Conserved Gsx2/Ind homeodomain monomer versus homodimer DNA binding defines regulatory outcomes in flies and mice.


Journal

Genes & development
ISSN: 1549-5477
Titre abrégé: Genes Dev
Pays: United States
ID NLM: 8711660

Informations de publication

Date de publication:
01 01 2021
Historique:
received: 28 07 2020
accepted: 19 11 2020
pubmed: 19 12 2020
medline: 5 8 2021
entrez: 18 12 2020
Statut: ppublish

Résumé

How homeodomain proteins gain sufficient specificity to control different cell fates has been a long-standing problem in developmental biology. The conserved Gsx homeodomain proteins regulate specific aspects of neural development in animals from flies to mammals, and yet they belong to a large transcription factor family that bind nearly identical DNA sequences in vitro. Here, we show that the mouse and fly Gsx factors unexpectedly gain DNA binding specificity by forming cooperative homodimers on precisely spaced and oriented DNA sites. High-resolution genomic binding assays revealed that Gsx2 binds both monomer and homodimer sites in the developing mouse ventral telencephalon. Importantly, reporter assays showed that Gsx2 mediates opposing outcomes in a DNA binding site-dependent manner: Monomer Gsx2 binding represses transcription, whereas homodimer binding stimulates gene expression. In

Identifiants

pubmed: 33334823
pii: gad.343053.120
doi: 10.1101/gad.343053.120
pmc: PMC7778271
doi:

Substances chimiques

Drosophila Proteins 0
Gsh2 protein, mouse 0
Homeodomain Proteins 0
IND protein, Drosophila 0

Types de publication

Journal Article Research Support, N.I.H., Extramural Research Support, Non-U.S. Gov't Research Support, U.S. Gov't, Non-P.H.S.

Langues

eng

Sous-ensembles de citation

IM

Pagination

157-174

Subventions

Organisme : NIGMS NIH HHS
ID : R01 GM079428
Pays : United States
Organisme : NINDS NIH HHS
ID : R01 NS044080
Pays : United States
Organisme : NINDS NIH HHS
ID : R01 NS069893
Pays : United States
Organisme : NIGMS NIH HHS
ID : T32 GM063483
Pays : United States

Informations de copyright

© 2021 Salomone et al.; Published by Cold Spring Harbor Laboratory Press.

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Auteurs

Joseph Salomone (J)

Graduate Program in Molecular and Developmental Biology, Cincinnati Children's Hospital Research Foundation, Cincinnati, Ohio 45229, USA.
Medical-Scientist Training Program, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.

Shenyue Qin (S)

Division of Developmental Biology, Cincinnati Children's Hospital Medical Center, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.

Temesgen D Fufa (TD)

Ophthalmic Genetics and Visual Function Branch, National Eye Institute, National Institutes of Health, Bethesda, Maryland 20892, USA.

Brittany Cain (B)

Department of Biomedical Engineering, University of Cincinnati, Cincinnati, Ohio 45219, USA.

Edward Farrow (E)

Medical-Scientist Training Program, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.

Bin Guan (B)

Ophthalmic Genetics and Visual Function Branch, National Eye Institute, National Institutes of Health, Bethesda, Maryland 20892, USA.

Robert B Hufnagel (RB)

Ophthalmic Genetics and Visual Function Branch, National Eye Institute, National Institutes of Health, Bethesda, Maryland 20892, USA.

Masato Nakafuku (M)

Division of Developmental Biology, Cincinnati Children's Hospital Medical Center, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.
Department of Pediatrics, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.

Hee-Woong Lim (HW)

Department of Pediatrics, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.
Division of Biomedical Informatics, Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio 45229, USA.
Department of Biomedical Informatics, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.

Kenneth Campbell (K)

Division of Developmental Biology, Cincinnati Children's Hospital Medical Center, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.
Department of Pediatrics, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.

Brian Gebelein (B)

Division of Developmental Biology, Cincinnati Children's Hospital Medical Center, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.
Department of Pediatrics, University of Cincinnati College of Medicine, Cincinnati, Ohio 45229, USA.

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Classifications MeSH