Disentangling the genetics of lean mass.


Journal

The American journal of clinical nutrition
ISSN: 1938-3207
Titre abrégé: Am J Clin Nutr
Pays: United States
ID NLM: 0376027

Informations de publication

Date de publication:
01 02 2019
Historique:
received: 22 03 2018
accepted: 11 09 2018
pubmed: 6 2 2019
medline: 23 10 2019
entrez: 6 2 2019
Statut: ppublish

Résumé

Lean body mass (LM) plays an important role in mobility and metabolic function. We previously identified five loci associated with LM adjusted for fat mass in kilograms. Such an adjustment may reduce the power to identify genetic signals having an association with both lean mass and fat mass. To determine the impact of different fat mass adjustments on genetic architecture of LM and identify additional LM loci. We performed genome-wide association analyses for whole-body LM (20 cohorts of European ancestry with n = 38,292) measured using dual-energy X-ray absorptiometry) or bioelectrical impedance analysis, adjusted for sex, age, age2, and height with or without fat mass adjustments (Model 1 no fat adjustment; Model 2 adjustment for fat mass as a percentage of body mass; Model 3 adjustment for fat mass in kilograms). Seven single-nucleotide polymorphisms (SNPs) in separate loci, including one novel LM locus (TNRC6B), were successfully replicated in an additional 47,227 individuals from 29 cohorts. Based on the strengths of the associations in Model 1 vs Model 3, we divided the LM loci into those with an effect on both lean mass and fat mass in the same direction and refer to those as "sumo wrestler" loci (FTO and MC4R). In contrast, loci with an impact specifically on LM were termed "body builder" loci (VCAN and ADAMTSL3). Using existing available genome-wide association study databases, LM increasing alleles of SNPs in sumo wrestler loci were associated with an adverse metabolic profile, whereas LM increasing alleles of SNPs in "body builder" loci were associated with metabolic protection. In conclusion, we identified one novel LM locus (TNRC6B). Our results suggest that a genetically determined increase in lean mass might exert either harmful or protective effects on metabolic traits, depending on its relation to fat mass.

Sections du résumé

Background
Lean body mass (LM) plays an important role in mobility and metabolic function. We previously identified five loci associated with LM adjusted for fat mass in kilograms. Such an adjustment may reduce the power to identify genetic signals having an association with both lean mass and fat mass.
Objectives
To determine the impact of different fat mass adjustments on genetic architecture of LM and identify additional LM loci.
Methods
We performed genome-wide association analyses for whole-body LM (20 cohorts of European ancestry with n = 38,292) measured using dual-energy X-ray absorptiometry) or bioelectrical impedance analysis, adjusted for sex, age, age2, and height with or without fat mass adjustments (Model 1 no fat adjustment; Model 2 adjustment for fat mass as a percentage of body mass; Model 3 adjustment for fat mass in kilograms).
Results
Seven single-nucleotide polymorphisms (SNPs) in separate loci, including one novel LM locus (TNRC6B), were successfully replicated in an additional 47,227 individuals from 29 cohorts. Based on the strengths of the associations in Model 1 vs Model 3, we divided the LM loci into those with an effect on both lean mass and fat mass in the same direction and refer to those as "sumo wrestler" loci (FTO and MC4R). In contrast, loci with an impact specifically on LM were termed "body builder" loci (VCAN and ADAMTSL3). Using existing available genome-wide association study databases, LM increasing alleles of SNPs in sumo wrestler loci were associated with an adverse metabolic profile, whereas LM increasing alleles of SNPs in "body builder" loci were associated with metabolic protection.
Conclusions
In conclusion, we identified one novel LM locus (TNRC6B). Our results suggest that a genetically determined increase in lean mass might exert either harmful or protective effects on metabolic traits, depending on its relation to fat mass.

