Chd1 protects genome integrity at promoters to sustain hypertranscription in embryonic stem cells.


Journal

Nature communications
ISSN: 2041-1723
Titre abrégé: Nat Commun
Pays: England
ID NLM: 101528555

Informations de publication

Date de publication:
11 08 2021
Historique:
received: 12 06 2019
accepted: 20 07 2021
entrez: 12 8 2021
pubmed: 13 8 2021
medline: 24 8 2021
Statut: epublish

Résumé

Stem and progenitor cells undergo a global elevation of nascent transcription, or hypertranscription, during key developmental transitions involving rapid cell proliferation. The chromatin remodeler Chd1 mediates hypertranscription in pluripotent cells but its mechanism of action remains poorly understood. Here we report a novel role for Chd1 in protecting genome integrity at promoter regions by preventing DNA double-stranded break (DSB) accumulation in ES cells. Chd1 interacts with several DNA repair factors including Atm, Parp1, Kap1 and Topoisomerase 2β and its absence leads to an accumulation of DSBs at Chd1-bound Pol II-transcribed genes and rDNA. Genes prone to DNA breaks in Chd1 KO ES cells are longer genes with GC-rich promoters, a more labile nucleosomal structure and roles in chromatin regulation, transcription and signaling. These results reveal a vulnerability of hypertranscribing stem cells to accumulation of endogenous DNA breaks, with important implications for developmental and cancer biology.

Identifiants

pubmed: 34381042
doi: 10.1038/s41467-021-25088-3
pii: 10.1038/s41467-021-25088-3
pmc: PMC8357957
doi:

Substances chimiques

Chd1 protein, mouse 0
Chromatin 0
DNA, Ribosomal 0
DNA-Binding Proteins 0
Poly-ADP-Ribose Binding Proteins 0
DNA Topoisomerases, Type II EC 5.99.1.3
Top2b protein, mouse EC 5.99.1.3

Types de publication

Journal Article Research Support, N.I.H., Extramural Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

4859

Subventions

Organisme : NIGMS NIH HHS
ID : R01 GM123556
Pays : United States
Organisme : Cancer Research UK
ID : C5759/A25254
Pays : United Kingdom
Organisme : NIGMS NIH HHS
ID : R01 GM113014
Pays : United States
Organisme : Department of Health
Pays : United Kingdom
Organisme : NCI NIH HHS
ID : R01 CA163336
Pays : United States
Organisme : CIHR
ID : 420231
Pays : Canada
Organisme : NCI NIH HHS
ID : P30 CA082103
Pays : United States

Informations de copyright

© 2021. The Author(s).

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Auteurs

Aydan Bulut-Karslioglu (A)

Eli and Edythe Broad Center of Regeneration Medicine and Stem Cell Research, Center for Reproductive Sciences and Diabetes Center, University of California, San Francisco, San Francisco, CA, USA. Aydan.karslioglu@molgen.mpg.de.
Max Planck Institute for Molecular Genetics, Berlin, Germany. Aydan.karslioglu@molgen.mpg.de.

Hu Jin (H)

Carl R. Woese Institute for Genomic Biology, Urbana, IL, USA.
Department of Physics, University of Illinois, Urbana-Champaign, Urbana, IL, USA.
Department of Biomedical Informatics, Harvard Medical School, Boston, MA, USA.

Yun-Kyo Kim (YK)

Lunenfeld-Tanenbaum Research Institute and Department of Molecular Genetics, University of Toronto, Toronto, ON, Canada.

Brandon Cho (B)

Lunenfeld-Tanenbaum Research Institute and Department of Molecular Genetics, University of Toronto, Toronto, ON, Canada.

Marcela Guzman-Ayala (M)

Eli and Edythe Broad Center of Regeneration Medicine and Stem Cell Research, Center for Reproductive Sciences and Diabetes Center, University of California, San Francisco, San Francisco, CA, USA.
Senti Biosciences, South San Francisco, CA, USA.

Andrew J K Williamson (AJK)

Stoller Biomarker Discovery Centre, The University of Manchester, Manchester, UK.
Thermo Fisher Scientific, Stafford House, UK.

Miroslav Hejna (M)

Carl R. Woese Institute for Genomic Biology, Urbana, IL, USA.
Department of Physics, University of Illinois, Urbana-Champaign, Urbana, IL, USA.

Maximilian Stötzel (M)

Max Planck Institute for Molecular Genetics, Berlin, Germany.

Anthony D Whetton (AD)

Stoller Biomarker Discovery Centre, The University of Manchester, Manchester, UK.

Jun S Song (JS)

Carl R. Woese Institute for Genomic Biology, Urbana, IL, USA.
Department of Physics, University of Illinois, Urbana-Champaign, Urbana, IL, USA.

Miguel Ramalho-Santos (M)

Eli and Edythe Broad Center of Regeneration Medicine and Stem Cell Research, Center for Reproductive Sciences and Diabetes Center, University of California, San Francisco, San Francisco, CA, USA. mrsantos@lunenfeld.ca.
Lunenfeld-Tanenbaum Research Institute and Department of Molecular Genetics, University of Toronto, Toronto, ON, Canada. mrsantos@lunenfeld.ca.

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Classifications MeSH