Assessment of fecal bacterial viability and diversity in fresh and frozen fecal microbiota transplant (FMT) product in horses.


Journal

BMC veterinary research
ISSN: 1746-6148
Titre abrégé: BMC Vet Res
Pays: England
ID NLM: 101249759

Informations de publication

Date de publication:
10 Jul 2024
Historique:
received: 13 03 2023
accepted: 01 07 2024
medline: 11 7 2024
pubmed: 11 7 2024
entrez: 10 7 2024
Statut: epublish

Résumé

Currently, lack of standardization for fecal microbiota transplantation (FMT) in equine practice has resulted in highly variable techniques, and there is no data on the bacterial metabolic activity or viability of the administered product. The objectives of this study were to compare the total and potentially metabolically active bacterial populations in equine FMT, and assess the effect of different frozen storage times, buffers, and temperatures on an equine FMT product. Fresh feces collected from three healthy adult horses was subjected to different storage methods. This included different preservation solutions (saline plus glycerol or saline only), temperature (-20 °C or -80 °C), and time (fresh, 30, 60, or 90 days). Samples underwent DNA extraction to assess total bacterial populations (both live and dead combined) and RNA extraction followed by reverse transcription to cDNA as a proxy to assess viable bacteria, then 16s rRNA gene amplicon sequencing using the V1-V2 region. The largest difference in population indices and taxonomic composition at the genus level was seen when evaluating the results of DNA-based (total) and cDNA-based (potentially metabolically active) extraction method. At the community level, alpha diversity (observed species, Shannon diversity) was significantly decreased in frozen samples for DNA-based analysis (P < 0.05), with less difference seen for cDNA-based sequencing. Using DNA-based analysis, length of storage had a significant impact (P < 0.05) on the bacterial community profiles. For potentially metabolically active populations, storage overall had less of an effect on the bacterial community composition, with a significant effect of buffer (P < 0.05). Individual horse had the most significant effect within both DNA and cDNA bacterial communities. Frozen storage of equine FMT material can preserve potentially metabolically active bacteria of the equine fecal microbiome, with saline plus glycerol preservation more effective than saline alone. Larger studies are needed to determine if these findings apply to other individual horses. The ability to freeze FMT material for use in equine patients could allow for easier clinical use of fecal transplant in horses with disturbances in their intestinal microbiome.

Sections du résumé

BACKGROUND BACKGROUND
Currently, lack of standardization for fecal microbiota transplantation (FMT) in equine practice has resulted in highly variable techniques, and there is no data on the bacterial metabolic activity or viability of the administered product. The objectives of this study were to compare the total and potentially metabolically active bacterial populations in equine FMT, and assess the effect of different frozen storage times, buffers, and temperatures on an equine FMT product. Fresh feces collected from three healthy adult horses was subjected to different storage methods. This included different preservation solutions (saline plus glycerol or saline only), temperature (-20 °C or -80 °C), and time (fresh, 30, 60, or 90 days). Samples underwent DNA extraction to assess total bacterial populations (both live and dead combined) and RNA extraction followed by reverse transcription to cDNA as a proxy to assess viable bacteria, then 16s rRNA gene amplicon sequencing using the V1-V2 region.
RESULTS RESULTS
The largest difference in population indices and taxonomic composition at the genus level was seen when evaluating the results of DNA-based (total) and cDNA-based (potentially metabolically active) extraction method. At the community level, alpha diversity (observed species, Shannon diversity) was significantly decreased in frozen samples for DNA-based analysis (P < 0.05), with less difference seen for cDNA-based sequencing. Using DNA-based analysis, length of storage had a significant impact (P < 0.05) on the bacterial community profiles. For potentially metabolically active populations, storage overall had less of an effect on the bacterial community composition, with a significant effect of buffer (P < 0.05). Individual horse had the most significant effect within both DNA and cDNA bacterial communities.
CONCLUSIONS CONCLUSIONS
Frozen storage of equine FMT material can preserve potentially metabolically active bacteria of the equine fecal microbiome, with saline plus glycerol preservation more effective than saline alone. Larger studies are needed to determine if these findings apply to other individual horses. The ability to freeze FMT material for use in equine patients could allow for easier clinical use of fecal transplant in horses with disturbances in their intestinal microbiome.

