Mitovesicles secreted into the extracellular space of brains with mitochondrial dysfunction impair synaptic plasticity.

Alzheimer’s disease Down syndrome Exosome Extracellular vesicle Long-term potentiation MAO-B Microvesicle Mitochondria Mitovesicle Neurodegenerative disease

Journal

Molecular neurodegeneration
ISSN: 1750-1326
Titre abrégé: Mol Neurodegener
Pays: England
ID NLM: 101266600

Informations de publication

Date de publication:
14 Apr 2024
Historique:
received: 31 08 2023
accepted: 18 03 2024
medline: 15 4 2024
pubmed: 15 4 2024
entrez: 14 4 2024
Statut: epublish

Résumé

Hypometabolism tied to mitochondrial dysfunction occurs in the aging brain and in neurodegenerative disorders, including in Alzheimer's disease, in Down syndrome, and in mouse models of these conditions. We have previously shown that mitovesicles, small extracellular vesicles (EVs) of mitochondrial origin, are altered in content and abundance in multiple brain conditions characterized by mitochondrial dysfunction. However, given their recent discovery, it is yet to be explored what mitovesicles regulate and modify, both under physiological conditions and in the diseased brain. In this study, we investigated the effects of mitovesicles on synaptic function, and the molecular players involved. Hippocampal slices from wild-type mice were perfused with the three known types of EVs, mitovesicles, microvesicles, or exosomes, isolated from the brain of a mouse model of Down syndrome or of a diploid control and long-term potentiation (LTP) recorded. The role of the monoamine oxidases type B (MAO-B) and type A (MAO-A) in mitovesicle-driven LTP impairments was addressed by treatment of mitovesicles with the irreversible MAO inhibitors pargyline and clorgiline prior to perfusion of the hippocampal slices. Mitovesicles from the brain of the Down syndrome model reduced LTP within minutes of mitovesicle addition. Mitovesicles isolated from control brains did not trigger electrophysiological effects, nor did other types of brain EVs (microvesicles and exosomes) from any genotype tested. Depleting mitovesicles of their MAO-B, but not MAO-A, activity eliminated their ability to alter LTP. Mitovesicle impairment of LTP is a previously undescribed paracrine-like mechanism by which EVs modulate synaptic activity, demonstrating that mitovesicles are active participants in the propagation of cellular and functional homeostatic changes in the context of neurodegenerative disorders.

Sections du résumé

BACKGROUND BACKGROUND
Hypometabolism tied to mitochondrial dysfunction occurs in the aging brain and in neurodegenerative disorders, including in Alzheimer's disease, in Down syndrome, and in mouse models of these conditions. We have previously shown that mitovesicles, small extracellular vesicles (EVs) of mitochondrial origin, are altered in content and abundance in multiple brain conditions characterized by mitochondrial dysfunction. However, given their recent discovery, it is yet to be explored what mitovesicles regulate and modify, both under physiological conditions and in the diseased brain. In this study, we investigated the effects of mitovesicles on synaptic function, and the molecular players involved.
METHODS METHODS
Hippocampal slices from wild-type mice were perfused with the three known types of EVs, mitovesicles, microvesicles, or exosomes, isolated from the brain of a mouse model of Down syndrome or of a diploid control and long-term potentiation (LTP) recorded. The role of the monoamine oxidases type B (MAO-B) and type A (MAO-A) in mitovesicle-driven LTP impairments was addressed by treatment of mitovesicles with the irreversible MAO inhibitors pargyline and clorgiline prior to perfusion of the hippocampal slices.
RESULTS RESULTS
Mitovesicles from the brain of the Down syndrome model reduced LTP within minutes of mitovesicle addition. Mitovesicles isolated from control brains did not trigger electrophysiological effects, nor did other types of brain EVs (microvesicles and exosomes) from any genotype tested. Depleting mitovesicles of their MAO-B, but not MAO-A, activity eliminated their ability to alter LTP.
CONCLUSIONS CONCLUSIONS
Mitovesicle impairment of LTP is a previously undescribed paracrine-like mechanism by which EVs modulate synaptic activity, demonstrating that mitovesicles are active participants in the propagation of cellular and functional homeostatic changes in the context of neurodegenerative disorders.

Identifiants

pubmed: 38616258
doi: 10.1186/s13024-024-00721-z
pii: 10.1186/s13024-024-00721-z
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

34

Subventions

Organisme : NIA NIH HHS
ID : AG017617
Pays : United States
Organisme : NIA NIH HHS
ID : AG056732
Pays : United States
Organisme : NIA NIH HHS
ID : AG057517
Pays : United States
Organisme : NIDA NIH HHS
ID : DA044489
Pays : United States
Organisme : NIH HHS
ID : R01NS110024
Pays : United States
Organisme : Alzheimer's Association
ID : AARF-22-923826
Pays : United States

Informations de copyright

© 2024. The Author(s).

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Auteurs

Pasquale D'Acunzo (P)

Center for Dementia Research, Nathan S. Kline Institute for Psychiatric Research, 10962, Orangeburg, NY, USA.
Department of Psychiatry, New York University Grossman School of Medicine, 10016, New York, NY, USA.

Elentina K Argyrousi (EK)

Department of Pathology and Cell Biology, Taub Institute for Research on Alzheimer's Disease and the Aging Brain, Columbia University, 10027, New York, NY, USA.
Department of Medicine, Columbia University, 10027, New York, NY, USA.

Jonathan M Ungania (JM)

Center for Dementia Research, Nathan S. Kline Institute for Psychiatric Research, 10962, Orangeburg, NY, USA.

Yohan Kim (Y)

Center for Dementia Research, Nathan S. Kline Institute for Psychiatric Research, 10962, Orangeburg, NY, USA.
Department of Psychiatry, New York University Grossman School of Medicine, 10016, New York, NY, USA.

Steven DeRosa (S)

Center for Dementia Research, Nathan S. Kline Institute for Psychiatric Research, 10962, Orangeburg, NY, USA.

Monika Pawlik (M)

Center for Dementia Research, Nathan S. Kline Institute for Psychiatric Research, 10962, Orangeburg, NY, USA.

Chris N Goulbourne (CN)

Center for Dementia Research, Nathan S. Kline Institute for Psychiatric Research, 10962, Orangeburg, NY, USA.

Ottavio Arancio (O)

Department of Pathology and Cell Biology, Taub Institute for Research on Alzheimer's Disease and the Aging Brain, Columbia University, 10027, New York, NY, USA.
Department of Medicine, Columbia University, 10027, New York, NY, USA.

Efrat Levy (E)

Center for Dementia Research, Nathan S. Kline Institute for Psychiatric Research, 10962, Orangeburg, NY, USA. Efrat.Levy@NKI.rfmh.org.
Department of Psychiatry, New York University Grossman School of Medicine, 10016, New York, NY, USA. Efrat.Levy@NKI.rfmh.org.
Department of Biochemistry & Molecular Pharmacology, New York University Grossman School of Medicine, 10027, New York, NY, USA. Efrat.Levy@NKI.rfmh.org.
NYU Neuroscience Institute, New York University Grossman School of Medicine, 10016, New York, NY, USA. Efrat.Levy@NKI.rfmh.org.

Classifications MeSH