Description, molecular characteristics and Wolbachia endosymbionts of Onchocerca borneensis Uni, Mat Udin & Takaoka n. sp. (Nematoda: Filarioidea) from the Bornean bearded pig Sus barbatus Müller (Cetartiodactyla: Suidae) of Sarawak, Malaysia.


Journal

Parasites & vectors
ISSN: 1756-3305
Titre abrégé: Parasit Vectors
Pays: England
ID NLM: 101462774

Informations de publication

Date de publication:
06 Feb 2020
Historique:
received: 29 08 2019
accepted: 15 01 2020
entrez: 8 2 2020
pubmed: 8 2 2020
medline: 6 6 2020
Statut: epublish

Résumé

The genus Onchocerca Diesing, 1841 includes species of medical importance, such as O. volvulus (Leuckart, 1893), which causes river blindness in the tropics. Recently, zoonotic onchocercosis has been reported in humans worldwide. In Japan, O. dewittei japonica Uni, Bain & Takaoka, 2001 from wild boars is a causative agent for this zoonosis. Many filarioid nematodes are infected with Wolbachia endosymbionts which exhibit various evolutionary relationships with their hosts. While investigating the filarial fauna of Borneo, we discovered an undescribed Onchocerca species in the bearded pig Sus barbatus Müller (Cetartiodactyla: Suidae). We isolated Onchocerca specimens from bearded pigs and examined their morphology. For comparative material, we collected fresh specimens of O. d. dewittei Bain, Ramachandran, Petter & Mak, 1977 from banded pigs (S. scrofa vittatus Boie) in Peninsular Malaysia. Partial sequences of three different genes (two mitochondrial genes, cox1 and 12S rRNA, and one nuclear ITS region) of these filarioids were analysed. By multi-locus sequence analyses based on six genes (16S rDNA, ftsZ, dnaA, coxA, fbpA and gatB) of Wolbachia, we determined the supergroups in the specimens from bearded pigs and those of O. d. dewittei. Onchocerca borneensis Uni, Mat Udin & Takaoka n. sp. is described on the basis of morphological characteristics and its genetic divergence from congeners. Molecular characteristics of the new species revealed its close evolutionary relationship with O. d. dewittei. Calculated p-distance for the cox1 gene sequences between O. borneensis n. sp. and O. d. dewittei was 5.9%, while that between O. d. dewittei and O. d. japonica was 7.6%. No intraspecific genetic variation was found for the new species. Wolbachia strains identified in the new species and O. d. dewittei belonged to supergroup C and are closely related. Our molecular analyses of filarioids from Asian suids indicate that the new species is sister to O. d. dewittei. On the basis of its morphological and molecular characteristics, we propose to elevate O. d. japonica to species level as O. japonica Uni, Bain & Takaoka, 2001. Coevolutionary relationships exist between the Wolbachia strains and their filarial hosts in Borneo and Peninsular Malaysia.

Sections du résumé

BACKGROUND BACKGROUND
The genus Onchocerca Diesing, 1841 includes species of medical importance, such as O. volvulus (Leuckart, 1893), which causes river blindness in the tropics. Recently, zoonotic onchocercosis has been reported in humans worldwide. In Japan, O. dewittei japonica Uni, Bain & Takaoka, 2001 from wild boars is a causative agent for this zoonosis. Many filarioid nematodes are infected with Wolbachia endosymbionts which exhibit various evolutionary relationships with their hosts. While investigating the filarial fauna of Borneo, we discovered an undescribed Onchocerca species in the bearded pig Sus barbatus Müller (Cetartiodactyla: Suidae).
METHODS METHODS
We isolated Onchocerca specimens from bearded pigs and examined their morphology. For comparative material, we collected fresh specimens of O. d. dewittei Bain, Ramachandran, Petter & Mak, 1977 from banded pigs (S. scrofa vittatus Boie) in Peninsular Malaysia. Partial sequences of three different genes (two mitochondrial genes, cox1 and 12S rRNA, and one nuclear ITS region) of these filarioids were analysed. By multi-locus sequence analyses based on six genes (16S rDNA, ftsZ, dnaA, coxA, fbpA and gatB) of Wolbachia, we determined the supergroups in the specimens from bearded pigs and those of O. d. dewittei.
RESULTS RESULTS
Onchocerca borneensis Uni, Mat Udin & Takaoka n. sp. is described on the basis of morphological characteristics and its genetic divergence from congeners. Molecular characteristics of the new species revealed its close evolutionary relationship with O. d. dewittei. Calculated p-distance for the cox1 gene sequences between O. borneensis n. sp. and O. d. dewittei was 5.9%, while that between O. d. dewittei and O. d. japonica was 7.6%. No intraspecific genetic variation was found for the new species. Wolbachia strains identified in the new species and O. d. dewittei belonged to supergroup C and are closely related.
CONCLUSIONS CONCLUSIONS
Our molecular analyses of filarioids from Asian suids indicate that the new species is sister to O. d. dewittei. On the basis of its morphological and molecular characteristics, we propose to elevate O. d. japonica to species level as O. japonica Uni, Bain & Takaoka, 2001. Coevolutionary relationships exist between the Wolbachia strains and their filarial hosts in Borneo and Peninsular Malaysia.

