Discovery and systematic characterization of risk variants and genes for coronary artery disease in over a million participants.


Journal

Nature genetics
ISSN: 1546-1718
Titre abrégé: Nat Genet
Pays: United States
ID NLM: 9216904

Informations de publication

Date de publication:
12 2022
Historique:
received: 26 04 2021
accepted: 17 10 2022
pubmed: 7 12 2022
medline: 15 12 2022
entrez: 6 12 2022
Statut: ppublish

Résumé

The discovery of genetic loci associated with complex diseases has outpaced the elucidation of mechanisms of disease pathogenesis. Here we conducted a genome-wide association study (GWAS) for coronary artery disease (CAD) comprising 181,522 cases among 1,165,690 participants of predominantly European ancestry. We detected 241 associations, including 30 new loci. Cross-ancestry meta-analysis with a Japanese GWAS yielded 38 additional new loci. We prioritized likely causal variants using functionally informed fine-mapping, yielding 42 associations with less than five variants in the 95% credible set. Similarity-based clustering suggested roles for early developmental processes, cell cycle signaling and vascular cell migration and proliferation in the pathogenesis of CAD. We prioritized 220 candidate causal genes, combining eight complementary approaches, including 123 supported by three or more approaches. Using CRISPR-Cas9, we experimentally validated the effect of an enhancer in MYO9B, which appears to mediate CAD risk by regulating vascular cell motility. Our analysis identifies and systematically characterizes >250 risk loci for CAD to inform experimental interrogation of putative causal mechanisms for CAD.

Identifiants

pubmed: 36474045
doi: 10.1038/s41588-022-01233-6
pii: 10.1038/s41588-022-01233-6
pmc: PMC9729111
doi:

Types de publication

Meta-Analysis Journal Article Research Support, Non-U.S. Gov't Research Support, U.S. Gov't, Non-P.H.S. Research Support, N.I.H., Extramural

Langues

eng

Sous-ensembles de citation

IM

Pagination

1803-1815

Subventions

Organisme : Department of Health
ID : BRC-1215-20014
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : R01 HL086694
Pays : United States
Organisme : NIDDK NIH HHS
ID : UM1 DK105554
Pays : United States
Organisme : NHLBI NIH HHS
ID : HHSN268201700001I
Pays : United States
Organisme : British Heart Foundation
ID : RG/14/5/30893
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : HHSN268201700002I
Pays : United States
Organisme : NHLBI NIH HHS
ID : HHSN268201700005I
Pays : United States
Organisme : NHLBI NIH HHS
ID : K08 HL153950
Pays : United States
Organisme : NHLBI NIH HHS
ID : R01 HL087641
Pays : United States
Organisme : British Heart Foundation
ID : RG/13/13/30194
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_QA137853
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : R35 HL135824
Pays : United States
Organisme : NHLBI NIH HHS
ID : R01 HL125863
Pays : United States
Organisme : Medical Research Council
ID : MR/L003120/1
Pays : United Kingdom
Organisme : Medical Research Council
ID : MC_PC_17228
Pays : United Kingdom
Organisme : Medical Research Council
ID : MR/S502443/1
Pays : United Kingdom
Organisme : NHGRI NIH HHS
ID : K08 HG010155
Pays : United States
Organisme : NHLBI NIH HHS
ID : K08 HL153937
Pays : United States
Organisme : Wellcome Trust
ID : 203141/Z/16/Z
Pays : United Kingdom
Organisme : Department of Health
Pays : United Kingdom
Organisme : British Heart Foundation
ID : FS/14/66/3129
Pays : United Kingdom
Organisme : British Heart Foundation
ID : RG/18/13/33946
Pays : United Kingdom
Organisme : NHGRI NIH HHS
ID : T32 HG000040
Pays : United States
Organisme : NHLBI NIH HHS
ID : HHSN268201700004I
Pays : United States
Organisme : NCRR NIH HHS
ID : UL1 RR025005
Pays : United States
Organisme : British Heart Foundation
ID : SP/13/2/30111
Pays : United Kingdom
Organisme : British Heart Foundation
ID : SP/16/4/32697
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : T32 HL007604
Pays : United States
Organisme : NHLBI NIH HHS
ID : R01 HL146860
Pays : United States
Organisme : NHLBI NIH HHS
ID : R01 HL059367
Pays : United States
Organisme : NHGRI NIH HHS
ID : U01 HG004402
Pays : United States
Organisme : British Heart Foundation
ID : SP/19/2/34462
Pays : United Kingdom
Organisme : British Heart Foundation
ID : RE/13/1/30181
Pays : United Kingdom
Organisme : British Heart Foundation
ID : FS/14/55/30806
Pays : United Kingdom
Organisme : NHLBI NIH HHS
ID : HHSN268201700003I
Pays : United States
Organisme : British Heart Foundation
ID : SP/09/002
Pays : United Kingdom
Organisme : Medical Research Council
ID : G0800270
Pays : United Kingdom

