CRISPR-Cas provides limited phage immunity to a prevalent gut bacterium in gnotobiotic mice.


Journal

The ISME journal
ISSN: 1751-7370
Titre abrégé: ISME J
Pays: England
ID NLM: 101301086

Informations de publication

Date de publication:
03 2023
Historique:
received: 20 05 2022
accepted: 06 01 2023
revised: 22 12 2022
pmc-release: 01 03 2024
pubmed: 12 1 2023
medline: 25 2 2023
entrez: 11 1 2023
Statut: ppublish

Résumé

Many bacteria and archaea harbor the adaptive CRISPR-Cas system, which stores small nucleotide fragments from previous invasions of nucleic acids via viruses or plasmids. This molecular archive blocks further invaders carrying identical or similar nucleotide sequences. However, few of these systems have been confirmed experimentally to be active in gut bacteria. Here, we demonstrate experimentally that the type I-C CRISPR-Cas system of the prevalent gut bacterium Eggerthella lenta can specifically target and cleave foreign DNA in vitro by using a plasmid transformation assay. We also show that the CRISPR-Cas system acquires new immunities (spacers) from the genome of a virulent E. lenta phage using traditional phage assays in vitro but also in vivo using gnotobiotic (GB) mice. Both high phage titer and an increased number of spacer acquisition events were observed when E. lenta was exposed to a low multiplicity of infection in vitro, and three phage genes were found to contain protospacer hotspots. Fewer new spacer acquisitions were detected in vivo than in vitro. Longitudinal analysis of phage-bacteria interactions showed sustained coexistence in the gut of GB mice, with phage abundance being approximately one log higher than the bacteria. Our findings show that while the type I-C CRISPR-Cas system is active in vitro and in vivo, a highly virulent phage in vitro was still able to co-exist with its bacterial host in vivo. Taken altogether, our results suggest that the CRISPR-Cas defense system of E. lenta provides only partial immunity in the gut.

Identifiants

pubmed: 36631688
doi: 10.1038/s41396-023-01358-4
pii: 10.1038/s41396-023-01358-4
pmc: PMC9938214
doi:

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

432-442

Subventions

Organisme : CIHR
Pays : Canada

Informations de copyright

© 2023. The Author(s), under exclusive licence to International Society for Microbial Ecology.