Identifiants

pubmed: 38987780
doi: 10.1186/s12917-024-04166-w
pii: 10.1186/s12917-024-04166-w
doi:

Substances chimiques

RNA, Ribosomal, 16S 0
DNA, Bacterial 0

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

306

Subventions

Organisme : Raymond Firestone Trust
ID : 580-5805-1-461612-xxxx-2000-5872
Organisme : Raymond Firestone Trust
ID : 580-5805-1-461612-xxxx-2000-5872

Informations de copyright

© 2024. The Author(s).

Références

Gough E, Shaikh H, Manges AR. Systematic review of intestinal microbiota transplantation (fecal bacteriotherapy) for recurrent Clostridium difficile infection. Clin Infect Dis. 2011;53(10):994–1002.
pubmed: 22002980 doi: 10.1093/cid/cir632
Allen-Vercoe E, Reid G, Viner N, Gloor GB, Hota S, Kim P, et al. A Canadian working group report on fecal microbial therapy: microbial ecosystems therapeutics. Can J Gastroenterol. 2012;26(7):457–62.
pubmed: 22803022 pmcid: 3395448 doi: 10.1155/2012/213828
Leffler DA, Lamont JT. Clostridium difficile infection. N Engl J Med. 2015;372(16):1539–48.
pubmed: 25875259 doi: 10.1056/NEJMra1403772
Hamilton M, Weingarden A, Sadowsky M, Khoruts A. Standardized frozen preparation for transplantation of fecal microbiota for recurrent Clostridium difficile infection. Am J Gastroenterol. 2012;107(5):761–7.
pubmed: 22290405 doi: 10.1038/ajg.2011.482
Aggeletopoulou I, Konstantakis C, Assimakopoulos SF, Triantos C. The role of the gut microbiota in the treatment of inflammatory bowel diseases. Microb Pathog. 2019;137:103774.
pubmed: 31586663 doi: 10.1016/j.micpath.2019.103774
Rokkas T, Gisbert JP, Gasbarrini A, Hold GL, Tilg H, Malfertheiner P, et al. A network meta-analysis of randomized controlled trials exploring the role of fecal microbiota transplantation in recurrent Clostridium difficile infection. United Eur Gastroenterol J. 2019;7(8):1051–63.
doi: 10.1177/2050640619854587
Youngster I, Sauk J, Pindar C, Wilson RG, Kaplan JL, Smith MB, et al. Fecal microbiota transplant for relapsing Clostridium difficile infection using a frozen inoculum from unrelated donors: a randomized, open-label, controlled pilot study. Clin Infect Dis. 2014;58(11):1515–22.
pubmed: 24762631 pmcid: 4017893 doi: 10.1093/cid/ciu135
Bakken JS, Borody T, Brandt LJ, Brill JV, Demarco DC, Franzos MA, et al. Treating Clostridium difficile infection with fecal microbiota transplantation. Clin Gastroenterol Hepatol. 2011;9(12):1044–9.
pubmed: 21871249 pmcid: 3223289 doi: 10.1016/j.cgh.2011.08.014
Satokari R, Mattila E, Kainulainen V, Arkkila PET. Simple faecal preparation and efficacy of frozen inoculum in faecal microbiota transplantation for recurrent Clostridium difficile infection – an observational cohort study. Aliment Pharmacol Ther. 2015;41(1):46–53.
pubmed: 25355279 doi: 10.1111/apt.13009
Li N, Tian HL, Chen QY, Yang B, Ma CL, Lin ZL, et al. Efficacy analysis of fecal microbiota transplantation in the treatment of 2010 patients with intestinal disorders. Chin J Gastrointest Surg. 2019;22(9):861–8.
Tuniyazi M, Wang W, Zhang N. A systematic review of current applications of fecal microbiota transplantation in horses. Vet Sci. 2023;10(4):290.
pubmed: 37104445 pmcid: 10141098
Buffie CG, Pamer EG. Microbiota-mediated colonization resistance against intestinal pathogens. Nat Rev Immunol. 2013;13(11):790–801.
pubmed: 24096337 pmcid: 4194195 doi: 10.1038/nri3535
Abt MC, Pamer EG. Commensal bacteria mediated defenses against pathogens. Curr Opin Immunol. 2014;0:16–22.
pmcid: 4132187 doi: 10.1016/j.coi.2014.03.003