Investigateurs

John Danesh (J)
Paul S de Vries (PS)
Moritz von Scheidt (M)

Commentaires et corrections

Type : CommentIn

Informations de copyright

© 2022. The Author(s).

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Auteurs

Krishna G Aragam (KG)

Cardiovascular Research Center, Massachusetts General Hospital, Boston, MA, USA. karagam@broadinstitute.org.
Center for Genomic Medicine, Massachusetts General Hospital, Boston, MA, USA. karagam@broadinstitute.org.
Cardiovascular Disease Initiative, Broad Institute of MIT and Harvard, Cambridge, MA, USA. karagam@broadinstitute.org.
Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA. karagam@broadinstitute.org.

Tao Jiang (T)

BHF Cardiovascular Epidemiology Unit, Department of Public Health and Primary Care, University of Cambridge, Cambridge, UK.

Anuj Goel (A)

Radcliffe Department of Medicine, Division of Cardiovascular Medicine, University of Oxford, Oxford, UK.
Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.

Stavroula Kanoni (S)

William Harvey Research Institute, Barts and the London School of Medicine and Dentistry, Queen Mary University of London, London, UK.

Brooke N Wolford (BN)

Department of Computational Medicine and Bioinformatics, University of Michigan, Ann Arbor, MI, USA.

Deepak S Atri (DS)

Cardiovascular Disease Initiative, Broad Institute of MIT and Harvard, Cambridge, MA, USA.
Divisions of Cardiovascular Medicine and Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, MA, USA.

Elle M Weeks (EM)

Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Minxian Wang (M)

Cardiovascular Disease Initiative, Broad Institute of MIT and Harvard, Cambridge, MA, USA.
Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

George Hindy (G)

Department of Population Medicine, Qatar University College of Medicine, Doha, Qatar.

Wei Zhou (W)

Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.
Department of Computational Medicine and Bioinformatics, University of Michigan, Ann Arbor, MI, USA.
Analytic and Translational Genetics Unit, Massachusetts General Hospital, Boston, MA, USA.
Stanley Center for Psychiatric Research, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Christopher Grace (C)

Radcliffe Department of Medicine, Division of Cardiovascular Medicine, University of Oxford, Oxford, UK.
Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.

Carolina Roselli (C)

Cardiovascular Disease Initiative, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Nicholas A Marston (NA)

TIMI Study Group, Division of Cardiovascular Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, MA, USA.

Frederick K Kamanu (FK)

TIMI Study Group, Division of Cardiovascular Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, MA, USA.

Ida Surakka (I)

Department of Internal Medicine, Division of Cardiology, University of Michigan, Ann Arbor, MI, USA.

Loreto Muñoz Venegas (LM)

Institute for Cardiogenetics, University of Lübeck, Lübeck, Germany.
German Research Center for Cardiovascular Research (DZHK), Partner Site Hamburg/Lübeck/Kiel, Lübeck, Germany.

Paul Sherliker (P)

Medical Research Council Population Health Research Unit, CTSU-Nuffield Department of Population Health, Medical Sciences Division, University of Oxford, Oxford, UK.

