GLUT4 translocation and dispersal operate in multiple cell types and are negatively correlated with cell size in adipocytes.


Journal

Scientific reports
ISSN: 2045-2322
Titre abrégé: Sci Rep
Pays: England
ID NLM: 101563288

Informations de publication

Date de publication:
29 11 2022
Historique:
received: 08 08 2022
accepted: 18 11 2022
entrez: 29 11 2022
pubmed: 30 11 2022
medline: 2 12 2022
Statut: epublish

Résumé

The regulated translocation of the glucose transporter, GLUT4, to the surface of adipocytes and muscle is a key action of insulin. This is underpinned by the delivery and fusion of GLUT4-containing vesicles with the plasma membrane. Recent studies have revealed that a further action of insulin is to mediate the dispersal of GLUT4 molecules away from the site of GLUT4 vesicle fusion with the plasma membrane. Although shown in adipocytes, whether insulin-stimulated dispersal occurs in other cells and/or is exhibited by other proteins remains a matter of debate. Here we show that insulin stimulates GLUT4 dispersal in the plasma membrane of adipocytes, induced pluripotent stem cell-derived cardiomyocytes and HeLa cells, suggesting that this phenomenon is specific to GLUT4 expressed in all cell types. By contrast, insulin-stimulated dispersal of TfR was not observed in HeLa cells, suggesting that the mechanism may be unique to GLUT4. Consistent with dispersal being an important physiological mechanism, we observed that insulin-stimulated GLUT4 dispersal is reduced under conditions of insulin resistance. Adipocytes of different sizes have been shown to exhibit distinct metabolic properties: larger adipocytes exhibit reduced insulin-stimulated glucose transport compared to smaller cells. Here we show that both GLUT4 delivery to the plasma membrane and GLUT4 dispersal are reduced in larger adipocytes, supporting the hypothesis that larger adipocytes are refractory to insulin challenge compared to their smaller counterparts, even within a supposedly homogeneous population of cells.

Identifiants

pubmed: 36446811
doi: 10.1038/s41598-022-24736-y
pii: 10.1038/s41598-022-24736-y
pmc: PMC9708847
doi:

Substances chimiques

Insulin 0

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

20535

Subventions

Organisme : British Heart Foundation
Pays : United Kingdom

Informations de copyright

© 2022. The Author(s).

