Skeletal muscle from TBC1D4 p.Arg684Ter variant carriers is severely insulin resistant but exhibits normal metabolic responses during exercise.


Journal

Nature metabolism
ISSN: 2522-5812
Titre abrégé: Nat Metab
Pays: Germany
ID NLM: 101736592

Informations de publication

Date de publication:
31 Oct 2024
Historique:
received: 03 11 2023
accepted: 27 09 2024
medline: 1 11 2024
pubmed: 1 11 2024
entrez: 1 11 2024
Statut: aheadofprint

Résumé

In the Greenlandic Inuit population, 4% are homozygous carriers of a genetic nonsense TBC1D4 p.Arg684Ter variant leading to loss of the muscle-specific isoform of TBC1D4 and an approximately tenfold increased risk of type 2 diabetes

Identifiants

pubmed: 39482542
doi: 10.1038/s42255-024-01153-1
pii: 10.1038/s42255-024-01153-1
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Subventions

Organisme : Det Frie Forskningsråd (Danish Council for Independent Research)
ID : #8020 00288B
Organisme : Det Frie Forskningsråd (Danish Council for Independent Research)
ID : #5053 00095B
Organisme : Det Frie Forskningsråd (Danish Council for Independent Research)
ID : #7016 00386B
Organisme : Novo Nordisk Fonden (Novo Nordisk Foundation)
ID : #NNF14OC0013057
Organisme : Novo Nordisk Fonden (Novo Nordisk Foundation)
ID : #NNF17OC0028136
Organisme : Novo Nordisk Fonden (Novo Nordisk Foundation)
ID : #NNF17SA0031406
Organisme : Novo Nordisk Fonden (Novo Nordisk Foundation)
ID : #NNF14OC0013057
Organisme : Novo Nordisk Fonden (Novo Nordisk Foundation)
ID : #NNF17OC0028136
Organisme : Novo Nordisk Fonden (Novo Nordisk Foundation)
ID : #NNF17OC0028136
Organisme : Novo Nordisk Foundation Center for Basic Metabolic Research (NovoNordisk Foundation Center for Basic Metabolic Research)
ID : #NNF18CC0034900

Informations de copyright

© 2024. The Author(s).