Satoshi Koyama (S)

Laboratory for Cardiovascular Genomics and Informatics, RIKEN Center for Integrative Medical Sciences, Tsurumi-ku, Yokohama, Japan.

Kazuyoshi Ishigaki (K)

Laboratory for Statistical and Translational Genetics, RIKEN Center for Integrative Medical Sciences, Tsurumi-ku, Yokohama, Japan.

Bjørn O Åsvold (BO)

Department of Public Health and Nursing, K.G. Jebsen Center for Genetic Epidemiology, Norwegian University of Science and Technology, NTNU, Trondheim, Norway.
HUNT Research Centre, Norwegian University of Science and Technology, Levanger, Norway.
Department of Endocrinology, Clinic of Medicine, St. Olavs Hospital, Trondheim, Norway.

Michael R Brown (MR)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, The University of Texas Health Science Center at Houston, Houston, TX, USA.

Ben Brumpton (B)

Department of Public Health and Nursing, K.G. Jebsen Center for Genetic Epidemiology, Norwegian University of Science and Technology, NTNU, Trondheim, Norway.
HUNT Research Centre, Norwegian University of Science and Technology, Levanger, Norway.

Paul S de Vries (PS)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, The University of Texas Health Science Center at Houston, Houston, TX, USA.

Olga Giannakopoulou (O)

William Harvey Research Institute, Barts and the London School of Medicine and Dentistry, Queen Mary University of London, London, UK.

Panagiota Giardoglou (P)

Department of Nutrition-Dietetics, School of Health Science and Education, Harokopio University, Athens, Greece.

Daniel F Gudbjartsson (DF)

deCODE Genetics/Amgen, Inc., Reykjavik, Iceland.
School of Engineering and Natural Sciences, University of Iceland, Reykjavik, Iceland.

Ulrich Güldener (U)

German Heart Centre Munich, Department of Cardiology, Technical University of Munich, Munich, Germany.

Syed M Ijlal Haider (SMI)

Institute for Cardiogenetics, University of Lübeck, Lübeck, Germany.
German Research Center for Cardiovascular Research (DZHK), Partner Site Hamburg/Lübeck/Kiel, Lübeck, Germany.

Anna Helgadottir (A)

deCODE Genetics/Amgen, Inc., Reykjavik, Iceland.

Maysson Ibrahim (M)

CTSU-Nuffield Department of Population Health, Medical Sciences Division, University of Oxford, Oxford, UK.

Adnan Kastrati (A)

German Heart Centre Munich, Department of Cardiology, Technical University of Munich, Munich, Germany.
German Research Center for Cardiovascular Research (DZHK e.V.), Partner Site Munich Heart Alliance, Munich, Germany.

Thorsten Kessler (T)

German Heart Centre Munich, Department of Cardiology, Technical University of Munich, Munich, Germany.
German Research Center for Cardiovascular Research (DZHK e.V.), Partner Site Munich Heart Alliance, Munich, Germany.

Theodosios Kyriakou (T)

Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.

Tomasz Konopka (T)

William Harvey Research Institute, Barts and the London School of Medicine and Dentistry, Queen Mary University of London, London, UK.

Ling Li (L)

German Heart Centre Munich, Department of Cardiology, Technical University of Munich, Munich, Germany.

Lijiang Ma (L)

Department of Genetics and Genomic Science, Icahn Institute for Genomics and Multiscale Biology, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
The Zena and Michael A. Wiener Cardiovascular Institute, Icahn School of Medicine at Mount Sinai, New York, NY, USA.

Thomas Meitinger (T)

German Research Center for Cardiovascular Research (DZHK e.V.), Partner Site Munich Heart Alliance, Munich, Germany.
Institute of Human Genetics, Helmholtz Zentrum München, German Research Center for Environmental Health, Neuherberg, Germany.
Klinikum Rechts der Isar, Institute of Human Genetics, Technical University of Munich, Munich, Germany.