Identifiants

pubmed: 30721968
pii: S0002-9165(22)03110-0
doi: 10.1093/ajcn/nqy272
pmc: PMC6500901
doi:

Substances chimiques

ADAMTSL3 protein, human 0
Extracellular Matrix Proteins 0
MC4R protein, human 0
RNA-Binding Proteins 0
Receptor, Melanocortin, Type 4 0
TNRC6B protein, human 0
VCAN protein, human 0
Versicans 126968-45-4
Alpha-Ketoglutarate-Dependent Dioxygenase FTO EC 1.14.11.33
FTO protein, human EC 1.14.11.33
ADAMTS Proteins EC 3.4.24.-

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

276-287

Subventions

Organisme : Medical Research Council
ID : MC_UU_00007/10
Pays : United Kingdom
Organisme : NIA NIH HHS
ID : P30 AG024827
Pays : United States
Organisme : NCCDPHP CDC HHS
ID : U01 DP006266
Pays : United States
Organisme : Medical Research Council
ID : MR/N003284/1
Pays : United Kingdom
Organisme : NCATS NIH HHS
ID : UL1 TR002369
Pays : United States
Organisme : NIDDK NIH HHS
ID : R01 DK110113
Pays : United States
Organisme : NIAMS NIH HHS
ID : R01 AR041398
Pays : United States
Organisme : Cancer Research UK
ID : 14136
Pays : United Kingdom
Organisme : Medical Research Council
ID : G0401527
Pays : United Kingdom
Organisme : NIDDK NIH HHS
ID : P30 DK020541
Pays : United States
Organisme : NIA NIH HHS
ID : U24 AG051129
Pays : United States
Organisme : NIA NIH HHS
ID : R01 AG015819
Pays : United States
Organisme : NIAMS NIH HHS
ID : U01 AR066160
Pays : United States
Organisme : Medical Research Council
ID : G1000143
Pays : United Kingdom
Organisme : NIA NIH HHS
ID : R01 AG017917
Pays : United States
Organisme : NIDDK NIH HHS
ID : R01 DK107786
Pays : United States
Organisme : Medical Research Council
ID : MC_UU_12015/1
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : R01 HL105756
Pays : United States

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Auteurs

David Karasik (D)

Hebrew SeniorLife Institute for Aging Research and Harvard Medical School, Boston, MA.
Azrieli Faculty of Medicine, Bar-Ilan University, Safed, Israel.

M Carola Zillikens (MC)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.
Netherlands Genomics Initiative-Sponsored Netherlands Consortium for Healthy Aging, Leiden, The Netherlands.

Yi-Hsiang Hsu (YH)

Hebrew SeniorLife Institute for Aging Research and Harvard Medical School, Boston, MA.
Department of Medicine, Harvard Medical School, Boston, MA.
Molecular and Integrative Physiological Sciences Program, Harvard School of Public Health, Boston, MA.

Ali Aghdassi (A)

Department of Medicine A, University of Greifswald, Greifswald, Germany.

Kristina Akesson (K)

Department of Clinical Sciences Malmö, Lund University, Malmö, Sweden.
Department of Orthopedics, Skåne University Hospital, Malmö, Sweden.

Najaf Amin (N)

Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands.

Inês Barroso (I)

Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom.
NIHR Cambridge Biomedical Research Centre.
University of Cambridge Metabolic Research Laboratories, Institute of Metabolic Science, Addenbrooke's Hospital, Cambridge, United Kingdom.

David A Bennett (DA)

Rush Alzheimer's Disease Center, Rush University Medical Center, Chicago, IL.

Lars Bertram (L)

Lübeck Interdisciplinary Platform for Genome Analytics, University of Lübeck, Lübeck, Germany.

Murielle Bochud (M)

University Institute for Social and Preventive Medicine, Lausanne University Hospital, Lausanne, Switzerland.

Ingrid B Borecki (IB)

Division of Statistical Genomics, Department of Genetics.
Division of Biostatistics, Washington University School of Medicine, St. Louis, MO.

Linda Broer (L)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.
Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands.

Aron S Buchman (AS)

Rush Alzheimer's Disease Center, Rush University Medical Center, Chicago, IL.

Liisa Byberg (L)

Department of Surgical Sciences, Uppsala University, Uppsala, Sweden.

Harry Campbell (H)

Centre for Global Health Research, Usher Institute of Population Health Sciences and Informatics, University of Edinburgh, Teviot Place, Edinburgh, United Kingdom.

Natalia Campos-Obando (N)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.

Jane A Cauley (JA)

Department of Epidemiology, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA.

Peggy M Cawthon (PM)

California Pacific Medical Center Research Institute, San Francisco, CA.

John C Chambers (JC)

Cardiology, Ealing Hospital NHS Trust, London, United Kingdom.
Department of Epidemiology and Biostatistics, School of Public Health, Imperial College London, London, United Kingdom.
Imperial College Healthcare NHS Trust, London, United Kingdom.
NIHR Cardiovascular Biomedical Research Unit, Royal Brompton and Harefield NHS Foundation Trust and Imperial College London, London, United Kingdom.