Références

Moltke, I. et al. A common Greenlandic TBC1D4 variant confers muscle insulin resistance and type 2 diabetes. Nature 512, 190–193 (2014).
pubmed: 25043022 doi: 10.1038/nature13425
Jorgensen, M. E. et al. Diabetes and impaired glucose tolerance among the inuit population of Greenland. Diabetes Care 25, 1766–1771 (2002).
pubmed: 12351475 doi: 10.2337/diacare.25.10.1766
Dahl Petersen, I. K., Bjerregaard, P. & Jørgensen, M. E. Physical activity patterns in Greenland: a country in transition. Scand. J. Public Health 39, 678–686 (2011).
pubmed: 21948977 doi: 10.1177/1403494811420486
Dahl-Petersen, I. K., Bjerregaard, P., Brage, S. & Jørgensen, M. E. Physical activity energy expenditure is associated with 2-h insulin independently of obesity among Inuit in Greenland. Diabetes Res. Clin. Pract. 102, 242–249 (2013).
pubmed: 24176243 doi: 10.1016/j.diabres.2013.10.004
Schnurr, T. M. et al. Physical activity attenuates postprandial hyperglycaemia in homozygous TBC1D4 loss-of-function mutation carriers. Diabetologia 64, 1795–1804 (2021).
pubmed: 33912980 pmcid: 8245392 doi: 10.1007/s00125-021-05461-z
Baus, D. et al. Identification of a novel AS160 splice variant that regulates GLUT4 translocation and glucose-uptake in rat muscle cells. Cell. Signal. 20, 2237–2246 (2008).
pubmed: 18771725 doi: 10.1016/j.cellsig.2008.08.010
Espelage, L., Al-Hasani, H. & Chadt, A. RabGAPs in skeletal muscle function and exercise. J. Mol. Endocrinol. 64, R1–R19 (2020).
pubmed: 31627187 doi: 10.1530/JME-19-0143
Mafakheri, S., Chadt, A. & Al-Hasani, H. Regulation of RabGAPs involved in insulin action. Biochem. Soc. Trans. 46, 683–690 (2018).
pubmed: 29784647 doi: 10.1042/BST20170479
Karlsson, H. K. R. et al. Insulin-stimulated phosphorylation of the Akt substrate AS160 is impaired in skeletal muscle of type 2 diabetic subjects. Diabetes 54, 1692–1697 (2005).
pubmed: 15919790 doi: 10.2337/diabetes.54.6.1692
Højlund, K. et al. Impaired insulin-stimulated phosphorylation of akt and AS160 in skeletal muscle of women with polycystic ovary syndrome is reversed by pioglitazone treatment. Diabetes 57, 357–366 (2008).
pubmed: 17977950 doi: 10.2337/db07-0706
Vind, B. F. et al. Impaired insulin-induced site-specific phosphorylation of TBC1 domain family, member 4 (TBC1D4) in skeletal muscle of type 2 diabetes patients is restored by endurance exercise-training. Diabetologia 54, 157–167 (2011).
pubmed: 20938636 doi: 10.1007/s00125-010-1924-4
Middelbeek, R. J. W. et al. Insulin stimulation regulates AS160 and TBC1D1 phosphorylation sites in human skeletal muscle. Nutr. Diabetes 3, e74 (2013).
pubmed: 23752133 pmcid: 3697402 doi: 10.1038/nutd.2013.13
Heller, S. R. et al. Risk of hypoglycaemia in types 1 and 2 diabetes: effects of treatment modalities and their duration. Diabetologia 50, 1140–1147 (2007).
doi: 10.1007/s00125-007-0599-y
Lupoli, R. et al. Role of the entero-insular axis in the pathogenesis of idiopathic reactive hypoglycemia: a pilot study. J. Clin. Endocrinol. Metab. 100, 4441–4446 (2015).
pubmed: 26502359 doi: 10.1210/jc.2015-3309
Ghosh, C., Mukhopadhyay, P., Ghosh, S. & Pradhan, M. Insulin sensitivity index (ISI0, 120) potentially linked to carbon isotopes of breath CO2 for pre-diabetes and type 2 diabetes. Sci. Rep. 5, 11959 (2015).
pubmed: 26148706 pmcid: 4493706 doi: 10.1038/srep11959
Heller, S. R. et al. A higher non-severe hypoglycaemia rate is associated with an increased risk of subsequent severe hypoglycaemia and major adverse cardiovascular events in individuals with type 2 diabetes in the LEADER study. Diabetologia 65, 55–64 (2022).
pubmed: 34704120 doi: 10.1007/s00125-021-05556-7
Overvad, M. et al. The effect of diabetes and the common diabetogenic TBC1D4 p.Arg684Ter variant on cardiovascular risk in Inuit in Greenland. Sci. Rep. 10, 22081 (2020).
pubmed: 33328529 pmcid: 7745023 doi: 10.1038/s41598-020-79132-1
Kanat, M., DeFronzo, R. A. & Abdul-Ghani, M. A. Treatment of prediabetes. World J. Diabetes 6, 1207 (2015).
pubmed: 26464759 pmcid: 4598604 doi: 10.4239/wjd.v6.i12.1207
DeFronzo, R. A. et al. Type 2 diabetes mellitus. Nat. Rev. Dis. Prim. 1, 15019 (2015).
