Novel alternative ribonucleotide excision repair pathways in human cells by DDX3X and specialized DNA polymerases.


Journal

Nucleic acids research
ISSN: 1362-4962
Titre abrégé: Nucleic Acids Res
Pays: England
ID NLM: 0411011

Informations de publication

Date de publication:
18 11 2020
Historique:
accepted: 08 10 2020
revised: 02 10 2020
received: 21 01 2020
pubmed: 3 11 2020
medline: 4 2 2021
entrez: 2 11 2020
Statut: ppublish

Résumé

Removal of ribonucleotides (rNMPs) incorporated into the genome by the ribonucleotide excision repair (RER) is essential to avoid genetic instability. In eukaryotes, the RNaseH2 is the only known enzyme able to incise 5' of the rNMP, starting the RER process, which is subsequently carried out by replicative DNA polymerases (Pols) δ or ϵ, together with Flap endonuclease 1 (Fen-1) and DNA ligase 1. Here, we show that the DEAD-box RNA helicase DDX3X has RNaseH2-like activity and can support fully reconstituted in vitro RER reactions, not only with Pol δ but also with the repair Pols β and λ. Silencing of DDX3X causes accumulation of rNMPs in the cellular genome. These results support the existence of alternative RER pathways conferring high flexibility to human cells in responding to the threat posed by rNMPs incorporation.

Identifiants

pubmed: 33137198
pii: 5952198
doi: 10.1093/nar/gkaa948
pmc: PMC7672437
doi:

Substances chimiques

Ribonucleotides 0
Adenosine Triphosphate 8L70Q75FXE
DNA polymerase beta2 EC 2.7.7.-
DNA Polymerase beta EC 2.7.7.7
ribonuclease HII EC 3.1.26.-
Ribonuclease H EC 3.1.26.4
DDX3X protein, human EC 3.6.1.-
DEAD-box RNA Helicases EC 3.6.4.13

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

11551-11565

Informations de copyright

© The Author(s) 2020. Published by Oxford University Press on behalf of Nucleic Acids Research.

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Auteurs

Valentina Riva (V)

Institute of Molecular Genetics IGM-CNR 'Luigi Luca Cavalli-Sforza', via Abbiategrasso 207, I-27100 Pavia, Italy.

Anna Garbelli (A)

Institute of Molecular Genetics IGM-CNR 'Luigi Luca Cavalli-Sforza', via Abbiategrasso 207, I-27100 Pavia, Italy.

Federica Casiraghi (F)

Institute of Molecular Genetics IGM-CNR 'Luigi Luca Cavalli-Sforza', via Abbiategrasso 207, I-27100 Pavia, Italy.

Francesca Arena (F)

Institute of Molecular Genetics IGM-CNR 'Luigi Luca Cavalli-Sforza', via Abbiategrasso 207, I-27100 Pavia, Italy.

Claudia Immacolata Trivisani (CI)

Department of Biotechnology, Chemistry and Pharmacy, University of Siena, Via A. De Gasperi 2, I-53100 Siena, Italy.

Assunta Gagliardi (A)

Department of Life Sciences, Via A. Moro 2, University of Siena, I-53100 Siena, Italy.

Luca Bini (L)

Department of Life Sciences, Via A. Moro 2, University of Siena, I-53100 Siena, Italy.

Martina Schroeder (M)

Kathleen Lonsdale Institute for Human Health Research, Biology Department, Maynooth University, Maynooth, Co. Kildare, Ireland.

Antonio Maffia (A)

Institute of Molecular Genetics IGM-CNR 'Luigi Luca Cavalli-Sforza', via Abbiategrasso 207, I-27100 Pavia, Italy.

Simone Sabbioneda (S)

Institute of Molecular Genetics IGM-CNR 'Luigi Luca Cavalli-Sforza', via Abbiategrasso 207, I-27100 Pavia, Italy.

Giovanni Maga (G)

Institute of Molecular Genetics IGM-CNR 'Luigi Luca Cavalli-Sforza', via Abbiategrasso 207, I-27100 Pavia, Italy.

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Classifications MeSH