397, #2022-01018
Organisme : the Bluefield Project, the Olav Thon Foundation, the Erling-Persson Family Foundation, Stiftelsen för Gamla Tjänarinnor, Hjärnfonden, Sweden
ID : #FO2022-0270
Organisme : the European Union Joint Programme - Neurodegenerative Disease Research
ID : JPND2021-00694)
Organisme : UK Dementia Research Institute at UCL
ID : UKDRI-1003
Organisme : Alzheimer's Research UK
ID : ARUK-RADF2021A-003
Organisme : NIHR Rare Disease Translational Research Collaboration
ID : BRC149/NS/MH
Organisme : MRC UK GENFI grant
ID : MR/M023664/1

Informations de copyright

© 2024. The Author(s).

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Auteurs

Imogen J Swift (IJ)

Department of Neurodegenerative Disease, Dementia Research Institute, UCL Institute of Neurology, Queen Square, London, UK.
Department of Neurodegenerative Disease, Dementia Research Centre, UCL Institute of Neurology, Queen Square, London, WC1N 3BG, UK.

Rosa Rademakers (R)

Department of Neuroscience, Mayo Clinic, Jacksonville, FL, USA.
VIB Center for Molecular Neurology, VIB, Antwerp, Belgium.
Department of Biomedical Sciences, University of Antwerp, Antwerp, Belgium.

NiCole Finch (N)

Department of Neuroscience, Mayo Clinic, Jacksonville, FL, USA.

Matt Baker (M)

Department of Neuroscience, Mayo Clinic, Jacksonville, FL, USA.

Roberta Ghidoni (R)

Molecular Markers Laboratory, IRCCS Istituto Centro San Giovanni Di Dio Fatebenefratelli, Brescia, Italy.

Luisa Benussi (L)

Molecular Markers Laboratory, IRCCS Istituto Centro San Giovanni Di Dio Fatebenefratelli, Brescia, Italy.

Giuliano Binetti (G)

MAC-Memory Clinic and Molecular Markers Laboratory, IRCCS Istituto Centro San Giovanni Di Dio Fatebenefratelli, Brescia, Italy.

Giacomina Rossi (G)

Unit of Neurology V and Neuropathology, Fondazione IRCCS Istituto Neurologico Carlo Besta, Milan, Italy.

Matthis Synofzik (M)

Division Translational Genomics of Neurodegenerative Diseases, Hertie-Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany.
German Center of Neurodegenerative Diseases (DZNE), Tübingen, Germany.

Carlo Wilke (C)

Division Translational Genomics of Neurodegenerative Diseases, Hertie-Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany.
German Center of Neurodegenerative Diseases (DZNE), Tübingen, Germany.

David Mengel (D)

Division Translational Genomics of Neurodegenerative Diseases, Hertie-Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany.
German Center of Neurodegenerative Diseases (DZNE), Tübingen, Germany.

Caroline Graff (C)

Department of Neurobiology, Care Sciences and Society, Center for Alzheimer Research, Division of Neurogeriatrics, BioclinicumKarolinska Institutet, Solna, Sweden.
Unit for Hereditary Dementias, Theme Inflammation and Aging, Karolinska University Hospital, Solna, Sweden.

Leonel T Takada (LT)

Department of Neurology, Hospital das Clinicas, University of São Paulo Medical School, São Paulo, Brazil.

Raquel Sánchez-Valle (R)

Alzheimer's Disease and Other Cognitive Disorders Unit, FRCB-IDIBAPS, Institut de Neurociències, Neurology Service, Hospital Clínic de Barcelona, Universitat de Barcelona (UB), 08036, Barcelona, Spain.

Anna Antonell (A)

Alzheimer's Disease and Other Cognitive Disorders Unit, FRCB-IDIBAPS, Institut de Neurociències, Neurology Service, Hospital Clínic de Barcelona, Universitat de Barcelona (UB), 08036, Barcelona, Spain.