Références

McGinn J, Marraffini LA. Molecular mechanisms of CRISPR-Cas spacer acquisition. Nat Rev Microbiol. 2019;17:7–12.
pubmed: 30171202 doi: 10.1038/s41579-018-0071-7
Brouns SJJ, Jore MM, Lundgren M, Westra ER, Slijkhuis RJH, Snijders APL, et al. Small CRISPR RNAs guide antiviral defense in prokaryotes. Science. 2008;321:960–4.
pubmed: 18703739 pmcid: 5898235 doi: 10.1126/science.1159689
Barrangou R, Fremaux C, Deveau H, Richards M, Boyaval P, Moineau S, et al. CRISPR provides acquired resistance against viruses in prokaryotes. Science. 2007;315:1709–12.
pubmed: 17379808 doi: 10.1126/science.1138140
Ishino Y, Shinagawa H, Makino K, Amemura M, Nakatura A. Nucleotide sequence of the iap gene, responsible for alkaline phosphatase isoenzyme conversion in Escherichia coli, and identification of the gene product. J Bacteriol. 1987;169:5429–33.
pubmed: 3316184 pmcid: 213968 doi: 10.1128/jb.169.12.5429-5433.1987
Bolotin A, Quinquis B, Sorokin A, Ehrlich SD. Clustered regularly interspaced short palindrome repeats (CRISPRs) have spacers of extrachromosomal origin. Microbiol (Read). 2005;151:2551–61.
doi: 10.1099/mic.0.28048-0
Makarova KS, Wolf YI, Iranzo J, Shmakov SA, Alkhnbashi OS, Brouns SJJ, et al. Evolutionary classification of CRISPR–Cas systems: a burst of class 2 and derived variants. Nat Rev Microbiol. 2019;18:67–83.
pubmed: 31857715 pmcid: 8905525 doi: 10.1038/s41579-019-0299-x
Makarova KS, Wolf YI, Alkhnbashi OS, Costa F, Shah SA, Saunders SJ, et al. An updated evolutionary classification of CRISPR-Cas systems. Nat Rev Microbiol. 2015;13:722–36.
pubmed: 26411297 pmcid: 5426118 doi: 10.1038/nrmicro3569
Hochstrasser ML, Taylor DW, Kornfeld JE, Nogales E, Doudna JA. DNA targeting by a minimal CRISPR RNA-guided cascade. Mol Cell. 2016;63:840–51.
pubmed: 27588603 pmcid: 5111854 doi: 10.1016/j.molcel.2016.07.027
Lee H, Zhou Y, Taylor DW, Sashital DG. Cas4-dependent prespacer processing ensures high-fidelity programming of CRISPR arrays. Mol Cell. 2018;70:48–59.e5.
pubmed: 29602742 pmcid: 5889325 doi: 10.1016/j.molcel.2018.03.003
Lee H, Dhingra Y, Sashital DG. The Cas4-Cas1-Cas2 complex mediates precise prespacer processing during CRISPR adaptation. eLife. 2019;8:1–26.
doi: 10.7554/eLife.44248
Rasmussen TS, Koefoed AK, Jakobsen RR, Deng L, Castro-Mejía JL, Brunse A, et al. Bacteriophage-mediated manipulation of the gut microbiome - promises and presents limitations. FEMS Microbiol Rev. 2020;44:507–21.
pubmed: 32495834 doi: 10.1093/femsre/fuaa020
Bernheim A, Sorek R. The pan-immune system of bacteria: antiviral defence as a community resource. Nat Rev Microbiol. 2020;18:113–9.
pubmed: 31695182 doi: 10.1038/s41579-019-0278-2
Louwen R, Staals RHJ, Endtz HP, van Baarlen P, van der Oost J. The role of CRISPR-Cas systems in virulence of pathogenic bacteria. Microbiol Mol Biol Rev. 2014;78:74–88.
pubmed: 24600041 pmcid: 3957734 doi: 10.1128/MMBR.00039-13
Grissa I, Vergnaud G, Pourcel C. CRISPRcompar: a website to compare clustered regularly interspaced short palindromic repeats. Nucleic Acids Res. 2008;36:W145–8.
pubmed: 18442988 pmcid: 2447796 doi: 10.1093/nar/gkn228
Tajkarimi M, Wexler HM. CRISPR-Cas systems in Bacteroides fragilis, an important pathobiont in the human gut microbiome. Front Microbiol. 2017;8:2234.
pubmed: 29218031 pmcid: 5704556 doi: 10.3389/fmicb.2017.02234
Soto-Perez P, Bisanz JE, Berry JD, Lam KN, Bondy-Denomy J, Turnbaugh PJ. CRISPR-Cas system of a prevalent human gut bacterium reveals hyper-targeting against phages in a human virome catalog. Cell Host Microbe. 2019;26:325–35.e5.