Identifiants

pubmed: 32028994
doi: 10.1186/s13071-020-3907-8
pii: 10.1186/s13071-020-3907-8
pmc: PMC7006428
doi:

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

50

Subventions

Organisme : The Ministry of Higher Education, Malaysia
ID : FRGS EP020-2012

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Auteurs

Shigehiko Uni (S)

Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603, Kuala Lumpur, Malaysia. unishigehiko@um.edu.my.
Department of Public Health, Faculty of Nursing, Kobe Women's University, Kobe, 650-0046, Japan. unishigehiko@um.edu.my.

Ahmad Syihan Mat Udin (AS)

Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Takeshi Agatsuma (T)

Department of Environmental Medicine, Kochi Medical School, Kochi University, Nankoku, 783-8505, Japan.

Kerstin Junker (K)

ARC-Onderstepoort Veterinary Institute, Private Bag X05, Onderstepoort, 0110, South Africa.

Weerachai Saijuntha (W)

Walai Rukhavej Botanical Research Institute, Mahasarakham University, Maha Sarakham, 44150, Thailand.

Naruemon Bunchom (N)

Department of Environmental Medicine, Kochi Medical School, Kochi University, Nankoku, 783-8505, Japan.
Walai Rukhavej Botanical Research Institute, Mahasarakham University, Maha Sarakham, 44150, Thailand.

Masako Fukuda (M)

Institute for Research Promotion, Oita University, Oita, 879-5593, Japan.

Coralie Martin (C)

UMR7245, MCAM, Muséum National d'Histoire Naturelle, 75005, Paris, France.

Emilie Lefoulon (E)

Molecular Parasitology Group, New England Biolabs, Inc, Ipswich, MA, 01938, USA.

Amandine Labat (A)

UMR7245, MCAM, Muséum National d'Histoire Naturelle, 75005, Paris, France.

Faisal Ali Anwarali Khan (FAA)

Department of Zoology, Faculty of Resource Sciences and Technology, Universiti Malaysia Sarawak, 943800, Kota Samarahan, Sarawak, Malaysia.

Van Lun Low (VL)

Tropical Infectious Diseases Research & Education Centre, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Phaik Leng Cheah (PL)

Department of Pathology, Faculty of Medicine, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Yvonne Ai-Lian Lim (YA)

Department of Parasitology, Faculty of Medicine, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Rosli Ramli (R)

Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Daicus Martin Belabut (DM)

Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Nur Afiqah Zainuri (NA)

Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Makoto Matsubayashi (M)

Department of International Prevention of Epidemics, Division of Veterinary Science, Graduate School of Life and Environmental Sciences, Osaka Prefecture University, Osaka, 598-8531, Japan.

Hasmahzaiti Omar (H)

Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Subha Bhassu (S)

Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603, Kuala Lumpur, Malaysia.
Centre for Biotechnology in Agriculture, CEBAR, University of Malaya, 50300, Kuala Lumpur, Malaysia.

Shoji Uga (S)

Department of Public Health, Faculty of Nursing, Kobe Women's University, Kobe, 650-0046, Japan.

Rosli Hashim (R)

Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Hiroyuki Takaoka (H)

Tropical Infectious Diseases Research & Education Centre, University of Malaya, 50603, Kuala Lumpur, Malaysia.

Mohd Sofian Azirun (MS)

Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603, Kuala Lumpur, Malaysia.

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