The immune response to RNA suppresses nucleic acid synthesis by limiting ribose 5-phosphate.

Antiviral Immunity Metabolism Nucleotide Biosynthesis PKR (Protein Kinase RNA-Activated) PPP (Pentose Phosphate Pathway) RPIA (Ribose 5-Phosphate Isomerase A)

Journal

The EMBO journal
ISSN: 1460-2075
Titre abrégé: EMBO J
Pays: England
ID NLM: 8208664

Informations de publication

Date de publication:
22 May 2024
Historique:
received: 06 09 2023
accepted: 19 03 2024
revised: 29 02 2024
medline: 23 5 2024
pubmed: 23 5 2024
entrez: 22 5 2024
Statut: aheadofprint

Résumé

During infection viruses hijack host cell metabolism to promote their replication. Here, analysis of metabolite alterations in macrophages exposed to poly I:C recognises that the antiviral effector Protein Kinase RNA-activated (PKR) suppresses glucose breakdown within the pentose phosphate pathway (PPP). This pathway runs parallel to central glycolysis and is critical to producing NADPH and pentose precursors for nucleotides. Changes in metabolite levels between wild-type and PKR-ablated macrophages show that PKR controls the generation of ribose 5-phosphate, in a manner distinct from its established function in gene expression but dependent on its kinase activity. PKR phosphorylates and inhibits the Ribose 5-Phosphate Isomerase A (RPIA), thereby preventing interconversion of ribulose- to ribose 5-phosphate. This activity preserves redox control but decreases production of ribose 5-phosphate for nucleotide biosynthesis. Accordingly, the PKR-mediated immune response to RNA suppresses nucleic acid production. In line, pharmacological targeting of the PPP during infection decreases the replication of the Herpes simplex virus. These results identify an immune response-mediated control of host cell metabolism and suggest targeting the RPIA as a potential innovative antiviral treatment.

Identifiants

pubmed: 38778156
doi: 10.1038/s44318-024-00100-w
pii: 10.1038/s44318-024-00100-w
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Subventions

Organisme : NHMRC
ID : 1143839
Organisme : NHMRC
ID : 1043398
Organisme : Perpetual Tructees
ID : CF07/2408
Organisme : Endeavour Fund
ID : UOOX1904

Informations de copyright

© 2024. The Author(s).

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Auteurs

Pushpak Bhattacharjee (P)

Centre for Innate Immunity and Infectious Diseases, Hudson Institute of Medical Research and Department of Molecular and Translational Sciences, Monash University, Clayton, VIC, 3168, Australia.

Die Wang (D)

Centre for Innate Immunity and Infectious Diseases, Hudson Institute of Medical Research and Department of Molecular and Translational Sciences, Monash University, Clayton, VIC, 3168, Australia.

Dovile Anderson (D)

Drug Delivery, Disposition and Dynamics, Monash Institute of Pharmaceutical Sciences, Monash University, Parkville, VIC, 3052, Australia.

Joshua N Buckler (JN)

Ferrier Research Institute, Victoria University of Wellington, Lower Hutt, 5010, New Zealand.

Eveline de Geus (E)

Centre for Innate Immunity and Infectious Diseases, Hudson Institute of Medical Research and Department of Molecular and Translational Sciences, Monash University, Clayton, VIC, 3168, Australia.

Feng Alex Yan (FA)

Australian Centre for Blood Diseases, Department of Clinical Hematology, Monash University, Clayton, VIC, 3004, Australia.

Galina Polekhina (G)

Department of Epidemiology & Preventive Medicine, Monash University, Melbourne, VIC, 3004, Australia.

Ralf Schittenhelm (R)

Monash Proteomics & Metabolomics Facility, Department of Biochemistry and Molecular Biology, Biomedicine Discovery Institute, Monash University, Clayton, VIC, 3800, Australia.

Darren J Creek (DJ)

Drug Delivery, Disposition and Dynamics, Monash Institute of Pharmaceutical Sciences, Monash University, Parkville, VIC, 3052, Australia.

Lawrence D Harris (LD)

Ferrier Research Institute, Victoria University of Wellington, Lower Hutt, 5010, New Zealand.

Anthony J Sadler (AJ)

Centre for Innate Immunity and Infectious Diseases, Hudson Institute of Medical Research and Department of Molecular and Translational Sciences, Monash University, Clayton, VIC, 3168, Australia. anthony.sadler@hudson.org.au.

Classifications MeSH