Zhao Chen (Z)

Mel and Enid Zuckerman College of Public Health, University of Arizona, Tucson, AZ.

Nam H Cho (NH)

Department of Preventive Medicine, Ajou University School of Medicine, Youngtong-Gu, Suwon, Korea.

Hyung Jin Choi (HJ)

Department of Internal Medicine.
Department of Anatomy and Cell Biology, Neuroscience Research Institute, Seoul National University College of Medicine, Seoul, Korea.
Wide River Institute of Immunology, Seoul National University, Hongcheon, Korea.

Wen-Chi Chou (WC)

Hebrew SeniorLife Institute for Aging Research and Harvard Medical School, Boston, MA.
Broad Institute, Cambridge, MA.

Steven R Cummings (SR)

California Pacific Medical Center Research Institute, San Francisco, CA.

Lisette C P G M de Groot (LCPGM)

Division of Human Nutrition, Wageningen University, AFSG, Wageningen, The Netherlands.

Phillip L De Jager (PL)

Center for Translational and Computational Neuroimmunology, Neurology, Columbia University Medical Center, New York, NY.
Cell Circuits Program, Broad Institute, Cambridge, MA.

Ilja Demuth (I)

Charité-Universitätsmedizin Berlin, Freie Universität Berlin, Humboldt-Universität zu Berlin, Berlin, Germany.
Berlin Institute of Health, Berlin, Germany.

Luda Diatchenko (L)

Regional Center for Neurosensory Disorders, School of Dentistry, University of North Carolina at Chapel Hill, Chapel Hill, NC.
Alan Edwards Centre for Research on Pain, McGill University, Montreal, Canada.

Michael J Econs (MJ)

Department of Medicine and Department of Medical and Molecular Genetics, Indiana University School of Medicine, Indianapolis, IN.

Gudny Eiriksdottir (G)

Icelandic Heart Association Holtasmari, Kopavogur, Iceland.

Anke W Enneman (AW)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.

Joel Eriksson (J)

Department of Internal Medicine and Clinical Nutrition, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden.

Johan G Eriksson (JG)

National Institute for Health and Welfare, Helsinki, Finland.
Department of General Practice and Primary Health Care, University of Helsinki, Helsinki, Finland.
Helsinki University Central Hospital, Unit of General Practice, Helsinki, Finland.
Folkhalsan Research Centre, Helsinki, Finland.

Karol Estrada (K)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.
Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands.
Translational Biology, Biogen, Cambridge, MA.

Daniel S Evans (DS)

California Pacific Medical Center Research Institute, San Francisco, CA.

Mary F Feitosa (MF)

Division of Statistical Genomics, Department of Genetics.

Mao Fu (M)

Program in Personalized and Genomic Medicine, and Department of Medicine, Division of Endocrinology, Diabetes and Nutrition, University of Maryland School of Medicine, Baltimore, MD.

Christian Gieger (C)

Research Unit of Molecular Epidemiology.
Institute of Epidemiology II.
Institute of Genetic Epidemiology, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.

Harald Grallert (H)

Research Unit of Molecular Epidemiology.
Institute of Epidemiology II.
CCG Type 2 Diabetes, Helmholtz Zentrum München, Neuherberg, Germany.
German Center for Diabetes Research, Neuherberg, Germany.
CCG Nutrigenomics and Type 2 Diabetes, Helmholtz Zentrum München, Neuherberg, Germany.

Vilmundur Gudnason (V)

Icelandic Heart Association Holtasmari, Kopavogur, Iceland.
University of Iceland, Faculty of Medicine, Reykjavik, Iceland.

Launer J Lenore (LJ)

Laboratory of Epidemiology and Population Sciences, Intramural Research Program, NIA, Bethesda, MD.

Caroline Hayward (C)

MRC Human Genetics Unit, IGMM, University of Edinburgh, Edinburgh, United Kingdom.

Albert Hofman (A)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.
Netherlands Genomics Initiative-Sponsored Netherlands Consortium for Healthy Aging, Leiden, The Netherlands.

Georg Homuth (G)

Interfaculty Institute for Genetics and Functional Genomics, University of Greifswald, Greifswald, Germany.

Kim M Huffman (KM)

Duke Molecular Physiology Institute and Division of Rheumatology, Department of Medicine, Duke University School of Medicine, Durham, NC.