pubmed: 31492655 pmcid: 6936622 doi: 10.1016/j.chom.2019.08.008
Cornuault JK, Moncaut E, Loux V, Mathieu A, Sokol H, Petit M-A, et al. The enemy from within: a prophage of Roseburia intestinalis systematically turns lytic in the mouse gut, driving bacterial adaptation by CRISPR spacer acquisition. ISME J. 2020;14:771–87.
pubmed: 31827247 doi: 10.1038/s41396-019-0566-x
Pan M, Nethery MA, Hidalgo-Cantabrana C, Barrangou R. Comprehensive mining and characterization of CRISPR-Cas systems in. Bifidobacterium Microorg. 2020;8:720.
doi: 10.3390/microorganisms8050720
Koh A, Molinaro A, Ståhlman M, Khan MT, Schmidt C, Mannerås-Holm L, et al. Microbially produced imidazole propionate impairs insulin signaling through mTORC1. Cell. 2018;175:947–61.e17.
pubmed: 30401435 doi: 10.1016/j.cell.2018.09.055
Qin J, Li Y, Cai Z, Li S, Zhu J, Zhang F, et al. A metagenome-wide association study of gut microbiota in type 2 diabetes. Nature. 2012;490:55–60.
pubmed: 23023125 doi: 10.1038/nature11450
Sprotte S, Rasmussen TS, Cho G, Brinks E, Lametsch R, Neve H, et al. Morphological and genetic characterization of Eggerthella lenta bacteriophage PMBT5. Viruses. 2022;14:1–16.
doi: 10.3390/v14081598
Rasmussen TS, Streidl T, Hitch TCAA, Wortmann E, Deptula P, Kofoed MVW, et al. Sporofaciens musculi gen. nov., sp. nov., a novel bacterium isolated from the caecum of an obese mouse. Int J Syst Evol Microbiol. 2021;71:004673.
Muhammed MK, Krych L, Nielsen DS, Vogensen FK. A high-throughput qPCR system for simultaneous quantitative detection of dairy Lactococcus lactis and Leuconostoc bacteriophages. PLoS One. 2017;12:e0174223.
pubmed: 28339484 pmcid: 5365131 doi: 10.1371/journal.pone.0174223
De Vos W. Gene cloning and expression in lactic streptococci. FEMS Microbiol Lett. 1987;46:281–95.
doi: 10.1111/j.1574-6968.1987.tb02466.x
Rao C, Chin D, Ensminger AW. Priming in a permissive type I-C CRISPR-Cas system reveals distinct dynamics of spacer acquisition and loss. RNA. 2017;23:1525–38.
pubmed: 28724535 pmcid: 5602111 doi: 10.1261/rna.062083.117
Burmistrz M, Rodriguez Martinez JI, Krochmal D, Staniec D, Pyrc K. Clustered regularly interspaced short palindromic repeat (CRISPR) RNAs in the Porphyromonas gingivalis CRISPR-Cas I-C system. J Bacteriol. 2017;199:e00275–17.
pubmed: 28893837 pmcid: 5686588 doi: 10.1128/JB.00275-17
Lemay M-L, Tremblay DM, Moineau S. Genome engineering of virulent lactococcal phages using CRISPR-Cas9. ACS Synth Biol. 2017;6:1351–8.
pubmed: 28324650 doi: 10.1021/acssynbio.6b00388
McKenzie RE, Almendros C, Vink JNA, Brouns SJJ. Using CAPTURE to detect spacer acquisition in native CRISPR arrays. Nat Protoc. 2019;14:976–90.
pubmed: 30742049 pmcid: 6831484 doi: 10.1038/s41596-018-0123-5
Rasmussen TS, de Vries L, Kot W, Hansen LH, Castro-Mejía JL, Vogensen FK, et al. Mouse vendor influence on the bacterial and viral gut composition exceeds the effect of diet. Viruses. 2019;11:435.
pubmed: 31086117 pmcid: 6563299 doi: 10.3390/v11050435
Krych Ł, Kot W, Bendtsen KMB, Hansen AK, Vogensen FK, Nielsen DS. Have you tried spermine? A rapid and cost-effective method to eliminate dextran sodium sulfate inhibition of PCR and RT-PCR. J Microbiol Methods. 2018;144:1–7.
pubmed: 29107603 doi: 10.1016/j.mimet.2017.10.015
Hui Y, Tamez-Hidalgo P, Cieplak T, Satessa GD, Kot W, Kjærulff S, et al. Supplementation of a lacto-fermented rapeseed-seaweed blend promotes gut microbial- and gut immune-modulation in weaner piglets. J Anim Sci Biotechnol. 2021;12:1–14.
