Investigation of the domain line tension in asymmetric vesicles prepared via hemifusion.

Asymmetric bilayers Asymmetric giant unilamellar vesicles Coexistence of liquid disordered and liquid ordered phases Domain line tension Hemifusion

Journal

Biochimica et biophysica acta. Biomembranes
ISSN: 1879-2642
Titre abrégé: Biochim Biophys Acta Biomembr
Pays: Netherlands
ID NLM: 101731713

Informations de publication

Date de publication:
01 06 2021
Historique:
received: 30 09 2020
revised: 28 01 2021
accepted: 12 02 2021
pubmed: 2 3 2021
medline: 9 9 2021
entrez: 1 3 2021
Statut: ppublish

Résumé

The plasma membrane (PM) is asymmetric in lipid composition. The distinct and characteristic lipid compositions of the exoplasmic and cytoplasmic leaflets lead to different lipid-lipid interactions and physical-chemical properties in each leaflet. The exoplasmic leaflet possesses an intrinsic ability to form coexisting ordered and disordered fluid domains, whereas the cytoplasmic leaflet seems to form a single fluid phase. To better understand the interleaflet interactions that influence domains, we compared asymmetric model membranes that capture salient properties of the PM with simpler symmetric membranes. Using asymmetric giant unilamellar vesicles (aGUVs) prepared by hemifusion with a supported lipid bilayer, we investigate the domain line tension that characterizes the behavior of coexisting ordered + disordered domains. The line tension can be related to the contact perimeter of the different phases. Compared to macroscopic phase separation, the appearance of modulated phases was found to be a robust indicator of a decrease in domain line tension. Symmetric GUVs of 1,2-distearoyl-sn-glycero-3-phosphocholine (DSPC)/1,2-dioleoyl-sn-glycero-3-phosphocholine (DOPC)/1-palmitoyl-2-oleoyl-glycero-3-phosphocholine (POPC)/cholesterol (chol) were formed into aGUVs by replacing the GUV outer leaflet with DOPC/chol = 0.8/0.2 in order to create a cytoplasmic leaflet model. These aGUVs revealed lower line tension for the ordered + disordered domains of the exoplasmic model leaflet.

Identifiants

pubmed: 33647248
pii: S0005-2736(21)00037-7
doi: 10.1016/j.bbamem.2021.183586
pmc: PMC8606045
mid: NIHMS1684440
pii:
doi:

Substances chimiques

Phosphatidylcholines 0
Unilamellar Liposomes 0
Cholesterol 97C5T2UQ7J
1,2-distearoyllecithin EAG959U971
1,2-oleoylphosphatidylcholine EDS2L3ODLV
1-palmitoyl-2-oleoylphosphatidylcholine TE895536Y5

Types de publication

Journal Article Research Support, N.I.H., Extramural Research Support, Non-U.S. Gov't Research Support, U.S. Gov't, Non-P.H.S.

Langues

eng

Sous-ensembles de citation

IM

Pagination

183586

Subventions

Organisme : NIGMS NIH HHS
ID : R01 GM105684
Pays : United States

Informations de copyright

Copyright © 2021 Elsevier B.V. All rights reserved.

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Auteurs

Thais A Enoki (TA)

Department of Molecular Biology and Genetics, Cornell University, Ithaca, NY, United States of America; Department of Chemistry, University of Tennessee, Knoxville, TN, United States of America. Electronic address: ta327@cornell.edu.

Joy Wu (J)

Department of Molecular Biology and Genetics, Cornell University, Ithaca, NY, United States of America.

Frederick A Heberle (FA)

Department of Chemistry, University of Tennessee, Knoxville, TN, United States of America.

Gerald W Feigenson (GW)

Department of Molecular Biology and Genetics, Cornell University, Ithaca, NY, United States of America.

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Classifications MeSH