Lise B Husted (LB)

Aarhus University Hospital, Endocrinology and Internal Medicine, Aarhus, Denmark.

Thomas Illig (T)

Research Unit of Molecular Epidemiology.
Department of Human Genetics, Hannover Medical School, Hanover, Germany.

Erik Ingelsson (E)

Department of Medicine, Division of Cardiovascular Medicine, Stanford University School of Medicine, Stanford, CA.

Till Ittermann (T)

Institute for Community Medicine, University of Greifswald, Greifswald, Germany.

John-Olov Jansson (JO)

Department of Physiology, Institute of Neuroscience and Physiology, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden.

Toby Johnson (T)

University Institute for Social and Preventive Medicine, Lausanne University Hospital, Lausanne, Switzerland.
Department of Medical Genetics, University of Lausanne, Lausanne, Switzerland.
Swiss Institute of Bioinformatics, Lausanne, Switzerland.

Reiner Biffar (R)

Centre of Oral Health, Department of Prosthetic Dentistry, Gerodontology and Biomaterials, University of Greifswald, Greifswald, Germany.

Joanne M Jordan (JM)

Thurston Arthritis Research Center, University of North Carolina at Chapel Hill, Chapel Hill, NC.

Antti Jula (A)

National Institute for Health and Welfare, Helsinki, Finland.

Magnus Karlsson (M)

Department of Clinical Sciences Malmö, Lund University, Malmö, Sweden.
Department of Orthopedics, Skåne University Hospital, Malmö, Sweden.

Kay-Tee Khaw (KT)

Department of Public Health and Primary Care, University of Cambridge, Cambridge, United Kingdom.

Tuomas O Kilpeläinen (TO)

MRC Epidemiology Unit, University of Cambridge School of Clinical Medicine, Cambridge Biomedical Campus, Cambridge, United Kingdom.
Novo Nordisk Foundation Center for Basic Metabolic Research, Section of Metabolic Genetics, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark.
Department of Environmental Medicine and Public Health, Icahn School of Medicine at Mount Sinai, New York, NY.

Norman Klopp (N)

Research Unit of Molecular Epidemiology.
Hannover Unified Biobank, Hannover Medical School, Hannover, Germany.

Jacqueline S L Kloth (JSL)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.

Daniel L Koller (DL)

Department of Medicine and Department of Medical and Molecular Genetics, Indiana University School of Medicine, Indianapolis, IN.

Jaspal S Kooner (JS)

Cardiology, Ealing Hospital NHS Trust, London, United Kingdom.
Imperial College Healthcare NHS Trust, London, United Kingdom.
Faculty of Medicine, National Heart & Lung Institute, Cardiovascular Science, Hammersmith Campus, Hammersmith Hospital, Imperial College London, United Kingdom.

William E Kraus (WE)

Duke Molecular Physiology Institute and Division of Cardiology, Department of Medicine, Duke University School of Medicine, Durham, NC.

Stephen Kritchevsky (S)

Sticht Center for Healthy Aging and Alzheimer's Prevention, Wake Forest School of Medicine, Winston-Salem, NC.

Zoltán Kutalik (Z)

University Institute for Social and Preventive Medicine, Lausanne University Hospital, Lausanne, Switzerland.
Swiss Institute of Bioinformatics, Lausanne, Switzerland.

Teemu Kuulasmaa (T)

Department of Medicine, University of Eastern Finland and Kuopio University Hospital, Kuopio, Finland.

Johanna Kuusisto (J)

Department of Medicine, University of Eastern Finland and Kuopio University Hospital, Kuopio, Finland.

Markku Laakso (M)

Department of Medicine, University of Eastern Finland and Kuopio University Hospital, Kuopio, Finland.

Jari Lahti (J)

Helsinki Collegium Advanced Studies, University of Helsinki, Helsinki, Finland.

Thomas Lang (T)

Department of Radiology and Biomedical Imaging, and School of Dentistry, UC San Francisco, San Francisco, CA.

Bente L Langdahl (BL)

Aarhus University Hospital, Endocrinology and Internal Medicine, Aarhus, Denmark.

Markus M Lerch (MM)

Department of Medicine A, University of Greifswald, Greifswald, Germany.