Références

Kahn, B. B. Adipose tissue, inter-organ communication, and the path to type 2 diabetes: The 2016 Banting Medal for scientific achievement lecture. Diabetes 68, 3–14. https://doi.org/10.2337/dbi18-0035 (2019).
doi: 10.2337/dbi18-0035 pubmed: 30573674
Saltiel, A. R. Insulin signaling in health and disease. J. Clin. Invest. 131, e142241. https://doi.org/10.1172/JCI142241 (2021).
doi: 10.1172/JCI142241 pubmed: 33393497 pmcid: 7773347
Gould, G. W., Brodsky, F. M. & Bryant, N. J. Building GLUT4 vesicles: CHC22 Clathrin’s human touch. Trends Cell Biol. 30, 705–719. https://doi.org/10.1016/j.tcb.2020.05.007 (2020).
doi: 10.1016/j.tcb.2020.05.007 pubmed: 32620516
Bryant, N. J., Govers, R. & James, D. E. Regulated transport of the glucose transporter GLUT4. Nat. Rev. Mol. Cell Biol. 3, 267–277. https://doi.org/10.1038/nrm782 (2002).
doi: 10.1038/nrm782 pubmed: 11994746
Bogan, J. S. & Kandror, K. V. Biogenesis and regulation of insulin-responsive vesicles containing GLUT4. Curr. Opin. Cell Biol. 22, 506–512. https://doi.org/10.1016/j.ceb.2010.03.012 (2010).
doi: 10.1016/j.ceb.2010.03.012 pubmed: 20417083 pmcid: 2910140
Klip, A., McGraw, T. E. & James, D. E. Thirty sweet years of GLUT4. J. Biol. Chem. 294, 11369–11381. https://doi.org/10.1074/jbc.REV119.008351 (2019).
doi: 10.1074/jbc.REV119.008351 pubmed: 31175156 pmcid: 6663870
Lizunov, V. A., Stenkula, K., Troy, A., Cushman, S. W. & Zimmerberg, J. Insulin regulates GLUT4 confinement in plasma membrane clusters in adipose cells. PLoS One 8, e57559. https://doi.org/10.1371/journal.pone.0057559 (2013).
doi: 10.1371/journal.pone.0057559 pubmed: 23520472 pmcid: 3592853
Lizunov, V. A. et al. Insulin stimulates fusion, but not tethering, of GLUT4 vesicles in skeletal muscle of HA-GLUT4-GFP transgenic mice. Am. J. Physiol. Endocrinol. Metab. 302, E950-960. https://doi.org/10.1152/ajpendo.00466.2011 (2012).
doi: 10.1152/ajpendo.00466.2011 pubmed: 22297303 pmcid: 3330721
Lizunov, V. A., Lisinski, I., Stenkula, K., Zimmerberg, J. & Cushman, S. W. Insulin regulates fusion of GLUT4 vesicles independent of Exo70-mediated tethering. J. Biol. Chem. 284, 7914–7919. https://doi.org/10.1074/jbc.M806460200 (2009).
doi: 10.1074/jbc.M806460200 pubmed: 19155211 pmcid: 2658084
Lizunov, V. A., Matsumoto, H., Zimmerberg, J., Cushman, S. W. & Frolov, V. A. Insulin stimulates the halting, tethering, and fusion of mobile GLUT4 vesicles in rat adipose cells. J. Cell Biol. 169, 481–489. https://doi.org/10.1083/jcb.200412069 (2005).
doi: 10.1083/jcb.200412069 pubmed: 15866888 pmcid: 2171949
Jiang, L. et al. Direct quantification of fusion rate reveals a distal role for AS160 in insulin-stimulated fusion of GLUT4 storage vesicles. J. Biol. Chem. 283, 8508–8516. https://doi.org/10.1074/jbc.M708688200 (2008).
doi: 10.1074/jbc.M708688200 pubmed: 18063571 pmcid: 2417169
Patki, V. et al. Insulin action on GLUT4 traffic visualized in single 3T3-L1 adipocytes by using ultra-fast microscopy. Mol. Biol. Cell 12, 129–141. https://doi.org/10.1091/mbc.12.1.129 (2001).
doi: 10.1091/mbc.12.1.129 pubmed: 11160828 pmcid: 30573
Suzuki, K. G., Fujiwara, T. K., Edidin, M. & Kusumi, A. Dynamic recruitment of phospholipase C gamma at transiently immobilized GPI-anchored receptor clusters induces IP3-Ca
doi: 10.1083/jcb.200609175 pubmed: 17517965 pmcid: 2064217
Kusumi, A. et al. Defining raft domains in the plasma membrane. Traffic 21, 106–137. https://doi.org/10.1111/tra.12718 (2020).
doi: 10.1111/tra.12718 pubmed: 31760668
Kusumi, A., Tsunoyama, T. A., Hirosawa, K. M., Kasai, R. S. & Fujiwara, T. K. Tracking single molecules at work in living cells. Nat. Chem. Biol. 10, 524–532. https://doi.org/10.1038/nchembio.1558 (2014).
doi: 10.1038/nchembio.1558 pubmed: 24937070
Trimble, W. S. & Grinstein, S. Barriers to the free diffusion of proteins and lipids in the plasma membrane. J. Cell Biol. 208, 259–271. https://doi.org/10.1083/jcb.201410071 (2015).
doi: 10.1083/jcb.201410071 pubmed: 25646084 pmcid: 4315255
Jaqaman, K. et al. Cytoskeletal control of CD36 diffusion promotes its receptor and signaling function. Cell 146, 593–606. https://doi.org/10.1016/j.cell.2011.06.049 (2011).
doi: 10.1016/j.cell.2011.06.049 pubmed: 21854984 pmcid: 3160624