doi: 10.1186/s40104-021-00601-2
Edgar RC. Search and clustering orders of magnitude faster than BLAST. Bioinformatics. 2010;26:2460–1.
pubmed: 20709691 doi: 10.1093/bioinformatics/btq461
Martin M. Cutadapt removes adapter sequences from high-throughput sequencing reads. EMBnet J. 2011;17:10.
doi: 10.14806/ej.17.1.200
Li H, Durbin R. Fast and accurate short read alignment with Burrows-Wheeler transform. Bioinformatics. 2009;25:1754–60.
pubmed: 19451168 pmcid: 2705234 doi: 10.1093/bioinformatics/btp324
Milne I, Stephen G, Bayer M, Cock PJAA, Pritchard L, Cardle L, et al. Using Tablet for visual exploration of second-generation sequencing data. Brief Bioinform. 2013;14:193–202.
pubmed: 22445902 doi: 10.1093/bib/bbs012
Johnson M, Zaretskaya I, Raytselis Y, Merezhuk Y, McGinnis S, Madden TL. NCBI BLAST: a better web interface. Nucleic Acids Res. 2008;36:W5–9.
pubmed: 18440982 pmcid: 2447716 doi: 10.1093/nar/gkn201
Crooks GE, Hon G, Chandonia J-M, Brenner SE. WebLogo: a sequence logo generator. Genome Res. 2004;14:1188–90.
pubmed: 15173120 pmcid: 419797 doi: 10.1101/gr.849004
Biswas A, Staals RHJ, Morales SE, Fineran PC, Brown CM. CRISPRDetect: A flexible algorithm to define CRISPR arrays. BMC Genomics. 2016;17:356.
pubmed: 27184979 pmcid: 4869251 doi: 10.1186/s12864-016-2627-0
Gussow AB, Park AE, Borges AL, Shmakov SA, Makarova KS, Wolf YI, et al. Machine-learning approach expands the repertoire of anti-CRISPR protein families. Nat Commun. 2020;11:3784.
pubmed: 32728052 pmcid: 7391736 doi: 10.1038/s41467-020-17652-0
Buchfink B, Xie C, Huson DH. Fast and sensitive protein alignment using DIAMOND. Nat Methods. 2015;12:59–60.
pubmed: 25402007 doi: 10.1038/nmeth.3176
Tisza MJ, Buck CB. A catalog of tens of thousands of viruses from human metagenomes reveals hidden associations with chronic diseases. Proc Natl Acad Sci USA. 2021;118:1–11.
doi: 10.1073/pnas.2023202118
Koppel N, Bisanz JE, Pandelia M-E, Turnbaugh PJ, Balskus EP. Discovery and characterization of a prevalent human gut bacterial enzyme sufficient for the inactivation of a family of plant toxins. eLife. 2018;7:33953.
doi: 10.7554/eLife.33953
Koskella B, Lin DM, Buckling A, Thompson JN. The costs of evolving resistance in heterogeneous parasite environments. Proc Biol Sci. 2012;279:1896–903.
pubmed: 22171085
Harcombe WR, Bull JJ. Impact of phages on two-species bacterial communities. Appl Environ Microbiol. 2005;71:5254–9.
pubmed: 16151111 pmcid: 1214695 doi: 10.1128/AEM.71.9.5254-5259.2005
Seed KD, Yen M, Shapiro BJ, Hilaire IJ, Charles RC, Teng JE, et al. Evolutionary consequences of intra-patient phage predation on microbial populations. eLife. 2014;3:e03497.
pubmed: 25161196 pmcid: 4141277 doi: 10.7554/eLife.03497
Hampton HG, Watson BNJ, Fineran PC. The arms race between bacteria and their phage foes. Nature. 2020;577:327–36.
pubmed: 31942051 doi: 10.1038/s41586-019-1894-8
Meaden S, Capria L, Alseth E, Gandon S, Biswas A, Lenzi L, et al. Phage gene expression and host responses lead to infection-dependent costs of CRISPR immunity. ISME J. 2021;15:534–44.
pubmed: 33011743 doi: 10.1038/s41396-020-00794-w
Lokareddy RK, Sankhala RS, Roy A, Afonine PV, Motwani T, Teschke CM, et al. Portal protein functions akin to a DNA-sensor that couples genome-packaging to icosahedral capsid maturation. Nat Commun. 2017;8:1–11.
doi: 10.1038/ncomms14310
Prevelige PE, Cortines JR. Phage assembly and the special role of the portal protein. Curr Opin Virol. 2018;31:66–73.
pubmed: 30274853 doi: 10.1016/j.coviro.2018.09.004