Costa MC, Arroyo LG, Allen-Vercoe E, Stämpfli HR, Kim PT, Sturgeon A et al. Comparison of the fecal microbiota of healthy horses and horses with colitis by high throughput sequencing of the V3-V5 region of the 16S rRNA gene. PLoS One. 2012;7(7).
Stewart HL, Southwood LL, Indugu N, Vecchiarelli B, Engiles JB, Pitta D. Differences in the equine faecal microbiota between horses presenting to a tertiary referral hospital for colic compared with an elective surgical procedure. Equine Vet J. 2019;51(3):336–42.
pubmed: 30153353 doi: 10.1111/evj.13010
Arnold C, Pilla R, Chaffin K, Lidbury J, Steiner J, Suchodolski J. Alterations in the fecal microbiome and metabolome of horses with antimicrobial-associated diarrhea compared to antibiotic-treated and non-treated healthy case controls. Animals. 2021;11(6):1807.
pubmed: 34204371 pmcid: 8235368 doi: 10.3390/ani11061807
Weese JS, Holcombe SJ, Embertson RM, Kurtz KA, Roessner HA, Jalali M, et al. Changes in the faecal microbiota of mares precede the development of post partum colic. Equine Vet J. 2015;47(6):641–9.
pubmed: 25257320 doi: 10.1111/evj.12361
Elzinga SE, Weese JS, Adams AA. Comparison of the fecal microbiota in horses with equine metabolic syndrome and metabolically normal controls fed a similar all-forage diet. J Equine Vet Sci. 2016;44:9–16.
doi: 10.1016/j.jevs.2016.05.010
Leclere M, Costa MC. Fecal microbiota in horses with asthma. J Vet Intern Med. 2020;34(2):996–1006.
pubmed: 32128892 pmcid: 7096608 doi: 10.1111/jvim.15748
Dias DPM, Sousa SS, Molezini FA, Ferreira HSD, de Campos R. Efficacy of faecal microbiota transplantation for treating acute colitis in horses undergoing colic surgery. Pesq Vet Bras. 2018;38(8):1564–9.
doi: 10.1590/1678-5150-pvb-5521
McKinney CA, Oliveira BCM, Bedenice D, Paradis MR, Mazan M, Sage S et al. The fecal microbiota of healthy donor horses and geriatric recipients undergoing fecal microbial transplantation for the treatment of diarrhea. PLoS ONE. 2020;15(3).
Di Pietro R. Development of a protocol with concentrated bacteria for fecal microbiota transplantation and impact on the equine fecal microbiota after antibiotic-induced dysbiosis. Published Online January 14, 2021. https://papyrus.bib.umontreal.ca/xmlui/handle/1866/24704 .
McKinney CA, Bedenice D, Pacheco AP, Oliveira BCM, Paradis MR, Mazan M, et al. Assessment of clinical and microbiota responses to fecal microbial transplantation in adult horses with diarrhea. PLoS ONE. 2021;16(1):e0244381.
pubmed: 33444319 pmcid: 7808643 doi: 10.1371/journal.pone.0244381
Costa M, Di Pietro R, Bessegatto JA, Pereira PFV, Stievani FC, Gomes RG, et al. Evaluation of changes in microbiota after fecal microbiota transplantation in 6 diarrheic horses. Can Vet J. 2021;62(10):1123–30.
pubmed: 34602643 pmcid: 8439339
Laustsen L, Edwards JE, Hermes GDA, Luthersson N, van Doorn DA, Okrathok, et al. Free faecal water: analysis of horse faecal microbiota and the impact of faecal microbial transplantation on symptom severity. Anim (Basel). 2021;11(10):2776.
Kinoshita Y, Niwa H, Uchida-Fujii E, Nukada T, Ueno T. Simultaneous daily fecal microbiota transplantation fails to prevent metronidazole-induced dysbiosis of equine gut microbiota. J Equine Vet Sci. 2022;114:104004.
pubmed: 35526726 doi: 10.1016/j.jevs.2022.104004
Di Pietro R, Arroyo LG, Leclere M, Costa M. Effects of concentrated fecal microbiota transplant on the equine fecal microbiota after antibiotic-induced dysbiosis. Can J Vet Res. 2023;87(2):85–96.
pubmed: 37020579 pmcid: 10069150
