Role of leucine zipper-like motifs in the oligomerization of Pseudomonas putida phasins.


Journal

Biochimica et biophysica acta. General subjects
ISSN: 1872-8006
Titre abrégé: Biochim Biophys Acta Gen Subj
Pays: Netherlands
ID NLM: 101731726

Informations de publication

Date de publication:
02 2019
Historique:
received: 15 06 2018
revised: 05 11 2018
accepted: 07 11 2018
pubmed: 13 11 2018
medline: 1 10 2019
entrez: 13 11 2018
Statut: ppublish

Résumé

Phasins are low molecular mass proteins that accumulate strongly in bacterial cells in response to the intracellular storage of polyhydroxyalkanoates (PHA). Although lacking catalytic activity, phasins are the major components of the surface of the PHA granules and could be potentially involved in the formation of a network-like protein layer surrounding the polyester inclusions. Structural models revealed phasins to possess coiled-coil regions that might be important in the establishment of protein-protein interactions. However, there is not experimental evidence of a coiled-coil mediated oligomerization in these proteins. Structure prediction analyses were used to characterize the coiled-coil motifs of phasins PhaF and PhaI -produced by the model bacterium Pseudomonas putida KT2440-. Their oligomerization was evaluated by biolayer interferometry and the in vivo two-hybrid (BACTH) system. The interaction ability of a series of coiled-coil mutated derivatives was also measured. The formation of PhaF and PhaI complexes was detected. A predicted short leucine zipper-like coiled-coil (ZIP), containing "ideal" residues located within the hydrophobic core, was shown responsible for the oligomers stability. The substitution of key residues (leucines or valines) in PhaI ZIP (ZIPI) for alanine reduced by four fold the oligomerization efficiency. These results indicate that coiled-coil motifs are essential for phasin interactions. Correct oligomerization requires the formation of a stable hydrophobic interface between both phasins. Our findings elucidate the oligomerization motif of PhaF and PhaI. This motif is present in most phasins from PHA-accumulating bacteria and offers a potentially important target for modulating the PHA granules stability.

Sections du résumé

BACKGROUND
Phasins are low molecular mass proteins that accumulate strongly in bacterial cells in response to the intracellular storage of polyhydroxyalkanoates (PHA). Although lacking catalytic activity, phasins are the major components of the surface of the PHA granules and could be potentially involved in the formation of a network-like protein layer surrounding the polyester inclusions. Structural models revealed phasins to possess coiled-coil regions that might be important in the establishment of protein-protein interactions. However, there is not experimental evidence of a coiled-coil mediated oligomerization in these proteins.
METHODS
Structure prediction analyses were used to characterize the coiled-coil motifs of phasins PhaF and PhaI -produced by the model bacterium Pseudomonas putida KT2440-. Their oligomerization was evaluated by biolayer interferometry and the in vivo two-hybrid (BACTH) system. The interaction ability of a series of coiled-coil mutated derivatives was also measured.
RESULTS
The formation of PhaF and PhaI complexes was detected. A predicted short leucine zipper-like coiled-coil (ZIP), containing "ideal" residues located within the hydrophobic core, was shown responsible for the oligomers stability. The substitution of key residues (leucines or valines) in PhaI ZIP (ZIPI) for alanine reduced by four fold the oligomerization efficiency.
CONCLUSIONS
These results indicate that coiled-coil motifs are essential for phasin interactions. Correct oligomerization requires the formation of a stable hydrophobic interface between both phasins.
GENERAL SIGNIFICANCE
Our findings elucidate the oligomerization motif of PhaF and PhaI. This motif is present in most phasins from PHA-accumulating bacteria and offers a potentially important target for modulating the PHA granules stability.

Identifiants

pubmed: 30419286
pii: S0304-4165(18)30350-7
doi: 10.1016/j.bbagen.2018.11.002
pii:
doi:

Substances chimiques

Bacterial Proteins 0
DNA-Binding Proteins 0
PHAP protein, Bacteria 0
Polyhydroxyalkanoates 0

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

362-370

Informations de copyright

Copyright © 2018 Elsevier B.V. All rights reserved.

Auteurs

Natalia A Tarazona (NA)

Department of Microbial and Plant Biotechnology, Centro de Investigaciones Biológicas, CSIC, Madrid 28040, Spain.

Beatriz Maestro (B)

Instituto de Biología Molecular y Celular, Universidad Miguel Hernández, Elche 03202, Spain.

Olga Revelles (O)

Department of Microbial and Plant Biotechnology, Centro de Investigaciones Biológicas, CSIC, Madrid 28040, Spain.

Jesús M Sanz (JM)

Department of Microbial and Plant Biotechnology, Centro de Investigaciones Biológicas, CSIC, Madrid 28040, Spain; Instituto de Biología Molecular y Celular, Universidad Miguel Hernández, Elche 03202, Spain.

M Auxiliadora Prieto (MA)

Department of Microbial and Plant Biotechnology, Centro de Investigaciones Biológicas, CSIC, Madrid 28040, Spain. Electronic address: auxi@cib.csic.es.

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Classifications MeSH