Stenkula, K. G., Lizunov, V. A., Cushman, S. W. & Zimmerberg, J. Insulin controls the spatial distribution of GLUT4 on the cell surface through regulation of its postfusion dispersal. Cell Metab. 12, 250–259. https://doi.org/10.1016/j.cmet.2010.08.005 (2010).
doi: 10.1016/j.cmet.2010.08.005 pubmed: 20816091 pmcid: 3427691
Valentine, J. M. et al. Beta3-Adrenergic receptor downregulation leads to adipocyte catecholamine resistance in obesity. J. Clin. Invest. 132, e153357. https://doi.org/10.1172/JCI153357 (2022).
doi: 10.1172/JCI153357 pubmed: 34847077 pmcid: 8759781
Gao, L., Chen, J., Gao, J., Wang, H. & Xiong, W. Super-resolution microscopy reveals the insulin-resistance-regulated reorganization of Glut4 on plasma membranes. J. Cell Sci. 130, 396–405. https://doi.org/10.1242/jcs.192450 (2017).
doi: 10.1242/jcs.192450 pubmed: 27888215
Koester Anna, M. et al. EFR3 and phosphatidylinositol 4-kinase IIIα regulate insulin-stimulated glucose transport and GLUT4 dispersal in 3T3-L1 adipocytes. Biosci. Rep. 42, bsr20221181. https://doi.org/10.1042/bsr20221181 (2022).
doi: 10.1042/bsr20221181 pubmed: 35735144 pmcid: 9272592
Brewer, P. D., Habtemichael, E. N., Romenskaia, I., Mastick, C. C. & Coster, A. C. Insulin-regulated GLUT4 translocation: Membrane protein trafficking with six distinctive steps. J. Biol. Chem. 289, 17280–17298. https://doi.org/10.1074/jbc.M114.555714 (2014).
doi: 10.1074/jbc.M114.555714 pubmed: 24778187 pmcid: 4067164
Leto, D. & Saltiel, A. R. Regulation of glucose transport by insulin: Traffic control of GLUT4. Nat. Rev. Mol. Cell Biol. 13, 383–396. https://doi.org/10.1038/nrm3351 (2012).
doi: 10.1038/nrm3351 pubmed: 22617471
Lizunov, V. A. et al. Impaired tethering and fusion of GLUT4 vesicles in insulin-resistant human adipose cells. Diabetes 62, 3114–3119. https://doi.org/10.2337/db12-1741 (2013).
doi: 10.2337/db12-1741 pubmed: 23801575 pmcid: 3749349
Santoro, A., McGraw, T. E. & Kahn, B. B. Insulin action in adipocytes, adipose remodeling, and systemic effects. Cell Metab. 33, 748–757. https://doi.org/10.1016/j.cmet.2021.03.019 (2021).
doi: 10.1016/j.cmet.2021.03.019 pubmed: 33826917 pmcid: 8078167
Morris, S. et al. Characterisation of GLUT4 trafficking in HeLa cells: Comparable kinetics and orthologous trafficking mechanisms to 3T3-L1 adipocytes. PeerJ 8, e8751. https://doi.org/10.7717/peerj.8751 (2020).
doi: 10.7717/peerj.8751 pubmed: 32185116 pmcid: 7060922
Roccisana, J., Sadler, J. B., Bryant, N. J. & Gould, G. W. Sorting of GLUT4 into its insulin-sensitive store requires the Sec1/Munc18 protein mVps45. Mol. Biol. Cell 24, 2389–2397. https://doi.org/10.1091/mbc.E13-01-0011 (2013).
doi: 10.1091/mbc.E13-01-0011 pubmed: 23741049 pmcid: 3727931
Bowman, P. R. T., Smith, G. L. & Gould, G. W. GLUT4 expression and glucose transport in human induced pluripotent stem cell-derived cardiomyocytes. PLoS One 14, e0217885. https://doi.org/10.1371/journal.pone.0217885 (2019).
doi: 10.1371/journal.pone.0217885 pubmed: 31344028 pmcid: 6657831
Brewer, P. D., Habtemichael, E. N., Romenskaia, I., Coster, A. C. & Mastick, C. C. Rab14 limits the sorting of GLUT4 from endosomes into insulin-sensitive regulated secretory compartments in adipocytes. Biochem. J. 473, 1315–1327. https://doi.org/10.1042/BCJ20160020 (2016).
doi: 10.1042/BCJ20160020 pubmed: 26936971
Habtemichael, E. N., Brewer, P. D., Romenskaia, I. & Mastick, C. C. Kinetic evidence that GLUT4 follows different endocytic pathways than the receptors for transferrin and ∝2-macroglobulin. J. Biol. Chem. 286, 10115–10125. https://doi.org/10.1074/jbc.M111.217935 (2011).
doi: 10.1074/jbc.M111.217935 pubmed: 21252237 pmcid: 3060463
Kozka, I. J., Clark, A. E. & Holman, G. D. Chronic treatment with insulin selectively down-regulates cell-surface GLUT4 glucose transporters in 3t3-L1 adipocytes. J. Biol. Chem. 266, 11726–11731 (1991).
doi: 10.1016/S0021-9258(18)99017-4 pubmed: 2050674
Maier, V. H. & Gould, G. W. Long-term insulin treatment of 3T3-L1 adipocytes results in mis-targeting of GLUT4: Implications for insulin-stimulated glucose transport. Diabetologia 43, 1273–1281. https://doi.org/10.1007/s001250051523 (2000).
doi: 10.1007/s001250051523 pubmed: 11079746
Koester, A. M. et al. EFR3 and Phosphatidylinositol 4-kinase III Regulate Insulin-stimulated Glucose Transport and GLUT4 Dispersal in 3T3-L1 Adipocytes (Bioscience Reports, In press, Oxford, 2022).