Sören Mucha (S)

Institute for Cardiogenetics, University of Lübeck, Lübeck, Germany.
German Research Center for Cardiovascular Research (DZHK), Partner Site Hamburg/Lübeck/Kiel, Lübeck, Germany.

Matthias Munz (M)

Institute for Cardiogenetics, University of Lübeck, Lübeck, Germany.
German Research Center for Cardiovascular Research (DZHK), Partner Site Hamburg/Lübeck/Kiel, Lübeck, Germany.

Federico Murgia (F)

CTSU-Nuffield Department of Population Health, Medical Sciences Division, University of Oxford, Oxford, UK.

Jonas B Nielsen (JB)

Department of Internal Medicine, Division of Cardiology, University of Michigan, Ann Arbor, MI, USA.
Department of Public Health and Nursing, K.G. Jebsen Center for Genetic Epidemiology, Norwegian University of Science and Technology, NTNU, Trondheim, Norway.

Markus M Nöthen (MM)

School of Medicine and University Hospital Bonn, Institute of Human Genetics, University of Bonn, Bonn, Germany.

Shichao Pang (S)

German Heart Centre Munich, Department of Cardiology, Technical University of Munich, Munich, Germany.

Tobias Reinberger (T)

Institute for Cardiogenetics, University of Lübeck, Lübeck, Germany.
German Research Center for Cardiovascular Research (DZHK), Partner Site Hamburg/Lübeck/Kiel, Lübeck, Germany.

Gavin Schnitzler (G)

Cardiovascular Disease Initiative, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Damian Smedley (D)

William Harvey Research Institute, Barts and the London School of Medicine and Dentistry, Queen Mary University of London, London, UK.

Gudmar Thorleifsson (G)

deCODE Genetics/Amgen, Inc., Reykjavik, Iceland.

Moritz von Scheidt (M)

German Heart Centre Munich, Department of Cardiology, Technical University of Munich, Munich, Germany.
German Research Center for Cardiovascular Research (DZHK e.V.), Partner Site Munich Heart Alliance, Munich, Germany.

Jacob C Ulirsch (JC)

Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.
Analytic and Translational Genetics Unit, Massachusetts General Hospital, Boston, MA, USA.
Program in Biological and Biomedical Sciences, Harvard Medical School, Boston, MA, USA.

David O Arnar (DO)

deCODE Genetics/Amgen, Inc., Reykjavik, Iceland.
Faculty of Medicine, University of Iceland, Reykjavik, Iceland.
Department of Internal Medicine, Division of Cardiology, Landspitali-National University Hospital of Iceland, Hringbraut, Reykjavik, Iceland.

Noël P Burtt (NP)

Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Maria C Costanzo (MC)

Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Jason Flannick (J)

Division of Genetics and Genomics, Boston Children's Hospital, Boston, MA, USA.

Kaoru Ito (K)

Laboratory for Cardiovascular Genomics and Informatics, RIKEN Center for Integrative Medical Sciences, Tsurumi-ku, Yokohama, Japan.

Dong-Keun Jang (DK)

Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Yoichiro Kamatani (Y)

Department of Computational Biology and Medical Sciences, Graduate School of Frontier Sciences, The University of Tokyo, Tokyo, Japan.

Amit V Khera (AV)

Center for Genomic Medicine, Massachusetts General Hospital, Boston, MA, USA.
Cardiovascular Disease Initiative, Broad Institute of MIT and Harvard, Cambridge, MA, USA.
Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Issei Komuro (I)

Department of Cardiovascular Medicine, The University of Tokyo, Tokyo, Japan.

Iftikhar J Kullo (IJ)

Department of Cardiovascular Medicine, Mayo Clinic, Rochester, MN, USA.

Luca A Lotta (LA)

Regeneron Genetics Center, Regeneron Pharmaceuticals, Tarrytown, NY, USA.

Christopher P Nelson (CP)

Department of Cardiovascular Sciences and NIHR Leicester Biomedical Research Centre, University of Leicester, Glenfield Hospital, Leicester, UK.