Joshua R Lewis (JR)

School of Medicine and Pharmacology, University of Western Australia, Perth, Australia.
Centre for Kidney Research, Children's Hospital at Westmead, School of Public Health, Sydney Medical School, The University of Sydney, Sydney, Australia.

Christina Lill (C)

Institute of Neurogenetics, University of Luebeck, Luebeck, Germany.

Lars Lind (L)

Department. of Medical Sciences, Uppsala University, Uppsala, Sweden.

Cecilia Lindgren (C)

Wellcome Trust Centre for Human Genetics, Oxford University, Oxford, United Kingdom.

Yongmei Liu (Y)

Department of Epidemiology and Prevention, Wake Forest School of Medicine, Winston-Salem, NC.

Gregory Livshits (G)

Department of Anatomy and Anthropology, Sackler Faculty of Medicine, Tel Aviv University, Tel Aviv, Israel.
Department of Twin Research and Genetic Epidemiology, King's College London, St Thomas' Campus, London, United Kingdom.

Östen Ljunggren (Ö)

Department. of Medical Sciences, Uppsala University, Uppsala, Sweden.

Ruth J F Loos (RJF)

Department of Environmental Medicine and Public Health, Icahn School of Medicine at Mount Sinai, New York, NY.
The Charles Bronfman Institute of Personalized Medicine.
Institute of Child Health and Development.
The Genetics of Obesity and Related Traits Program, Icahn School of Medicine at Mount Sinai, New York, NY.

Mattias Lorentzon (M)

Department of Internal Medicine, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden.

Jian'an Luan (J)

Department of Environmental Medicine and Public Health, Icahn School of Medicine at Mount Sinai, New York, NY.

Robert N Luben (RN)

Novo Nordisk Foundation Center for Basic Metabolic Research, Section of Metabolic Genetics, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark.

Ida Malkin (I)

Department of Anatomy and Anthropology, Sackler Faculty of Medicine, Tel Aviv University, Tel Aviv, Israel.

Fiona E McGuigan (FE)

Department of Clinical Sciences Malmö, Lund University, Malmö, Sweden.

Carolina Medina-Gomez (C)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.
Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands.

Thomas Meitinger (T)

Institute of Human Genetics, MRI, Technische Universität München, Munich, Germany.
Institute of Human Genetics, Helmholtz Zentrum München-German Research Center for Environmental Health, Neuherberg, Germany.

Håkan Melhus (H)

Department. of Medical Sciences, Uppsala University, Uppsala, Sweden.

Dan Mellström (D)

Department of Internal Medicine and Clinical Nutrition, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden.

Karl Michaëlsson (K)

Department of Surgical Sciences, Uppsala University, Uppsala, Sweden.

Braxton D Mitchell (BD)

Program in Personalized and Genomic Medicine, and Department of Medicine, Division of Endocrinology, Diabetes and Nutrition, University of Maryland School of Medicine, Baltimore, MD.
Geriatric Research and Education Clinical Center-Veterans Administration Medical Center, Baltimore, MD.

Andrew P Morris (AP)

Wellcome Trust Centre for Human Genetics, Oxford University, Oxford, United Kingdom.
Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom.

Leif Mosekilde (L)

Aarhus University Hospital, Endocrinology and Internal Medicine, Aarhus, Denmark.

Maria Nethander (M)

Department of Internal Medicine and Clinical Nutrition, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden.

Anne B Newman (AB)

Department of Epidemiology, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA.

Jeffery R O'Connell (JR)

Program in Personalized and Genomic Medicine, and Department of Medicine, Division of Endocrinology, Diabetes and Nutrition, University of Maryland School of Medicine, Baltimore, MD.

Ben A Oostra (BA)

Department of Clinical Genetics, Erasmus MC, Rotterdam, The Netherlands.
Centre for Medical Systems Biology & Netherlands Consortium on Healthy Aging, Leiden, The Netherlands.

Eric S Orwoll (ES)

Oregon Health & Science University, Portland, OR.

Aarno Palotie (A)

Institute for Molecular Medicine Finland, University of Helsinki, Helsinki, Finland.
Department of Medical Genetics, University of Helsinki and University Central Hospital, Helsinki, Finland.

Munro Peacock (M)

Department of Medicine and Department of Medical and Molecular Genetics, Indiana University School of Medicine, Indianapolis, IN.

Markus Perola (M)

National Institute for Health and Welfare, Helsinki, Finland.
University of Helsinki, Institute for Molecular Medicine, Finland and Diabetes and Obesity Research Program, Helsinki, Finland.
University of Tartu, Estonian Genome Center, Tartu, Estonia.