Ovesný, M., Křížek, P., Borkovec, J., Svindrych, Z. & Guy, M. H. ThunderSTORM: A comprehensive ImageJ plug-in for PALM and STORM data analysis and super-resolution imaging. Bioinformatics 30, 2389–2390. https://doi.org/10.1093/bioinformatics/btu202 (2014).
doi: 10.1093/bioinformatics/btu202 pubmed: 24771516 pmcid: 4207427
Campello, R., Moulavi, D. & Sander, J. In Advances in Knowledge Discovery and Data Mining (ed. Pei, J.) 160–172 (Springer, Berlin, 2013).
doi: 10.1007/978-3-642-37456-2_14
Campello, R., Moulavi, D., Zimek, A. & Sander, J. Hierarchical density estimates for data clustering, visualization, and outlier detection. ACM Trans. Knowl. Discov. Data 10, 1–51. https://doi.org/10.1145/2733381 (2015).
doi: 10.1145/2733381
Ester, M., Kriegel, H. P., Sander, J. & Xu, X. In Proceedings of the Second International Conference on Knowledge Discovery and Data Mining (1996).
Levet, F. et al. SR-Tesseler: A method to segment and quantify localization-based super-resolution microscopy data. Nat. Meth. 12, 1065–1071. https://doi.org/10.1038/Nmeth.3579 (2015).
doi: 10.1038/Nmeth.3579
van de Linde, S. Single-molecule localization microscopy analysis with ImageJ. J. Phys. D App. Phys. 52, 203002 (2019).
doi: 10.1088/1361-6463/ab092f
Drawnel, F. M. et al. Disease modeling and phenotypic drug screening for diabetic cardiomyopathy using human induced pluripotent stem cells. Cell Rep. 9, 810–820. https://doi.org/10.1016/j.celrep.2014.09.055 (2014).
doi: 10.1016/j.celrep.2014.09.055 pubmed: 25437537
Fazakerley, D. J., Lawrence, S. P., Lizunov, V. A., Cushman, S. W. & Holman, G. D. A common trafficking route for GLUT4 in cardiomyocytes in response to insulin, contraction and energy-status signalling. J. Cell Sci. 122, 727–734. https://doi.org/10.1242/jcs.041178 (2009).
doi: 10.1242/jcs.041178 pubmed: 19208760 pmcid: 2720923
Bryant, N. J. & Gould, G. W. Insulin stimulated GLUT4 translocation—Size is not everything!. Curr. Opin. Cell Biol. 65, 28–34. https://doi.org/10.1016/j.ceb.2020.02.006 (2020).
doi: 10.1016/j.ceb.2020.02.006 pubmed: 32182545
Haga, Y., Ishii, K. & Suzuki, T. N-Glycosylation is critical for the stability and intracellular trafficking of glucose transporter GLUT4. J. Biol. Chem. 286, 31320–31327. https://doi.org/10.1074/jbc.M111.253955 (2011).
doi: 10.1074/jbc.M111.253955 pubmed: 21757715 pmcid: 3173088
Tanner, L. I. & Lienhard, G. E. Insulin elicits a redistribution of transferrin receptors in 3T3-L1 adipocytes through an increase in the rate constant for receptor externalization. J. Biol. Chem. 262, 8975–8980 (1987).
doi: 10.1016/S0021-9258(18)48032-5 pubmed: 3298247
Sadacca, L. A., Bruno, J., Wen, J., Xiong, W. Y. & McGraw, T. E. Specialized sorting of GLUT4 and its recruitment to the cell surface are independently regulated by distinct Rabs. Mol. Biol. Cell 24, 2544–2557. https://doi.org/10.1091/mbc.E13-02-0103 (2013).
doi: 10.1091/mbc.E13-02-0103 pubmed: 23804653 pmcid: 3744946
Xiong, W. Y., Jordens, I., Gonzalez, E. & McGraw, T. E. GLUT4 is sorted to vesicles whose accumulation beneath and insertion into the plasma membrane are differentially regulated by insulin and selectively affected by insulin resistance. Mol. Biol. Cell 21, 1375–1386. https://doi.org/10.1091/mbc.E09-08-0751 (2010).
doi: 10.1091/mbc.E09-08-0751 pubmed: 20181829 pmcid: 2854095
Blot, V. & McGraw, T. E. Molecular mechanisms controlling GLUT4 intracellular retention. Mol. Biol. Cell 19, 3477–3487. https://doi.org/10.1091/mbc.E08-03-0236 (2008).
doi: 10.1091/mbc.E08-03-0236 pubmed: 18550797 pmcid: 2488284
Livingstone, C., James, D. E., Rice, J. E., Hanpeter, D. & Gould, G. W. Compartment ablation analysis of the insulin-responsive glucose transporter (GLUT4) in 3T3-L1 adipocytes. Biochem. J. 315(Pt 2), 487–495. https://doi.org/10.1042/bj3150487 (1996).
doi: 10.1042/bj3150487 pubmed: 8615819 pmcid: 1217222
Wollman, A. J. M., Kioumourtzoglou, D., Ward, R., Gould, G. W. & Bryant, N. J. Large scale, single-cell FRET-based glucose uptake measurements within heterogeneous populations. iScience. 25, 104023. https://doi.org/10.1016/j.isci.2022.104023 (2022).
doi: 10.1016/j.isci.2022.104023 pubmed: 35313696 pmcid: 8933717