Stewart HL, Pitta D, Indugu N, Vecchiarelli B, Engiles JB, Southwood LL. Characterization of the fecal microbiota of healthy horses. Am J Vet Res. 2018;79(8):811–9.
pubmed: 30058849 doi: 10.2460/ajvr.79.8.811
Takahashi M, Ishikawa D, Sasaki T, Lu YJ, Kuwahara-Arai K, Kamei M, et al. Faecal freezing preservation period influences colonization ability for faecal microbiota transplantation. J Appl Microbiol. 2019;126(3):973–84.
pubmed: 30489686 doi: 10.1111/jam.14167
Kopper JJ, Alexander TL, Kogan CJ, Berreta AR, Burbick CR. In vitro evaluation of the effect of storage at – 20°C and proximal gastrointestinal conditions on viability of equine fecal microbiota transplant. J Equine Vet Sci. 2021;98:103360.
pubmed: 33663713 doi: 10.1016/j.jevs.2020.103360
Loublier C, Taminiau B, Heinen J, Lecoq L, Amory H, Daube G, Cesarini C. Evaluation of bacterial composition and viability of equine feces after processing for transplantation. Microorganisms. 2023;11(2):231.
pubmed: 36838196 pmcid: 9966902 doi: 10.3390/microorganisms11020231
Kaplan-Shabtai V, Indugu N, Hennessy ML, Vecchiarelli B, Bender JS, Stefanovski D et al. Using structural equation modeling to understand interactions between bacterial and archaeal populations and volatile fatty acid proportions in the rumen. Front Microbiol. 2021;12.
Li R, Tun HM, Jahan M, Zhang Z, Kumar A, Dilantha Fernando WG, et al. Comparison of DNA-, PMA-, and RNA-based 16S rRNA Illumina sequencing for detection of live bacteria in water. Sci Rep. 2017;7:5752.
pubmed: 28720878 pmcid: 5515937 doi: 10.1038/s41598-017-02516-3
Gosalbes MJ, Durbán A, Pignatelli M, Abellan JJ, Jiménez-Hernández N, Pérez-Cobas AE et al. Metatranscriptomic approach to analyze the functional human gut microbiota. Quintana-Murci L, editor. PLoS One. 2011;6(3):e17447.
Holzhausen EA, Nikodemova M, Deblois CL, Barnet JH, Peppard PE, Suen G, et al. Assessing the impact of storage time on the stability of stool microbiota richness, diversity, and composition. Gut Pathogens. 2021;13(1):75.
pubmed: 34930464 pmcid: 8686582 doi: 10.1186/s13099-021-00470-0
Bundgaard-Nielsen C, Hagstrøm S, Sørensen S. Interpersonal variations in gut microbiota profiles supersedes the effects of differing fecal storage conditions. Sci Rep. 2018;8(1):17367.
pubmed: 30478355 pmcid: 6255890 doi: 10.1038/s41598-018-35843-0
Al KF, Bisanz JE, Gloor GB, Reid G, Burton JP. Evaluation of sampling and storage procedures on preserving the community structure of stool microbiota: a simple at-home toilet-paper collection method. J Microbiol Methods. 2018;144:117–21.
pubmed: 29155236 doi: 10.1016/j.mimet.2017.11.014
Fouhy F, Deane J, Rea MC, O’Sullivan Ó, Ross RP, O’Callaghan G, et al. The effects of freezing on faecal microbiota as determined using MiSeq sequencing and culture-based investigations. PLoS ONE. 2015;10(3):e0119355.
pubmed: 25748176 pmcid: 4352061 doi: 10.1371/journal.pone.0119355
Guo Y, Li SH, Kuang YS, He JR, Lu JH, Luo BJ, et al. Effect of short-term room temperature storage on the microbial community in infant fecal samples. Sci Rep. 2016;6:26648.
pubmed: 27226242 pmcid: 4880902 doi: 10.1038/srep26648
Theelen MJP, Luiken REC, Wagenaar JA, van Sloet MM, Rossen JWA, Zomer AL. The equine faecal microbiota of healthy horses and ponies in the Netherlands: impact of host and environmental factors. Animals. 2021;11(6):1762.
pubmed: 34204691 pmcid: 8231505 doi: 10.3390/ani11061762
Indugu N, Hennessy M, Kaplan-Shabtai VS, de Assis Lage CF, Räisänen SE, Melgar A, et al. Comparing noninvasive sampling techniques with standard cannula sampling method for ruminal microbial analysis. JDS Commun. 2021;2(6):329–33.