Eberhardt RY, Bartholdson SJ, Punta M, Bateman A. The SHOCT domain: a widespread domain under-represented in model organisms. PLoS One. 2013;8:1–7.
doi: 10.1371/journal.pone.0057848
Campos-Olivas R, Louis JM, Clérot D, Gronenborn B, Gronenborn AM. The structure of a replication initiator unites diverse aspects of nucleic acid metabolism. Proc Natl Acad Sci USA. 2002;99:10310–5.
pubmed: 12130667 pmcid: 124910 doi: 10.1073/pnas.152342699
Reece RJ, Maxwell A, Wang JC. DNA gyrase: structure and function. Crit Rev Biochem Mol Biol. 1991;26:335–75.
pubmed: 1657531 doi: 10.3109/10409239109114072
Chitra SR, Ramalakshmi N, Arunkumar S, Manimegalai P. A comprehensive review on DNA gyrase inhibitors. Infect Disord Drug Targets. 2020;20:765–77.
pubmed: 33109068 doi: 10.2174/1871526520666200102110235
Hernandez CA, Koskella B. Phage resistance evolution in vitro is not reflective of in vivo outcome in a plant-bacteria-phage system. Evolution (N. Y). 2019;73:2461–75.
Alseth EO, Pursey E, Luján AM, McLeod I, Rollie C, Westra ER. Bacterial biodiversity drives the evolution of CRISPR-based phage resistance. Nature. 2019;574:549–52.
pubmed: 31645729 pmcid: 6837874 doi: 10.1038/s41586-019-1662-9
Lourenço M, Chaffringeon L, Lamy-Besnier Q, Pédron T, Campagne P, Eberl C, et al. The spatial heterogeneity of the gut limits predation and fosters coexistence of bacteria and bacteriophages. Cell Host Microbe. 2020;28:390–401.e5.
pubmed: 32615090 doi: 10.1016/j.chom.2020.06.002
McGinn J, Marraffini LA. CRISPR-Cas systems optimize their immune response by specifying the site of spacer integration. Mol Cell. 2016;64:616–23.
pubmed: 27618488 pmcid: 5096952 doi: 10.1016/j.molcel.2016.08.038
Heler R, Samai P, Modell JW, Weiner C, Goldberg GW, Bikard D, et al. Cas9 specifies functional viral targets during CRISPR-Cas adaptation. Nature. 2015;519:199–202.
pubmed: 25707807 pmcid: 4385744 doi: 10.1038/nature14245
Kieper SN, Almendros C, Behler J, McKenzie RE, Nobrega FL, Haagsma AC, et al. Cas4 facilitates PAM-compatible spacer selection during CRISPR adaptation. Cell Rep. 2018;22:3377–84.
pubmed: 29590607 pmcid: 5896167 doi: 10.1016/j.celrep.2018.02.103
Weissman JL, Stoltzfus A, Westra ER, Johnson PLF. Avoidance of self during CRISPR immunization. Trends Microbiol. 2020;28:543–53.
pubmed: 32544441 doi: 10.1016/j.tim.2020.02.005
Stern A, Keren L, Wurtzel O, Amitai G, Sorek R. Self-targeting by CRISPR: gene regulation or autoimmunity? Trends Genet. 2010;26:335–40.
pubmed: 20598393 pmcid: 2910793 doi: 10.1016/j.tig.2010.05.008
Stachler A-E, Turgeman-Grott I, Shtifman-Segal E, Allers T, Marchfelder A, Gophna U. High tolerance to self-targeting of the genome by the endogenous CRISPR-Cas system in an archaeon. Nucleic Acids Res. 2017;45:5208–16.
pubmed: 28334774 pmcid: 5435918 doi: 10.1093/nar/gkx150
Shah SA, Erdmann S, Mojica FJM, Garrett RA. Protospacer recognition motifs: mixed identities and functional diversity. RNA Biol. 2013;10:891–9.
pubmed: 23403393 pmcid: 3737346 doi: 10.4161/rna.23764
Sausset R, Petit MA, Gaboriau-Routhiau V, de Paepe M. New insights into intestinal phages. Mucosal Immunol. 2020;13:205–15.
pubmed: 31907364 pmcid: 7039812 doi: 10.1038/s41385-019-0250-5
Yang C, Ottemann KM. Control of bacterial colonization in the glands and crypts. Curr Opin Microbiol. 2019;47:38–44.
pubmed: 30502720 doi: 10.1016/j.mib.2018.11.004
Eriksen RS, Mitarai N, Sneppen K. Sustainability of spatially distributed bacteria-phage systems. Sci Rep. 2020;10:3154.
pubmed: 32081858 pmcid: 7035299 doi: 10.1038/s41598-020-59635-7