Daniela Galimberti (D)

Dept. of Biomedical, Surgical and Dental Sciences, University of Milan, Milan, Italy.
Neurodegerative Diseases Center, Fondazione IRCCS Ca' Granda, Ospedale Maggiore Policlinico, Milan, Italy.

Chiara Fenoglio (C)

Dept. of Biomedical, Surgical and Dental Sciences, University of Milan, Milan, Italy.
Neurodegerative Diseases Center, Fondazione IRCCS Ca' Granda, Ospedale Maggiore Policlinico, Milan, Italy.

Maria Serpente (M)

Neurodegerative Diseases Center, Fondazione IRCCS Ca' Granda, Ospedale Maggiore Policlinico, Milan, Italy.

Marina Arcaro (M)

Neurodegerative Diseases Center, Fondazione IRCCS Ca' Granda, Ospedale Maggiore Policlinico, Milan, Italy.

Stefanie Schreiber (S)

Department of Neurology, Otto Von Guericke University, Magdeburg, Germany.

Stefan Vielhaber (S)

Department of Neurology, Otto Von Guericke University, Magdeburg, Germany.

Philipp Arndt (P)

Department of Neurology, Otto Von Guericke University, Magdeburg, Germany.

Isabel Santana (I)

Neurology Department, Centro Hospitalar E Universitário de Coimbra, Coimbra, Portugal.
Center for Neuroscience and Cell Biology, Centre for Innovative Biomedicine and Biotechnology, University of Coimbra, Coimbra, Portugal.
Faculty of Medicine, University of Coimbra, Coimbra, Portugal.

Maria Rosario Almeida (MR)

Faculty of Medicine, University of Coimbra, Coimbra, Portugal.

Fermín Moreno (F)

Cognitive Disorders Unit, Department of Neurology, Donostia University Hospital, San Sebastian, Gipuzkoa, Spain.
Neuroscience Area, Biodonostia Health Research Insitute, San Sebastian, Gipuzkoa, Spain.

Myriam Barandiaran (M)

Cognitive Disorders Unit, Department of Neurology, Donostia University Hospital, San Sebastian, Gipuzkoa, Spain.
Neuroscience Area, Biodonostia Health Research Insitute, San Sebastian, Gipuzkoa, Spain.

Alazne Gabilondo (A)

Cognitive Disorders Unit, Department of Neurology, Donostia University Hospital, San Sebastian, Gipuzkoa, Spain.
Neuroscience Area, Biodonostia Health Research Insitute, San Sebastian, Gipuzkoa, Spain.

Johannes Stubert (J)

Department of Obstetrics and Gynecology, Rostock University Medical Center, Rostock, Germany.

Estrella Gómez-Tortosa (E)

Department of Neurology, Fundación Jiménez Díaz, Madrid, Spain.

Pablo Agüero (P)

Department of Neurology, Fundación Jiménez Díaz, Madrid, Spain.

M José Sainz (MJ)

Department of Neurology, Fundación Jiménez Díaz, Madrid, Spain.

Tomohito Gohda (T)

Department of Nephrology, Faculty of Medicine, Juntendo University, Tokyo, Japan.

Maki Murakoshi (M)

Department of Nephrology, Faculty of Medicine, Juntendo University, Tokyo, Japan.

Nozomu Kamei (N)

Department of Endocrinology and Metabolism, Hiroshima Red Cross Hospital & Atomicbomb Survivors Hospital, Hiroshima, Japan.
Institute for Clinical Research, National Hospital Organization, Kure Medical Center and Chugoku Cancer Center, Hiroshima, Japan.

Sarah Kittel-Schneider (S)

Department of Psychiatry, University College Cork, Cork, Ireland.
Department of Psychiatry, Psychotherapy and Psychosomatic Medicine, University Hospital Würzburg, Würzburg, Germany.

Andreas Reif (A)

Department of Psychiatry, Psychotherapy and Psychosomatic Medicine, University Hospital Frankfurt, Frankfurt, Germany.

Johannes Weigl (J)

Department of Psychiatry, Psychotherapy and Psychosomatic Medicine, University Hospital Würzburg, Würzburg, Germany.
Department of Psychiatry, Hospital in Tauberbischofsheim, Tauberbischofsheim, Germany.