Loo, L. H. et al. Heterogeneity in the physiological states and pharmacological responses of differentiating 3T3-L1 preadipocytes. J. Cell. Biol. 187, 375–384. https://doi.org/10.1083/jcb.200904140 (2009).
doi: 10.1083/jcb.200904140 pubmed: 19948481 pmcid: 2779244
Rizzatti, V. et al. Lipid droplets characterization in adipocyte differentiated 3T3-L1 cells: size and optical density distribution. Eur. J. Histochem. 57, e24. https://doi.org/10.4081/ejh.2013.e24 (2013).
doi: 10.4081/ejh.2013.e24 pubmed: 24085273 pmcid: 3794355
Anthanont, P., Ramos, P., Jensen, M. D. & Hames, K. C. Family history of type 2 diabetes, abdominal adipocyte size and markers of the metabolic syndrome. Int. J. Obes. Lond. 41, 1621–1626. https://doi.org/10.1038/ijo.2017.171 (2017).
doi: 10.1038/ijo.2017.171 pubmed: 28736442 pmcid: 5818259
Bluher, M., Patti, M. E., Gesta, S., Kahn, B. B. & Kahn, C. R. Intrinsic heterogeneity in adipose tissue of fat-specific insulin receptor knock-out mice is associated with differences in patterns of gene expression. J. Biol. Chem. 279, 31891–31901. https://doi.org/10.1074/jbc.M404569200 (2004).
doi: 10.1074/jbc.M404569200 pubmed: 15131119
Stenkula, K. G. & Erlanson-Albertsson, C. Adipose cell size: importance in health and disease. Am. J. Physiol. Regul. Integr. Comp. Physiol. 315, R284–R295. https://doi.org/10.1152/ajpregu.00257.2017 (2018).
doi: 10.1152/ajpregu.00257.2017 pubmed: 29641234
Weyer, C., Foley, J. E., Bogardus, C., Tataranni, P. A. & Pratley, R. E. Enlarged subcutaneous abdominal adipocyte size, but not obesity itself, predicts type II diabetes independent of insulin resistance. Diabetologia 43, 1498–1506. https://doi.org/10.1007/s001250051560 (2000).
doi: 10.1007/s001250051560 pubmed: 11151758
Wree, A. et al. Adipocyte cell size, free fatty acids and apolipoproteins are associated with non-alcoholic liver injury progression in severely obese patients. Metabolism 63, 1542–1552. https://doi.org/10.1016/j.metabol.2014.09.001 (2014).
doi: 10.1016/j.metabol.2014.09.001 pubmed: 25267016
Yang Loureiro, Z., Solivan-Rivera, J. & Corvera, S. Adipocyte heterogeneity underlying adipose tissue functions. Endocrinology 163, BQAB138. https://doi.org/10.1210/endocr/bqab138 (2022).
doi: 10.1210/endocr/bqab138 pubmed: 34223880
Deutsch, A., Feng, D., Pessin, J. E. & Shinoda, K. The impact of single-cell genomics on adipose tissue research. Int. J. Mol. Sci. 21, 4773. https://doi.org/10.3390/ijms21134773 (2020).
doi: 10.3390/ijms21134773 pubmed: 32635651 pmcid: 7369959
Yamada, T., Kamiya, M. & Higuchi, M. Fat depot-specific effects of body fat distribution and adipocyte size on intramuscular fat accumulation in Wagyu cattle. Anim. Sci. J. 91, e13449. https://doi.org/10.1111/asj.13449 (2020).
doi: 10.1111/asj.13449 pubmed: 32815204
Franck, N. et al. Insulin-induced GLUT4 translocation to the plasma membrane is blunted in large compared with small primary fat cells isolated from the same individual. Diabetologia 50, 1716–1722. https://doi.org/10.1007/s00125-007-0713-1 (2007).
doi: 10.1007/s00125-007-0713-1 pubmed: 17572871
Kozka, I. J. et al. The effects of insulin on the level and activity of the GLUT4 present in human adipose-cells. Diabetologia 38, 661–666. https://doi.org/10.1007/s001250050334 (1995).
doi: 10.1007/s001250050334 pubmed: 7672486
Hansson, B. et al. Adipose cell size changes are associated with a drastic actin remodeling. Sci. Rep. 9, 12941. https://doi.org/10.1038/s41598-019-49418-0 (2019).
doi: 10.1038/s41598-019-49418-0 pubmed: 31506540 pmcid: 6736966
Hansson, B. et al. A hypothesis for insulin resistance in primary human adipocytes involving MRTF-A and suppression of PPARγ. Biochem. Biophys. Res. Commun. 533, 64–69. https://doi.org/10.1016/j.bbrc.2020.08.105 (2020).
doi: 10.1016/j.bbrc.2020.08.105 pubmed: 32921413
Hammarstedt, A., Graham, T. E. & Kahn, B. B. Adipose tissue dysregulation and reduced insulin sensitivity in non-obese individuals with enlarged abdominal adipose cells. Diabetol. Metab. Syndr. 4, 42. https://doi.org/10.1186/1758-5996-4-42 (2012).
doi: 10.1186/1758-5996-4-42 pubmed: 22992414 pmcid: 3523053
Lizunov, V. A. et al. Human adipose cells in vitro are either refractory or responsive to insulin, reflecting host metabolic state. PLoS One 10, e0119291. https://doi.org/10.1371/journal.pone.0119291 (2015).
doi: 10.1371/journal.pone.0119291 pubmed: 25768970 pmcid: 4359092