Eriksen RS, Svenningsen SL, Sneppen K, Mitarai N. A growing microcolony can survive and support persistent propagation of virulent phages. Proc Natl Acad Sci USA. 2017;115:337–42.
pubmed: 29259110 pmcid: 5777033 doi: 10.1073/pnas.1708954115
Abedon ST. Phage ‘delay’ towards enhancing bacterial escape from biofilms: a more comprehensive way of viewing resistance to bacteriophages. AIMS Microbiol. 2017;3:186–226.
pubmed: 31294157 pmcid: 6605007 doi: 10.3934/microbiol.2017.2.186
Weissman JL, Holmes R, Barrangou R, Moineau S, Fagan WF, Levin B, et al. Immune loss as a driver of coexistence during host-phage coevolution. ISME J. 2018;12:585–97.
pubmed: 29328063 pmcid: 5776473 doi: 10.1038/ismej.2017.194
Common J, Morley D, Westra ER, van Houte S. CRISPR-Cas immunity leads to a coevolutionary arms race between Streptococcus thermophilus and lytic phage. Philos Trans R Soc Lond B Biol Sci. 2019;374:20180098.
pubmed: 30905285 pmcid: 6452269 doi: 10.1098/rstb.2018.0098
Hanshew AS, Mason CJ, Raffa KF, Currie CR. Minimization of chloroplast contamination in 16S rRNA gene pyrosequencing of insect herbivore bacterial communities. J Microbiol Methods. 2013;95:149–55.
pubmed: 23968645 pmcid: 4133986 doi: 10.1016/j.mimet.2013.08.007
Dron M, Rahire M, Rochaix JD. Sequence of the chloroplast 16S rRNA gene and its surrounding regions of Chlamydomonas reinhardii. Nucleic Acids Res. 1982;10:7609–20.
pubmed: 6296784 pmcid: 327033 doi: 10.1093/nar/10.23.7609
Gupta RS, Patel S. Robust demarcation of the family Caryophanaceae (Planococcaceae) and its different genera including three novel genera based on phylogenomics and highly specific molecular signatures. Front Microbiol. 2019;10:2821.
pubmed: 32010063 doi: 10.3389/fmicb.2019.02821
Tanaka M, Onizuka S, Mishima R, Nakayama J. Cultural isolation of spore-forming bacteria in human feces using bile acids. Sci Rep. 2020;10:15041.
pubmed: 32929101 pmcid: 7490687 doi: 10.1038/s41598-020-71883-1
Browne HP, Almeida A, Kumar N, Vervier K, Adoum AT, Viciani E, et al. Host adaptation in gut Firmicutes is associated with sporulation loss and altered transmission cycle. Genome Biol. 2021;22:204.
pubmed: 34348764 pmcid: 8340488 doi: 10.1186/s13059-021-02428-6
Beno SM, Cheng RA, Orsi RH, Duncan DR, Guo X, Kovac J, et al. Paenibacillus odorifer, the predominant Paenibacillus species isolated from milk in the United States, demonstrates genetic and phenotypic conservation of psychrotolerance but clade-asociated differences in nitrogen metabolic pathways. mSphere. 2020;5:e00739–19.
pubmed: 31969477 pmcid: 7407005 doi: 10.1128/mSphere.00739-19
Paredes-Sabja D, Torres JA, Setlow P, Sarker MR. Clostridium perfringens spore germination: characterization of germinants and their receptors. J Bacteriol. 2008;190:1190–201.
pubmed: 18083820 doi: 10.1128/JB.01748-07
Shen A, Edwards AN, Sarker MR, Paredes-Sabja D. Sporulation and germination in clostridial pathogens. Microbiol Spectr. 2019;7:903–26.
doi: 10.1128/microbiolspec.GPP3-0017-2018
Setlow P. Germination of spores of Bacillus species: what we know and do not know. J Bacteriol. 2014;196:1297–305.
pubmed: 24488313 pmcid: 3993344 doi: 10.1128/JB.01455-13
Setlow P, Christie G. What’s new and notable in bacterial spore killing! World J Microbiol Biotechnol. 2021;37:144.
pubmed: 34351499 pmcid: 8342367 doi: 10.1007/s11274-021-03108-0
Setlow P. Spores of Bacillus subtilis: Their resistance to and killing by radiation, heat and chemicals. J Appl Microbiol. 2006;101:514–25.
pubmed: 16907802 doi: 10.1111/j.1365-2672.2005.02736.x