Jinlong Jian (J)

University of Pennsylvania, Gene Therapy Program, Philadelphia, USA.

Chuanju Liu (C)

Department of Orthopaedics & Rehabilitation, Yale University School of Medicine, New Haven, CT, USA.

Ginette Serrero (G)

A&G Pharmaceutical Inc, Columbia, MD, USA.
Program in Oncology, University of Maryland Greenebaum Comprehensive Cancer Center, Baltimore, MD, USA.

Thomas Greither (T)

Center for Reproductive Medicine and Andrology, Martin Luther University Halle-Wittenberg, Halle, Germany.

Gerit Theil (G)

Department of Urology, Martin Luther University Halle-Wittenberg, Halle, Germany.

Ebba Lohmann (E)

Department of Neurodegenerative Diseases, Hertie Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany.
DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany.

Stefano Gazzina (S)

Department of Neurological and Vision Sciences, Neurophysiology Unit, ASST SpedaliCivili, Brescia, Italy.

Silvia Bagnoli (S)

Department of Neurological and Psychiatric Sciences, University of Florence, Viale Morgagni, 85, 50134, Florence, Italy.

Giovanni Coppola (G)

Department of Neurology, University of California, Los Angeles, California, USA.
Department of Psychiatry Semel Institute for Neuroscience and Human Behavior, University of California, Los Angeles, California, USA.

Amalia Bruni (A)

Regional Neurogenetic Centre, ASPCZ, Lamezia Terme, Italy.

Mirja Quante (M)

Department of Neonatology, Tuebingen University Hospital, Tuebingen, Germany.

Wieland Kiess (W)

Leipzig Research Center for Civilization Diseases - LIFE, University of Leipzig, Leipzig, Germany.
Hospital for Children and Adolescents, University of Leipzig, Leipzig, Germany.
Center of Pediatric Research (CPL), University of Leipzig, Leipzig, Germany.

Andreas Hiemisch (A)

Leipzig Research Center for Civilization Diseases - LIFE, University of Leipzig, Leipzig, Germany.
Center of Pediatric Research (CPL), University of Leipzig, Leipzig, Germany.

Anne Jurkutat (A)

Leipzig Research Center for Civilization Diseases - LIFE, University of Leipzig, Leipzig, Germany.

Matthew S Block (MS)

Department of Oncology, Mayo Clinic, Rochester, MN, USA.

Aaron M Carlson (AM)

Department of Neurology, University of Colorado Anschutz Medical Campus, Denver, CO, USA.

Geir Bråthen (G)

Department of Neurology and Clinical Neurophysiology, Trondheim University Hospital, Trondheim, Norway.
Department of Neuromedicine and Movement Science, Faculty of Medicine and Health Sciences, NTNU. , Trondheim, Norway.

Sigrid Botne Sando (SB)

Department of Neurology and Clinical Neurophysiology, Trondheim University Hospital, Trondheim, Norway.
Department of Neuromedicine and Movement Science, Faculty of Medicine and Health Sciences, NTNU. , Trondheim, Norway.

Gøril Rolfseng Grøntvedt (GR)

Department of Neurology and Clinical Neurophysiology, Trondheim University Hospital, Trondheim, Norway.
Department of Neuromedicine and Movement Science, Faculty of Medicine and Health Sciences, NTNU. , Trondheim, Norway.

Camilla Lauridsen (C)

Department of Research, Trondheim University Hospital, Trondheim, Norway.

Amanda Heslegrave (A)

Department of Neurodegenerative Disease, Dementia Research Institute, UCL Institute of Neurology, Queen Square, London, UK.

Carolin Heller (C)

Department of Neurodegenerative Disease, Dementia Research Institute, UCL Institute of Neurology, Queen Square, London, UK.
Department of Neurodegenerative Disease, Dementia Research Centre, UCL Institute of Neurology, Queen Square, London, WC1N 3BG, UK.