Auteurs

Anna M Koester (AM)

Strathclyde Institute of Pharmacy and Biomedical Sciences, University of Strathclyde, Glasgow, UK.
Institute for Molecular, Cellular and Systems Biology, College of Medical Veterinary and Life Sciences, University of Glasgow, Glasgow, UK.

Angéline Geiser (A)

Strathclyde Institute of Pharmacy and Biomedical Sciences, University of Strathclyde, Glasgow, UK.

Peter R T Bowman (PRT)

Strathclyde Institute of Pharmacy and Biomedical Sciences, University of Strathclyde, Glasgow, UK.

Sebastian van de Linde (S)

Department of Physics, SUPA, University of Strathclyde, 107 Rottenrow East, Glasgow, G4 0NG, Scotland, UK.

Nikolaj Gadegaard (N)

James Watt School of Engineering, University of Glasgow, Glasgow, G12 8QQ, UK.

Nia J Bryant (NJ)

Department of Biology, University of York, York, YO10 5DD, UK. nia.bryant@york.ac.uk.

Gwyn W Gould (GW)

Strathclyde Institute of Pharmacy and Biomedical Sciences, University of Strathclyde, Glasgow, UK. gwyn.gould@strath.ac.uk.
Institute for Molecular, Cellular and Systems Biology, College of Medical Veterinary and Life Sciences, University of Glasgow, Glasgow, UK. gwyn.gould@strath.ac.uk.

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