A loss-of-function variant in SUV39H2 identified in autism-spectrum disorder causes altered H3K9 trimethylation and dysregulation of protocadherin β-cluster genes in the developing brain.


Journal

Molecular psychiatry
ISSN: 1476-5578
Titre abrégé: Mol Psychiatry
Pays: England
ID NLM: 9607835

Informations de publication

Date de publication:
12 2021
Historique:
received: 27 01 2021
accepted: 10 06 2021
revised: 21 05 2021
pubmed: 16 7 2021
medline: 15 3 2022
entrez: 15 7 2021
Statut: ppublish

Résumé

Recent evidence has documented the potential roles of histone-modifying enzymes in autism-spectrum disorder (ASD). Aberrant histone H3 lysine 9 (H3K9) dimethylation resulting from genetic variants in histone methyltransferases is known for neurodevelopmental and behavioral anomalies. However, a systematic examination of H3K9 methylation dynamics in ASD is lacking. Here we resequenced nine genes for histone methyltransferases and demethylases involved in H3K9 methylation in individuals with ASD and healthy controls using targeted next-generation sequencing. We identified a novel rare variant (A211S) in the SUV39H2, which was predicted to be deleterious. The variant showed strongly reduced histone methyltransferase activity in vitro. In silico analysis showed that the variant destabilizes the hydrophobic core and allosterically affects the enzyme activity. The Suv39h2-KO mice displayed hyperactivity and reduced behavioral flexibility in learning the tasks that required complex behavioral adaptation, which is relevant for ASD. The Suv39h2 deficit evoked an elevated expression of a subset of protocadherin β (Pcdhb) cluster genes in the embryonic brain, which is attributable to the loss of H3K9 trimethylation (me3) at the gene promoters. Reduced H3K9me3 persisted in the cerebellum of Suv39h2-deficient mice to an adult stage. Congruently, reduced expression of SUV39H1 and SUV39H2 in the postmortem brain samples of ASD individuals was observed, underscoring the role of H3K9me3 deficiency in ASD etiology. The present study provides direct evidence for the role of SUV39H2 in ASD and suggests a molecular cascade of SUV39H2 dysfunction leading to H3K9me3 deficiency followed by an untimely, elevated expression of Pcdhb cluster genes during early neurodevelopment.

Identifiants

pubmed: 34262135
doi: 10.1038/s41380-021-01199-7
pii: 10.1038/s41380-021-01199-7
doi:

Substances chimiques

Histones 0
Protocadherins 0
Histone-Lysine N-Methyltransferase EC 2.1.1.43
Suv39h2 protein, mouse EC 2.1.1.43

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

7550-7559

Informations de copyright

© 2021. The Author(s), under exclusive licence to Springer Nature Limited.

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Auteurs

Shabeesh Balan (S)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan. shabeeshbalan@riken.jp.
Neuroscience Research Laboratory, Institute of Mental Health and Neurosciences (IMHANS), Kozhikode, Kerala, India. shabeeshbalan@riken.jp.

Yoshimi Iwayama (Y)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan.

Tetsuo Ohnishi (T)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan.
Department of Molecular Neuroscience, Medical Research Institute, Tokyo Medical and Dental University, Tokyo, Japan.

Mikiko Fukuda (M)

Cellular Memory Laboratory, RIKEN Cluster for Pioneering Research, Wako, Saitama, Japan.

Atsuko Shirai (A)

Cellular Memory Laboratory, RIKEN Cluster for Pioneering Research, Wako, Saitama, Japan.

Ayumi Yamada (A)

Cellular Memory Laboratory, RIKEN Cluster for Pioneering Research, Wako, Saitama, Japan.

Sara Weirich (S)

Department of Biochemistry, Institute of Biochemistry and Technical Biochemistry, University of Stuttgart, Allmandring 31, Stuttgart, Germany.

Maren Kirstin Schuhmacher (MK)

Department of Biochemistry, Institute of Biochemistry and Technical Biochemistry, University of Stuttgart, Allmandring 31, Stuttgart, Germany.

Kalarickal Vijayan Dileep (KV)

Laboratory for Structural Bioinformatics, RIKEN Center for Biosystems Dynamics Research, Yokohama, Kanagawa, Japan.

Toshihiro Endo (T)

Phenovance Research and Technology LLC, Kashiwa, Chiba, Japan.

Yasuko Hisano (Y)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan.

Kaoru Kotoshiba (K)

Cellular Memory Laboratory, RIKEN Cluster for Pioneering Research, Wako, Saitama, Japan.

Tomoko Toyota (T)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan.

Takeshi Otowa (T)

Department of Neuropsychiatry, NTT Medical Center Tokyo, Tokyo, Japan.

Hitoshi Kuwabara (H)

Department of Psychiatry, Hamamatsu University School of Medicine, Hamamatsu, Shizuoka, Japan.

Mamoru Tochigi (M)

Department of Neuropsychiatry, Teikyo University School of Medicine, Tokyo, Japan.

Akiko Watanabe (A)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan.

Hisako Ohba (H)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan.

Motoko Maekawa (M)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan.

Manabu Toyoshima (M)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan.

Tsukasa Sasaki (T)

Laboratory of Health Education, Graduate School of Education, The University of Tokyo, Tokyo, Japan.

Kazuhiko Nakamura (K)

Department of Neuropsychiatry, Hirosaki University School of Medicine, Hirosaki, Aomori, Japan.

Masatsugu Tsujii (M)

Faculty of Contemporary Sociology, Chukyo University, Toyota, Aichi, Japan.

Hideo Matsuzaki (H)

Research Center for Child Mental Development, University of Fukui, Yoshida-gun, Fukui, Japan.

Kam Y J Zhang (KYJ)

Laboratory for Structural Bioinformatics, RIKEN Center for Biosystems Dynamics Research, Yokohama, Kanagawa, Japan.

Albert Jeltsch (A)

Department of Biochemistry, Institute of Biochemistry and Technical Biochemistry, University of Stuttgart, Allmandring 31, Stuttgart, Germany.

Yoichi Shinkai (Y)

Cellular Memory Laboratory, RIKEN Cluster for Pioneering Research, Wako, Saitama, Japan. yshinkai@riken.jp.

Takeo Yoshikawa (T)

Laboratory for Molecular Psychiatry, RIKEN Center for Brain Science, Wako, Saitama, Japan. takeo.yoshikawa@riken.jp.

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