Emily Abel (E)

Department of Neurodegenerative Disease, Dementia Research Institute, UCL Institute of Neurology, Queen Square, London, UK.

Alba Gómez-Núñez (A)

Alzheimer's Disease and Other Cognitive Disorders Unit, FRCB-IDIBAPS, Institut de Neurociències, Neurology Service, Hospital Clínic de Barcelona, Universitat de Barcelona (UB), 08036, Barcelona, Spain.

Roger Puey (R)

Alzheimer's Disease and Other Cognitive Disorders Unit, FRCB-IDIBAPS, Institut de Neurociències, Neurology Service, Hospital Clínic de Barcelona, Universitat de Barcelona (UB), 08036, Barcelona, Spain.

Andrea Arighi (A)

Neurodegerative Diseases Center, Fondazione IRCCS Ca' Granda, Ospedale Maggiore Policlinico, Milan, Italy.

Enmanuela Rotondo (E)

Neurodegerative Diseases Center, Fondazione IRCCS Ca' Granda, Ospedale Maggiore Policlinico, Milan, Italy.

Lize C Jiskoot (LC)

Department of Neurology and Alzheimer Center Erasmus MC, Erasmus MC University Medical Center, Rotterdam, Netherlands.

Lieke H H Meeter (LHH)

Department of Neurology and Alzheimer Center Erasmus MC, Erasmus MC University Medical Center, Rotterdam, Netherlands.

João Durães (J)

Neurology Department, Centro Hospitalar E Universitário de Coimbra, Coimbra, Portugal.

Marisa Lima (M)

Neurology Department, Centro Hospitalar E Universitário de Coimbra, Coimbra, Portugal.

Miguel Tábuas-Pereira (M)

Neurology Department, Centro Hospitalar E Universitário de Coimbra, Coimbra, Portugal.
Faculty of Medicine, University of Coimbra, Coimbra, Portugal.

João Lemos (J)

Faculty of Medicine, University of Coimbra, Coimbra, Portugal.

Bradley Boeve (B)

Department of Neurology, Mayo Clinic, Rochester, MN, USA.

Ronald C Petersen (RC)

Department of Neurology, Mayo Clinic, Rochester, MN, USA.

Dennis W Dickson (DW)

Department of Neuroscience, Mayo Clinic, Jacksonville, FL, USA.

Neill R Graff-Radford (NR)

Department of Neurology, Mayo Clinic, Jacksonville, FL, USA.

Isabelle LeBer (I)

Sorbonne UniversitéInserm U1127, CNRS UMR 7225, Institut du Cerveau Et La Moelle Épinière (ICM), AP-HP - Hôpital Pitié-Salpêtrière, Paris, France.

Leila Sellami (L)

Sorbonne UniversitéInserm U1127, CNRS UMR 7225, Institut du Cerveau Et La Moelle Épinière (ICM), AP-HP - Hôpital Pitié-Salpêtrière, Paris, France.
Centre de Référence Des Démences Rares Ou Précoces, IM2A, Département de Neurologie, AP-HP - Hôpital Pitié-Salpêtrière, Paris, France.

Foudil Lamari (F)

UF de Biochimie Des Maladies Neurométaboliques Et Neurodégénératives, Service de Biochimie Métabolique, AP-HP - Hôpital Pitié-Salpêtrière, Paris, France.

Fabienne Clot (F)

UF de Neurogénétique Moléculaire Et Cellulaire, Département de Génétique, AP-HP, Hôpitaux Universitaires La Pitié Salpêtrière-Charles Foix, Paris, France.

Barbara Borroni (B)

Department of Clinical and Experimental Sciences, University of Brescia, Brescia, Italy.

Valentina Cantoni (V)

Department of Clinical and Experimental Sciences, University of Brescia, Brescia, Italy.

Jasmine Rivolta (J)

Department of Clinical and Experimental Sciences, University of Brescia, Brescia, Italy.

Alberto Lleó (A)

Neurology Department. Hospital Sant Pau, Memory Unit, Barcelona, Spain.
Centro de Investigación Biomédica en Red en Enfermedades Neurodegenerativas (CIBERNED), 28031, Madrid, Spain.
Autonomous University of Barcelona, 08913, Barcelona, Spain.