pubmed: 36337103 pmcid: 9623630 doi: 10.3168/jdsc.2021-0094
Li F, Henderson G, Sun X, Cox F, Janssen PH, Guan LL. Taxonomic assessment of rumen microbiota using total RNA and targeted amplicon sequencing approaches. Front Microbiol. 2016;7:987.
pubmed: 27446027 pmcid: 4916217
Shakya M, Lo CC, Chain PSG. Advances and challenges in metatranscriptomic analysis. Front Genet. 2019;10.
Lettat A, Benchaar C. Diet-induced alterations in total and metabolically active microbes within the rumen of dairy cows. PLoS ONE. 2013;8(4):e60978.
pubmed: 23593365 pmcid: 3622600 doi: 10.1371/journal.pone.0060978
Lee S, Kemp PF. Single-cell RNA content of natural marine planktonic bacteria measured by hybridization with multiple 16S rRNA-targeted fluorescent probes. Limnol Oceanogr. 1994;39(4):869–79.
doi: 10.4319/lo.1994.39.4.0869
Knecht H, Neulinger SC, Heinsen FA, Knecht C, Schilhabel A, Schmitz RA, et al. Effects of β-Lactam antibiotics and fluoroquinolones on human gut microbiota in relation to Clostridium difficile Associated Diarrhea. PLoS ONE. 2014;9(2):e89417.
pubmed: 24586762 pmcid: 3938479 doi: 10.1371/journal.pone.0089417
Kumar SS, Ghosh AR. Assessment of bacterial viability: a comprehensive review on recent advances and challenges. Microbiology. 2019;165(6):593–610.
pubmed: 30843781 doi: 10.1099/mic.0.000786
Beckers KF, Schulz CJ, Childers GW. Rapid regrowth and detection of microbial contaminants in equine fecal microbiome samples. PLoS ONE. 2017;12(11):e0187044.
pubmed: 29091944 pmcid: 5665523 doi: 10.1371/journal.pone.0187044
Martin de Bustamante M, Plummer C, MacNicol J, Gomez D. Impact of ambient temperature sample storage on the equine fecal microbiota. Animals. 2021;11(3):819.
pubmed: 33803934 pmcid: 8001224 doi: 10.3390/ani11030819
Gavriliuc S, Stothart MR, Henry A, Poissant J. Long-term storage of feces at – 80°C versus – 20°C is negligible for 16S rRNA amplicon profiling of the equine bacterial microbiome. PeerJ. 2021;9:e10837.
pubmed: 33854827 pmcid: 7953882 doi: 10.7717/peerj.10837
Ma J, Sheng L, Hong Y, Xi C, Gu Y, Zheng N et al. Variations of gut microbiome profile under different storage conditions and preservation periods: a multi-dimensional evaluation. Front Microbiol. 2020;11.
Willette JA, Pitta D, Indugu N, Vecchiarelli B, Hennessy ML, Dobbie T, et al. Experimental crossover study on the effects of withholding feed for 24 h on the equine faecal bacterial microbiota in healthy mares. BMC Vet Res. 2021;17(1):3.
pubmed: 33402190 pmcid: 7786913 doi: 10.1186/s12917-020-02706-8
Pitta DW, Indugu N, Vecchiarelli B, Rico DE, Harvatine KJ. Alterations in ruminal bacterial populations at induction and recovery from diet-induced milk fat depression in dairy cows. J Dairy Sci. 2018;101(1):295–309.
pubmed: 29103706 doi: 10.3168/jds.2016-12514
De Weirdt R, Hernandez-Sanabria E, Fievez V, Mees E, Geirnaert A, Van Herreweghen F, et al. Mucosa-associated biohydrogenating microbes protect the simulated colon microbiome from stress associated with high concentrations of poly-unsaturated fat. Environ Microbiol. 2017;19(2):722–39.
pubmed: 27883264 doi: 10.1111/1462-2920.13622
Bhattacharya S. Cryoprotectants and their usage in cryopreservation process. Cryopreservation Biotechnology in Biomedical and Biological sciences. IntechOpen; 2018. pp. 7–19.
Hubálek Z. Protectants used in the cryopreservation of microorganisms. Cryobiology. 2003;46(3):205–29.
pubmed: 12818211 doi: 10.1016/S0011-2240(03)00046-4