pubmed: 27189025 doi: 10.1038/nrdp.2015.19
James, D. E., Stöckli, J. & Birnbaum, M. J. The aetiology and molecular landscape of insulin resistance. Nat. Rev. Mol. Cell Biol. 22, 751–771 (2021).
pubmed: 34285405 doi: 10.1038/s41580-021-00390-6
Larsen, J. K. et al. Illumination of the endogenous insulin-regulated TBC1D4 interactome in human skeletal muscle. Diabetes 71, 906–920 (2022).
pubmed: 35192682 pmcid: 9074744 doi: 10.2337/db21-0855
Xie, B. et al. The inactivation of RabGAP function of AS160 promotes lysosomal degradation of glut4 and causes postprandial hyperglycemia and hyperinsulinemia. Diabetes 65, 3327–3340 (2016).
pubmed: 27554475 doi: 10.2337/db16-0416
Larance, M. et al. Characterization of the role of the Rab GTPase-activating protein AS160 in insulin-regulated GLUT4 trafficking. J. Biol. Chem. 280, 37803–37813 (2005).
pubmed: 16154996 doi: 10.1074/jbc.M503897200
Peck, G. R. et al. Interaction of the Akt substrate, AS160, with the glucose transporter 4 vesicle marker protein, insulin-regulated aminopeptidase. Mol. Endocrinol. 20, 2576–2583 (2006).
pubmed: 16762977 doi: 10.1210/me.2005-0476
Eickelschulte, S. et al. AKT/AMPK-mediated phosphorylation of TBC1D4 disrupts the interaction with insulin-regulated aminopeptidase. J. Biol. Chem. 296, 100637 (2021).
pubmed: 33872597 pmcid: 8131924 doi: 10.1016/j.jbc.2021.100637
Chadt, A. et al. Deletion of both rab-GTPase-activating proteins TBC1D1 and TBC1D4 in mice eliminates insulin- and AICAR-stimulated glucose transport. Diabetes 64, 746–759 (2015).
pubmed: 25249576 doi: 10.2337/db14-0368
Lansey, M. N., Walker, N. N., Hargett, S. R., Stevens, J. R. & Keller, S. R. Deletion of Rab GAP AS160 modifies glucose uptake and GLUT4 translocation in primary skeletal muscles and adipocytes and impairs glucose homeostasis. Am. J. Physiol. Endocrinol. Metab. 303, E1273–E1286 (2012).
pubmed: 23011063 pmcid: 3517634 doi: 10.1152/ajpendo.00316.2012
Arias, E. B., Zheng, X., Agrawal, S. & Cartee, G. D. Whole body glucoregulation and tissue-specific glucose uptake in a novel Akt substrate of 160 kDa knockout rat model. PLoS ONE 14, e0216236 (2019).
pubmed: 31034517 pmcid: 6488193 doi: 10.1371/journal.pone.0216236
Treebak, J. T. et al. Potential role of TBC1D4 in enhanced post-exercise insulin action in human skeletal muscle. Diabetologia 52, 891–900 (2009).
pubmed: 19252894 pmcid: 3627047 doi: 10.1007/s00125-009-1294-y
Vendelbo, M. H. et al. Insulin resistance after a 72-h fast is associated with impaired AS160 phosphorylation and accumulation of lipid and glycogen in human skeletal muscle. Am. J. Physiol. Endocrinol. Metab. 302, E190–E200 (2012).
pubmed: 22028408 doi: 10.1152/ajpendo.00207.2011
Bergström, J. & Hultman, E. Muscle glycogen synthesis after exercise: an enhancing factor localized to the muscle cells in man.Nature 210, 309–310 (1966).
pubmed: 5954569 doi: 10.1038/210309a0
Richter, E. A., Mikines, K. J., Galbo, H. & Kiens, B. Effect of exercise on insulin action in human skeletal muscle. J. Appl. Physiol. 66, 876–885 (1989).
pubmed: 2496078 doi: 10.1152/jappl.1989.66.2.876
Kjøbsted, R. et al. Enhanced muscle insulin sensitivity after contraction/exercise is mediated by AMPK. Diabetes 66, 598–612 (2017).
pubmed: 27797909 doi: 10.2337/db16-0530
Kjøbsted, R. et al. TBC1D4 is necessary for enhancing muscle insulin sensitivity in response to AICAR and contraction. Diabetes 68, 1756–1766 (2019).
pubmed: 31175100 doi: 10.2337/db18-0769
Zheng, A. et al. Exercise-induced improvement in insulin-stimulated glucose uptake by rat skeletal muscle is absent in male AS160-Knockout rats, partially restored by muscle expression of phosphomutated AS160, and fully restored by muscle expression of wild-type AS160. Diabetes 71, 219–232 (2022).
pubmed: 34753801 doi: 10.2337/db21-0601
Cartee, G. D. & Wojtaszewski, J. F. P. Role of Akt substrate of 160 kDa in insulin-stimulated and contraction-stimulated glucose transport. Appl. Physiol. Nutr. Metab. 32, 557–566 (2007).
pubmed: 17510697 doi: 10.1139/H07-026
Kjøbsted, R. et al. AMPK and TBC1D1 regulate muscle glucose uptake after, but not during, exercise and contraction. Diabetes 68, 1427–1440 (2019).