Robert Roberts (R)

Cardiovascular Genomics and Genetics, University of Arizona College of Medicin, Phoenix, AZ, USA.

Gudmundur Thorgeirsson (G)

deCODE Genetics/Amgen, Inc., Reykjavik, Iceland.
Faculty of Medicine, University of Iceland, Reykjavik, Iceland.
Department of Internal Medicine, Division of Cardiology, Landspitali-National University Hospital of Iceland, Hringbraut, Reykjavik, Iceland.

Unnur Thorsteinsdottir (U)

deCODE Genetics/Amgen, Inc., Reykjavik, Iceland.
Faculty of Medicine, University of Iceland, Reykjavik, Iceland.

Thomas R Webb (TR)

Department of Cardiovascular Sciences and NIHR Leicester Biomedical Research Centre, University of Leicester, Glenfield Hospital, Leicester, UK.

Aris Baras (A)

Regeneron Genetics Center, Regeneron Pharmaceuticals, Tarrytown, NY, USA.

Johan L M Björkegren (JLM)

Department of Genetics and Genomic Sciences, Institute of Genomics and Multiscale Biology, Icahn School of Medicine at Mount Sinai, New York, NY, USA.
Integrated Cardio Metabolic Centre, Karolinska Institutet, Karolinska Universitetssjukhuset, Huddinge, Sweden.
Clinical Gene Networks AB, Stockholm, Sweden.

Eric Boerwinkle (E)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, The University of Texas Health Science Center at Houston, Houston, TX, USA.
Human Genome Sequencing Center, Baylor College of Medicine, Houston, TX, USA.

George Dedoussis (G)

Department of Nutrition-Dietetics, School of Health Science and Education, Harokopio University, Athens, Greece.

Hilma Holm (H)

deCODE Genetics/Amgen, Inc., Reykjavik, Iceland.

Kristian Hveem (K)

Department of Public Health and Nursing, K.G. Jebsen Center for Genetic Epidemiology, Norwegian University of Science and Technology, NTNU, Trondheim, Norway.
HUNT Research Centre, Norwegian University of Science and Technology, Levanger, Norway.

Olle Melander (O)

Department of Clinical Sciences in Malmö, Lund University, Malmö, Sweden.

Alanna C Morrison (AC)

Human Genetics Center, Department of Epidemiology, Human Genetics, and Environmental Sciences, School of Public Health, The University of Texas Health Science Center at Houston, Houston, TX, USA.

Marju Orho-Melander (M)

Department of Clinical Sciences in Malmö, Lund University, Malmö, Sweden.

Loukianos S Rallidis (LS)

Second Department of Cardiology, Medical School, National and Kapodistrian University of Athens, University General Hospital Attikon, Athens, Greece.

Arno Ruusalepp (A)

Department of Cardiac Surgery, Tartu University Hospital and Institute of Clinical Medicine, Tartu University, Tartu, Estonia.

Marc S Sabatine (MS)

TIMI Study Group, Division of Cardiovascular Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, MA, USA.

Kari Stefansson (K)

deCODE Genetics/Amgen, Inc., Reykjavik, Iceland.
Faculty of Medicine, University of Iceland, Reykjavik, Iceland.

Pierre Zalloua (P)

Harvard T.H.Chan School of Public Health, Boston, MA, USA.
College of Medicine and Health Sciences, Khalifa University, Abu Dhabi, UAE.

Patrick T Ellinor (PT)

Cardiovascular Research Center, Massachusetts General Hospital, Boston, MA, USA.
Cardiovascular Disease Initiative, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Martin Farrall (M)

Radcliffe Department of Medicine, Division of Cardiovascular Medicine, University of Oxford, Oxford, UK.
Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.