Auteurs

Torben Sølbeck Rasmussen (TS)

Section of Microbiology and Fermentation, Department of Food Science, Faculty of Science, University of Copenhagen, 1958, Frederiksberg, Denmark. torben@food.ku.dk.

Anna Kirstine Koefoed (AK)

Section of Microbiology and Fermentation, Department of Food Science, Faculty of Science, University of Copenhagen, 1958, Frederiksberg, Denmark.

Ling Deng (L)

Section of Microbiology and Fermentation, Department of Food Science, Faculty of Science, University of Copenhagen, 1958, Frederiksberg, Denmark.

Musemma K Muhammed (MK)

Section of Microbiology and Fermentation, Department of Food Science, Faculty of Science, University of Copenhagen, 1958, Frederiksberg, Denmark.

Geneviève M Rousseau (GM)

Département de biochimie, de microbiologie et de bio-informatique, Faculté des sciences et de 1enie, Université Laval, Québec, QC, G1V 0A6, Canada.
Groupe de recherche en écologie buccale, Faculté de médecine dentaire, Université Laval, Québec, QC, G1V 0A6, Canada.

Witold Kot (W)

Section of Microbial Ecology and Biotechnology, Department of Plant and Environmental Sciences, University of Copenhagen, 1871, Frederiksberg, Denmark.

Sabrina Sprotte (S)

Department of Microbiology and Biotechnology, Max Rubner-Institut, 24103, Kiel, Germany.

Horst Neve (H)

Department of Microbiology and Biotechnology, Max Rubner-Institut, 24103, Kiel, Germany.

Charles M A P Franz (CMAP)

Department of Microbiology and Biotechnology, Max Rubner-Institut, 24103, Kiel, Germany.

Axel Kornerup Hansen (AK)

Section of Experimental Animal Models, Department of Veterinary and Animal Sciences, University of Copenhagen, 1871, Frederiksberg, Denmark.

Finn Kvist Vogensen (FK)

Section of Microbiology and Fermentation, Department of Food Science, Faculty of Science, University of Copenhagen, 1958, Frederiksberg, Denmark.

Sylvain Moineau (S)

Département de biochimie, de microbiologie et de bio-informatique, Faculté des sciences et de 1enie, Université Laval, Québec, QC, G1V 0A6, Canada.
Groupe de recherche en écologie buccale, Faculté de médecine dentaire, Université Laval, Québec, QC, G1V 0A6, Canada.
Félix d'Hérelle Reference Center for Bacterial Viruses, Faculté de médecine dentaire, Université Laval, Québec, QC, G1V 0A6, Canada.

Dennis Sandris Nielsen (DS)

Section of Microbiology and Fermentation, Department of Food Science, Faculty of Science, University of Copenhagen, 1958, Frederiksberg, Denmark. dn@food.ku.dk.

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