Juan Fortea (J)

Neurology Department. Hospital Sant Pau, Memory Unit, Barcelona, Spain.
Centro de Investigación Biomédica en Red en Enfermedades Neurodegenerativas (CIBERNED), 28031, Madrid, Spain.
Autonomous University of Barcelona, 08913, Barcelona, Spain.

Daniel Alcolea (D)

Neurology Department. Hospital Sant Pau, Memory Unit, Barcelona, Spain.
Centro de Investigación Biomédica en Red en Enfermedades Neurodegenerativas (CIBERNED), 28031, Madrid, Spain.
Autonomous University of Barcelona, 08913, Barcelona, Spain.

Ignacio Illán-Gala (I)

Neurology Department. Hospital Sant Pau, Memory Unit, Barcelona, Spain.
Centro de Investigación Biomédica en Red en Enfermedades Neurodegenerativas (CIBERNED), 28031, Madrid, Spain.
Autonomous University of Barcelona, 08913, Barcelona, Spain.

Lucie Andres-Cerezo (L)

Institute of Rheumatology, Na Slupi 4, 12850, Prague 2, Prague, Czech Republic.

Philip Van Damme (P)

Laboratory of Neurobiology, Flanders Interuniversity Institute for Biotechnology, Katholieke Universiteit Leuven, Campus Gasthuisberg, 3000, Louvain, Belgium.

Jordi Clarimon (J)

Centro de Investigación Biomédica en Red Sobre Enfermedades Neurodegenerativas (CIBERNED), Madrid, Spain.
Memory Unit, Department of Neurology, Institut d'Investigacions Biomèdiques Sant Pau - Hospital de Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain.

Petra Steinacker (P)

Department of Neurology, Martin-Luther University Halle-Wittenberg, University Clinic Halle, Halle (Saale), Germany.

Emily Feneberg (E)

Department of Neurology, Klinikum Rechts Der Isar, Technical University of Munich, Munich, Germany.

Markus Otto (M)

Department of Neurology, Martin-Luther University Halle-Wittenberg, University Clinic Halle, Halle (Saale), Germany.

Emma L van der Ende (EL)

Department of Neurology and Alzheimer Center Erasmus MC, Erasmus MC University Medical Center, Rotterdam, Netherlands.

John C van Swieten (JC)

Department of Neurology and Alzheimer Center Erasmus MC, Erasmus MC University Medical Center, Rotterdam, Netherlands.

Harro Seelaar (H)

Department of Neurology and Alzheimer Center Erasmus MC, Erasmus MC University Medical Center, Rotterdam, Netherlands.

Henrik Zetterberg (H)

Department of Neurodegenerative Disease, Dementia Research Institute, UCL Institute of Neurology, Queen Square, London, UK.
UK Dementia Research Institute at University College London, UCL Queen Square Institute of Neurology, University College London, London, UK.
Institute of Neuroscience and Physiology, Sahlgrenska Academy at the University of Gothenburg, 43180, Mölndal, Sweden.
Department of Psychiatry and Neurochemistry, Institute of Neuroscience and Physiology, Sahlgrenska Academy at the University of Gothenburg, Mölndal, Sweden.
Clinical Neurochemistry Laboratory, Sahlgrenska University Hospital, Mölndal, Sweden.
Hong Kong Center for Neurodegenerative Diseases, Hong Kong, China.

Aitana Sogorb-Esteve (A)

Department of Neurodegenerative Disease, Dementia Research Institute, UCL Institute of Neurology, Queen Square, London, UK.
Department of Neurodegenerative Disease, Dementia Research Centre, UCL Institute of Neurology, Queen Square, London, WC1N 3BG, UK.

Jonathan D Rohrer (JD)

Department of Neurodegenerative Disease, Dementia Research Centre, UCL Institute of Neurology, Queen Square, London, WC1N 3BG, UK. j.rohrer@ucl.ac.uk.

Classifications MeSH