Annette Peters (A)

Research Unit of Molecular Epidemiology.
Institute of Epidemiology II.

Richard L Prince (RL)

School of Medicine and Pharmacology, University of Western Australia, Perth, Australia.
Department of Endocrinology and Diabetes, Sir Charles Gardiner Hospital, Perth, Australia.

Bruce M Psaty (BM)

Cardiovascular Health Research Unit, Departments of Medicine, Epidemiology, and Health Services, University of Washington, Seattle, WA.
Kaiser Permanente Washington Health Research Institute, Seattle, WA.

Katri Räikkönen (K)

Department of Psychology and Logopedics, University of Helsinki, Helsinki, Finland.

Stuart H Ralston (SH)

Molecular Medicine Centre, MRC Institute of Genetics and Molecular Medicine, Western General Hospital, Edinburgh, United Kingdom.

Samuli Ripatti (S)

Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom.
Institute for Molecular Medicine Finland, University of Helsinki, Helsinki, Finland.
Hjelt Institute, University of Helsinki, Helsinki, Finland.

Fernando Rivadeneira (F)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.
Netherlands Genomics Initiative-Sponsored Netherlands Consortium for Healthy Aging, Leiden, The Netherlands.
Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands.

John A Robbins (JA)

Department of Medicine, University California at Davis, Sacramento, CA.

Jerome I Rotter (JI)

Institute for Translational Genomic and Population Sciences, Los Angeles Biomedical Research Institute and Department of Pediatrics, Harbor UCLA Medical Center, Torrance, CA.

Igor Rudan (I)

Department of Epidemiology, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, PA.

Veikko Salomaa (V)

National Institute for Health and Welfare, Helsinki, Finland.

Suzanne Satterfield (S)

Department of Preventive Medicine, University of Tennessee Health Science Center, Memphis, TN.

Sabine Schipf (S)

Institute for Community Medicine, University of Greifswald, Greifswald, Germany.

Chan Soo Shin (CS)

Department of Internal Medicine.

Albert V Smith (AV)

Icelandic Heart Association Holtasmari, Kopavogur, Iceland.
University of Iceland, Faculty of Medicine, Reykjavik, Iceland.

Shad B Smith (SB)

Center for Translational Pain Medicine, Department of Anesthesiology, Duke University, Durham, NC.

Nicole Soranzo (N)

Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom.

Timothy D Spector (TD)

Department of Twin Research and Genetic Epidemiology, King's College London, St Thomas' Campus, London, United Kingdom.

Alena Stancáková (A)

Department of Medicine, University of Eastern Finland and Kuopio University Hospital, Kuopio, Finland.

Kari Stefansson (K)

University of Iceland, Faculty of Medicine, Reykjavik, Iceland.
deCODE Genetics, Reykjavik, Iceland.

Elisabeth Steinhagen-Thiessen (E)

Charité-Universitätsmedizin Berlin, Freie Universität Berlin, Humboldt-Universität zu Berlin, Berlin, Germany.

Lisette Stolk (L)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.
Netherlands Genomics Initiative-Sponsored Netherlands Consortium for Healthy Aging, Leiden, The Netherlands.

Elizabeth A Streeten (EA)

Program in Personalized and Genomic Medicine, and Department of Medicine, Division of Endocrinology, Diabetes and Nutrition, University of Maryland School of Medicine, Baltimore, MD.
Geriatric Research and Education Clinical Center-Veterans Administration Medical Center, Baltimore, MD.

Unnur Styrkarsdottir (U)

deCODE Genetics, Reykjavik, Iceland.

Karin M A Swart (KMA)

Department of Epidemiology and Biostatistics, and the EMGO+ Institute; VU University Medical Center, Amsterdam, The Netherlands.

Patricia Thompson (P)

Department of Pathology, Stony Brook University, Stony Brook, NY.

Cynthia A Thomson (CA)

Mel and Enid Zuckerman College of Public Health, University of Arizona, Tucson, AZ.

Gudmar Thorleifsson (G)

deCODE Genetics, Reykjavik, Iceland.

Unnur Thorsteinsdottir (U)

University of Iceland, Faculty of Medicine, Reykjavik, Iceland.
deCODE Genetics, Reykjavik, Iceland.