pubmed: 31010958 doi: 10.2337/db19-0050
Steenberg, D. E. et al. A single bout of one-legged exercise to local exhaustion decreases insulin action in nonexercised muscle leading to decreased whole-body insulin action. Diabetes 69, 578–590 (2020).
pubmed: 31974138 doi: 10.2337/db19-1010
Kjøbsted, R. et al. TBC1D4-S711 controls skeletal muscle insulin sensitization after exercise and contraction. Diabetes 72, 857–871 (2023).
pubmed: 37074686 doi: 10.2337/db22-0666
Needham, E. J. et al. Personalized phosphoproteomics identifies functional signaling. Nat. Biotechnol. 40, 576–584 (2022).
pubmed: 34857927 doi: 10.1038/s41587-021-01099-9
An, D. et al. TBC1D1 regulates insulin- and contraction-induced glucose transport in mouse skeletal muscle. Diabetes 59, 1358–1365 (2010).
pubmed: 20299473 pmcid: 2874696 doi: 10.2337/db09-1266
Vichaiwong, K. et al. Contraction regulates site-specific phosphorylation of TBC1D1 in skeletal muscle. Biochem. J. 431, 311–320 (2010).
pubmed: 20701589 doi: 10.1042/BJ20101100
Stöckli, J. et al. The RabGAP TBC1D1 plays a central role in exercise-regulated glucose metabolism in skeletal muscle. Diabetes 64, 1914–1922 (2015).
pubmed: 25576050 doi: 10.2337/db13-1489
Zisman, A. et al. Targeted disruption of the glucose transporter 4 selectively in muscle causes insulin resistance and glucose intolerance. Nat. Med. 6, 924–928 (2000).
pubmed: 10932232 doi: 10.1038/78693
Wendt, C. D. et al. Contraction-mediated glucose transport in skeletal muscle is regulated by a framework of AMPK, TBC1D1/4, and Rac1. Diabetes 70, 2796–2809 (2021).
pubmed: 34561225 doi: 10.2337/db21-0587
Pillon, N. J. et al. Transcriptomic profiling of skeletal muscle adaptations to exercise and inactivity. Nat. Commun. 11, 470 (2020).
pubmed: 31980607 pmcid: 6981202 doi: 10.1038/s41467-019-13869-w
Klip, A., McGraw, T. E. & James, D. E. Thirty sweet years of GLUT4. J. Biol. Chem. 294, 11369–11381 (2019).
pubmed: 31175156 pmcid: 6663870 doi: 10.1074/jbc.REV119.008351
Ploug, T., Van Deurs, B., Ai, H., Cushman, S. W. & Ralston, E. Analysis of GLUT4 distribution in whole skeletal muscle fibers: Identification of distinct storage compartments that are recruited by insulin and muscle contractions. J. Cell Biol. 142, 1429–1446 (1998).
pubmed: 9744875 pmcid: 2141761 doi: 10.1083/jcb.142.6.1429
Wasserman, D. H., Kang, L., Ayala, J. E., Fueger, P. T. & Lee-Young, R. S. The physiological regulation of glucose flux into muscle in vivo. J. Exp. Biol. 214, 254–262 (2011).
pubmed: 21177945 doi: 10.1242/jeb.048041
Kramer, H. F. et al. AS160 regulates insulin- and contraction-stimulated glucose uptake in mouse skeletal muscle. J. Biol. Chem. 281, 31478–31485 (2006).
pubmed: 16935857 doi: 10.1016/S0021-9258(19)84060-7
Chen, S., Wasserman, D. H., MacKintosh, C. & Sakamoto, K. Mice with AS160/TBC1D4-Thr649Ala knockin mutation are glucose intolerant with reduced insulin sensitivity and altered GLUT4 trafficking. Cell Metab. 13, 68–79 (2011).
pubmed: 21195350 pmcid: 3081066 doi: 10.1016/j.cmet.2010.12.005
Yang, X. et al. Tissue-specific splicing and dietary interaction of a mutant As160 allele determine muscle metabolic fitness in rodents. Diabetes 70, 1826–1842 (2021).
pubmed: 33980689 doi: 10.2337/db21-0039
Wang, H. Y. et al. AS160 deficiency causes whole-body insulin resistance via composite effects in multiple tissues. Biochem. J 449, 479–489 (2013).
pubmed: 23078342 doi: 10.1042/BJ20120702
Mastrototaro, L. & Roden, M. Insulin resistance and insulin sensitizing agents. Metabolism. 125, 154892 (2021).
pubmed: 34563556 doi: 10.1016/j.metabol.2021.154892
Richter, E. A. & Hargreaves, M. Exercise, GLUT4, and skeletal muscle glucose uptake. Physiol. Rev. 93, 993–1017 (2013).
pubmed: 23899560 doi: 10.1152/physrev.00038.2012
Bjerregaard, P. et al. Inuit health in Greenland: a population survey of life style and disease in Greenland and among Inuit living in Denmark. Int. J. Circumpolar Health 62, 3–79 (2003).
pubmed: 14527126
Ekblom-Bak, E., Björkman, F., Hellenius, M. L. & Ekblom, B. A new submaximal cycle ergometer test for prediction of VO2max. Scand. J. Med. Sci. Sports 24, 319–326 (2014).