John Danesh (J)

BHF Cardiovascular Epidemiology Unit, Department of Public Health and Primary Care, University of Cambridge, Cambridge, UK.
National Institute for Health and Care Research Cambridge Biomedical Research Centre, Cambridge University Hospitals, Cambridge, UK.
The National Institute for Health and Care Research Blood and Transplant Unit (NIHR BTRU) in Donor Health and Genomics, University of Cambridge, Cambridge, UK.
Human Genetics, Wellcome Sanger Institute, Saffron Walden, UK.
Health Data Research UK Cambridge, Wellcome Genome Campus and University of Cambridge, Cambridge, UK.
British Heart Foundation Centre of Research Excellence, Division of Cardiovascular Medicine, Addenbrooke's Hospital, Cambridge, UK.

Christian T Ruff (CT)

TIMI Study Group, Division of Cardiovascular Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, MA, USA.

Hilary K Finucane (HK)

Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.
Analytic and Translational Genetics Unit, Massachusetts General Hospital, Boston, MA, USA.
Stanley Center for Psychiatric Research, Broad Institute of MIT and Harvard, Cambridge, MA, USA.

Jemma C Hopewell (JC)

CTSU-Nuffield Department of Population Health, Medical Sciences Division, University of Oxford, Oxford, UK.

Robert Clarke (R)

CTSU-Nuffield Department of Population Health, Medical Sciences Division, University of Oxford, Oxford, UK.

Rajat M Gupta (RM)

Cardiovascular Disease Initiative, Broad Institute of MIT and Harvard, Cambridge, MA, USA.
Program in Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, MA, USA.
Divisions of Cardiovascular Medicine and Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, MA, USA.

Jeanette Erdmann (J)

Institute for Cardiogenetics, University of Lübeck, Lübeck, Germany.
German Research Center for Cardiovascular Research (DZHK), Partner Site Hamburg/Lübeck/Kiel, Lübeck, Germany.

Nilesh J Samani (NJ)

Faculty of Medicine, University of Iceland, Reykjavik, Iceland.

Heribert Schunkert (H)

German Heart Centre Munich, Department of Cardiology, Technical University of Munich, Munich, Germany.
German Research Center for Cardiovascular Research (DZHK e.V.), Partner Site Munich Heart Alliance, Munich, Germany.

Hugh Watkins (H)

Radcliffe Department of Medicine, Division of Cardiovascular Medicine, University of Oxford, Oxford, UK.
Wellcome Centre for Human Genetics, University of Oxford, Oxford, UK.

Cristen J Willer (CJ)

Department of Computational Medicine and Bioinformatics, University of Michigan, Ann Arbor, MI, USA.
Department of Internal Medicine, Division of Cardiology, University of Michigan, Ann Arbor, MI, USA.
Department of Human Genetics, University of Michigan, Ann Arbor, MI, USA.

Panos Deloukas (P)

William Harvey Research Institute, Barts and the London School of Medicine and Dentistry, Queen Mary University of London, London, UK.
Princess Al-Jawhara Al-Brahim Centre of Excellence in Research of Hereditary Disorders (PACER-HD), King Abdulaziz University, Jeddah, Saudi Arabia.

Sekar Kathiresan (S)

Verve Therapeutics, Cambridge, MA, USA.

Adam S Butterworth (AS)

BHF Cardiovascular Epidemiology Unit, Department of Public Health and Primary Care, University of Cambridge, Cambridge, UK. asb38@medschl.cam.ac.uk.
National Institute for Health and Care Research Cambridge Biomedical Research Centre, Cambridge University Hospitals, Cambridge, UK. asb38@medschl.cam.ac.uk.
The National Institute for Health and Care Research Blood and Transplant Unit (NIHR BTRU) in Donor Health and Genomics, University of Cambridge, Cambridge, UK. asb38@medschl.cam.ac.uk.
Health Data Research UK Cambridge, Wellcome Genome Campus and University of Cambridge, Cambridge, UK. asb38@medschl.cam.ac.uk.
British Heart Foundation Centre of Research Excellence, Division of Cardiovascular Medicine, Addenbrooke's Hospital, Cambridge, UK. asb38@medschl.cam.ac.uk.

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