Emmi Tikkanen (E)

National Institute for Health and Welfare, Helsinki, Finland.
Institute for Molecular Medicine Finland, University of Helsinki, Helsinki, Finland.

Gregory J Tranah (GJ)

California Pacific Medical Center Research Institute, San Francisco, CA.

André G Uitterlinden (AG)

Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands.
Netherlands Genomics Initiative-Sponsored Netherlands Consortium for Healthy Aging, Leiden, The Netherlands.
Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands.

Cornelia M van Duijn (CM)

Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands.
Centre for Medical Systems Biology & Netherlands Consortium on Healthy Aging, Leiden, The Netherlands.

Natasja M van Schoor (NM)

Department of Epidemiology and Biostatistics, and the EMGO+ Institute; VU University Medical Center, Amsterdam, The Netherlands.

Liesbeth Vandenput (L)

Department of Internal Medicine and Clinical Nutrition, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden.

Peter Vollenweider (P)

Department of Medicine, Internal Medicine, Lausanne University Hospital and Faculty of Biology and Medicine, Lausanne, Switzerland.

Henry Völzke (H)

Institute for Community Medicine, University of Greifswald, Greifswald, Germany.

Jean Wactawski-Wende (J)

Department of Epidemiology and Environmental Health, University at Buffalo, SUNY, Buffalo, NY.

Mark Walker (M)

Institute of Cellular Medicine (Diabetes), The Medical School, Newcastle University, Framlington Place, Newcastle upon Tyne, United Kingdom.

Nicholas J Wareham (N)

MRC Epidemiology Unit, University of Cambridge School of Clinical Medicine, Cambridge Biomedical Campus, Cambridge, United Kingdom.

Dawn Waterworth (D)

Genetics, GlaxoSmithKline, King of Prussia, PA.

Michael N Weedon (MN)

Genetics of Complex Traits, Royal Devon & Exeter Hospital, University of Exeter Medical School, Exeter, United Kingdom.

H-Erich Wichmann (HE)

Institute of Epidemiology II.
Institute of Medical Informatics, Biometry and Epidemiology, Ludwig-Maximilians-Universität, Neuherberg/Munich, Germany.
Institute of Medical Statistics and Epidemiology, Technical University, Munich, Germany.

Elisabeth Widen (E)

Institute for Molecular Medicine Finland, University of Helsinki, Helsinki, Finland.

Frances M K Williams (FMK)

Department of Twin Research and Genetic Epidemiology, King's College London, St Thomas' Campus, London, United Kingdom.

James F Wilson (JF)

Centre for Global Health Research, Usher Institute of Population Health Sciences and Informatics, University of Edinburgh, Teviot Place, Edinburgh, United Kingdom.

Nicole C Wright (NC)

Department of Epidemiology, University of Alabama at Birmingham, Birmingham, AL.

Laura M Yerges-Armstrong (LM)

Program in Personalized and Genomic Medicine, and Department of Medicine, Division of Endocrinology, Diabetes and Nutrition, University of Maryland School of Medicine, Baltimore, MD.
Genetics, GlaxoSmithKline, King of Prussia, PA.

Lei Yu (L)

Rush Alzheimer's Disease Center, Rush University Medical Center, Chicago, IL.

Weihua Zhang (W)

Cardiology, Ealing Hospital NHS Trust, London, United Kingdom.
Imperial College Healthcare NHS Trust, London, United Kingdom.

Jing Hua Zhao (JH)

MRC Epidemiology Unit, University of Cambridge School of Clinical Medicine, Cambridge Biomedical Campus, Cambridge, United Kingdom.

Yanhua Zhou (Y)

Department of Biostatistics, Boston University School of Public Health, Boston, MA.

Carrie M Nielson (CM)

Oregon Health & Science University, Portland, OR.

Tamara B Harris (TB)

Laboratory of Epidemiology and Population Sciences, Intramural Research Program, NIA, Bethesda, MD.

Serkalem Demissie (S)

Department of Biostatistics, Boston University School of Public Health, Boston, MA.

Douglas P Kiel (DP)

Hebrew SeniorLife Institute for Aging Research and Harvard Medical School, Boston, MA.
Department of Medicine, Harvard Medical School, Boston, MA.

Claes Ohlsson (C)

Department of Internal Medicine and Clinical Nutrition, Institute of Medicine, Sahlgrenska Academy, University of Gothenburg, Gothenburg, Sweden.

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