pubmed: 23126417 doi: 10.1111/sms.12014
Wojtaszewski, J. F. P., Hansen, B. F., Kiens, B. & Richter, E. A. Insulin signaling in human skeletal muscle: time course and effect of exercise. Diabetes 46, 1775–1781 (1997).
pubmed: 9356025 doi: 10.2337/diab.46.11.1775
Kristensen, J. M., Treebak, J. T., Schjerling, P., Goodyear, L. & Wojtaszewski, J. F. P. Two weeks of metformin treatment induces AMPK-dependent enhancement of insulin-stimulated glucose uptake in mouse soleus muscle. Am. J. Physiol. Endocrinol. Metab. 306, E1099–E1109 (2014).
pubmed: 24644243 pmcid: 4971810 doi: 10.1152/ajpendo.00417.2013
Albers, P. H. et al. Human muscle fiber type-specific insulin signaling: Impact of obesity and type 2 diabetes. Diabetes 64, 485–497 (2015).
pubmed: 25187364 doi: 10.2337/db14-0590
Bornø, A., Foged, L. & Van Hall, G. Glucose and glycerol concentrations and their tracer enrichment measurements using liquid chromatography tandem mass spectrometry. J. Mass Spectrom. 49, 980–988 (2014).
pubmed: 25303387 doi: 10.1002/jms.3407
Seidemann, J., Lowry, O. H. & Passonneau, J. V. A flexible system of enzymatic analysis. Academic Press, New York, 1972. 291 S., 32 Abb., 15 Tab., Preis $ 14.00. Starch (Stärke) 25, 322 (1973).
doi: 10.1002/star.19730250914
Højlund, K. et al. Dysregulation of glycogen synthase COOH- and NH
pubmed: 19837931 doi: 10.1210/jc.2009-0897
Rueden, C. T. et al. ImageJ2: ImageJ for the next generation of scientific image data. BMC Bioinf. 18, 529 (2017).
doi: 10.1186/s12859-017-1934-z
Dobin, A. & Gingeras, T. R. Mapping RNA-seq reads with STAR. Curr. Protoc. Bioinforma. 51, 11.14.1–11.14.19 (2015).
Wang, L. et al. Measure transcript integrity using RNA-seq data. BMC Bioinf. 17, 58 (2016).
doi: 10.1186/s12859-016-0922-z
Graubert, A., Aguet, F., Ravi, A., Ardlie, K. G. & Getz, G. RNA-SeQC 2: efficient RNA-seq quality control and quantification for large cohorts. Bioinformatics 37, 3048–3050 (2021).
pubmed: 33677499 pmcid: 8479667 doi: 10.1093/bioinformatics/btab135
Love, M. I., Huber, W. & Anders, S. Moderated estimation of fold change and dispersion for RNA-seq data with DESeq2. Genome Biol. 15, 550 (2014).
pubmed: 25516281 pmcid: 4302049 doi: 10.1186/s13059-014-0550-8
Udeshi, N. D., Mertins, P., Svinkina, T. & Carr, S. A. Large-scale identification of ubiquitination sites by mass spectrometry. Nat. Protoc. 8, 1950–1960 (2013).
pubmed: 24051958 pmcid: 4725055 doi: 10.1038/nprot.2013.120
Sinitcyn, P. et al. MaxDIA enables library-based and library-free data-independent acquisition proteomics. Nat. Biotechnol. 12, 1563–1573 (2021).
doi: 10.1038/s41587-021-00968-7
Subramanian, A. et al. Gene set enrichment analysis: a knowledge-based approach for interpreting genome-wide expression profiles. Proc. Natl Acad. Sci. USA 102, 15545–15550 (2005).
pubmed: 16199517 pmcid: 1239896 doi: 10.1073/pnas.0506580102
Mootha, V. K. et al. PGC-1α-responsive genes involved in oxidative phosphorylation are coordinately downregulated in human diabetes. Nat. Genet. 34, 267–273 (2003).
pubmed: 12808457 doi: 10.1038/ng1180
Ritchie, M. E. et al. Limma powers differential expression analyses for fRNA-sequencing and microarray studies. Nucleic Acids Res. 43, e47 (2015).
pubmed: 25605792 pmcid: 4402510 doi: 10.1093/nar/gkv007
Benjamini, Y. & Hochberg, Y. Controlling the false discovery rate: a practical and powerful approach to multiple testing. J. R. Stat. Soc. Ser. B 57, 289–300 (1995).
doi: 10.1111/j.2517-6161.1995.tb02031.x
Ochoa, D. et al. An atlas of human kinase regulation. Mol. Syst. Biol. 12, 888 (2016).
pubmed: 27909043 pmcid: 5199121 doi: 10.15252/msb.20167295
Abdul-Ghani, M. A., Molina-Carrion, M., Jani, R., Jenkinson, C. & DeFronzo, R. A. Adipocytes in subjects with impaired fasting glucose and impaired glucose tolerance are resistant to the anti-lipolytic effect of insulin. Acta Diabetol. 45, 147–150 (2008).
pubmed: 18357404 doi: 10.1007/s00592-008-0033-z
Gutt, M. et al. Validation of the insulin sensitivity index (ISI0,120): comparison with other measures. Diabetes Res. Clin. Pract. 47, 177–184 (2000).
pubmed: 10741566 doi: 10.1016/S0168-8227(99)00116-3