Gaci N, Chaudhary PP, Tottey W, Alric M, Brugère JF. Functional amplification and preservation of human gut microbiota. Microb Ecol Health Dis. 2017;28(1):1308070.
pubmed: 28572754 pmcid: 5443092
Deschamps C, Fournier E, Uriot O, Lajoie F, Verdier C, Comtet-Marre S, et al. Comparative methods for fecal sample storage to preserve gut microbial structure and function in an in vitro model of the human colon. Appl Microbiol Biotechnol. 2020;104(23):10233–47.
pubmed: 33085024 doi: 10.1007/s00253-020-10959-4
Cammarota G, Ianiro G, Kelly CR, Mullish BH, Allegretti JR, Kassam Z et al. International consensus conference on stool banking for faecal microbiota transplantation in clinical practice. Gut. 2019;68(12):2111–21.
Yu Z, Morrison M. Improved extraction of PCR-quality community DNA from digesta and fecal samples. Biotechniques. 2004;36(5):808–12.
pubmed: 15152600 doi: 10.2144/04365ST04
Song SJ, Lauber C, Costello EK, Lozupone CA, Humphrey G, Berg-Lyons D, et al. Cohabiting family members share microbiota with one another and with their dogs. Elife. 2013;2:e00458.
pubmed: 23599893 pmcid: 3628085 doi: 10.7554/eLife.00458
Bolyen E, Rideout JR, Dillon MR, Bokulich NA, Abnet CC, Al-Ghalith GA, et al. Reproducible, interactive, scalable and extensible microbiome data science using QIIME 2. Nat Biotechnol. 2019;37(8):852–7.
pubmed: 31341288 pmcid: 7015180 doi: 10.1038/s41587-019-0209-9
Callahan BJ, McMurdie PJ, Rosen MJ, Han AW, Johnson AJA, Holmes SP. DADA2: high-resolution sample inference from Illumina amplicon data. Nat Methods. 2016;13(7):581–3.
pubmed: 27214047 pmcid: 4927377 doi: 10.1038/nmeth.3869
Katoh K, Standley DM. MAFFT multiple sequence alignment Software Version 7: improvements in performance and usability. Mol Biol Evol. 2013;30(4):772–80.
pubmed: 23329690 pmcid: 3603318 doi: 10.1093/molbev/mst010
Price MN, Dehal PS, Arkin AP. FastTree 2–approximately maximum-likelihood trees for large alignments. PLoS ONE. 2010;5(3):e9490.
pubmed: 20224823 pmcid: 2835736 doi: 10.1371/journal.pone.0009490
DeSantis TZ, Hugenholtz P, Larsen N, Rojas M, Brodie EL, Keller K, et al. Greengenes, a chimera-checked 16S rRNA gene database and workbench compatible with ARB. Appl Environ Microbiol. 2006;72(7):5069–72.
pubmed: 16820507 pmcid: 1489311 doi: 10.1128/AEM.03006-05
Anderson MJ. A new method for non-parametric multivariate analysis of variance. Austral Ecol. 2001;26(1):32–46.

Auteurs

Alicia E Long (AE)

Department of Clinical Studies, New Bolton Center, University of Pennsylvania, Kennett Square, PA, USA. lalicia@vet.upenn.edu.

Dipti Pitta (D)

Department of Clinical Studies, New Bolton Center, University of Pennsylvania, Kennett Square, PA, USA.

Meagan Hennessy (M)

Department of Clinical Studies, New Bolton Center, University of Pennsylvania, Kennett Square, PA, USA.

Nagaraju Indugu (N)

Department of Clinical Studies, New Bolton Center, University of Pennsylvania, Kennett Square, PA, USA.

Bonnie Vecchiarelli (B)

Department of Clinical Studies, New Bolton Center, University of Pennsylvania, Kennett Square, PA, USA.

Daniela Luethy (D)

Department of Clinical Studies, New Bolton Center, University of Pennsylvania, Kennett Square, PA, USA.

Helen Aceto (H)

Department of Clinical Studies, New Bolton Center, University of Pennsylvania, Kennett Square, PA, USA.

Samuel Hurcombe (S)

Department of Clinical Studies, New Bolton Center, University of Pennsylvania, Kennett Square, PA, USA.
Veterinary Innovative Partners, New York, NY, USA.

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