Auteurs

Jonas M Kristensen (JM)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Rasmus Kjøbsted (R)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Trine J Larsen (TJ)

Greenland Center of Health Research, Institute of Institute of Health and Nature, University of Greenland, Nuuk, Greenland.

Christian S Carl (CS)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Janne R Hingst (JR)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Johan Onslev (J)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Jesper B Birk (JB)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Anette Thorup (A)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Dorte E Steenberg (DE)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Jonas R Knudsen (JR)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Nicolai S Henriksen (NS)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Elise J Needham (EJ)

Charles Perkins Centre, School of Life and Environmental Sciences, University of Sydney, Sydney, New South Wales, Australia.

Jens F Halling (JF)

Section for Cell Biology and Physiology, Department of Biology, University of Copenhagen, Copenhagen, Denmark.

Anders Gudiksen (A)

Section for Cell Biology and Physiology, Department of Biology, University of Copenhagen, Copenhagen, Denmark.

Carsten F Rundsten (CF)

Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark.

Kristian E Hanghøj (KE)

The Bioinformatics Centre, Department of Biology, University of Copenhagen, Copenhagen, Denmark.

Sara E Stinson (SE)

Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark.

Birgitte Hoier (B)

August Krogh Section for Human Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Camilla C Hansen (CC)

August Krogh Section for Human Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Thomas E Jensen (TE)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Ylva Hellsten (Y)

August Krogh Section for Human Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Henriette Pilegaard (H)

Section for Cell Biology and Physiology, Department of Biology, University of Copenhagen, Copenhagen, Denmark.

Niels Grarup (N)

Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark.

Jesper Olesen (J)

Queen Ingrid Primary Health Care Center, Nuuk, Greenland.

Sean J Humphrey (SJ)

Charles Perkins Centre, School of Life and Environmental Sciences, University of Sydney, Sydney, New South Wales, Australia.

David E James (DE)

Charles Perkins Centre, School of Life and Environmental Sciences, University of Sydney, Sydney, New South Wales, Australia.
Sydney Medical School, University of Sydney, Sydney, New South Wales, Australia.

Michael L Pedersen (ML)

Greenland Center of Health Research, Institute of Institute of Health and Nature, University of Greenland, Nuuk, Greenland.
Steno Diabetes Center Greenland, Nuuk, Greenland.

Erik A Richter (EA)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark.

Torben Hansen (T)

Novo Nordisk Foundation Center for Basic Metabolic Research, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark.

Marit E Jørgensen (ME)

Greenland Center of Health Research, Institute of Institute of Health and Nature, University of Greenland, Nuuk, Greenland.
Steno Diabetes Center Copenhagen, Gentofte, Denmark.
National Institute of Public Health, University of Southern Denmark, Odense, Denmark.

Jørgen F P Wojtaszewski (JFP)

August Krogh Section for Molecular Physiology, Department of Nutrition, Exercise and Sports, Faculty of Science, University of Copenhagen, Copenhagen, Denmark. jw@nexs.ku.dk.

Classifications MeSH