Simultaneous imaging of ER and cytosolic Ca2+ dynamics reveals long-distance ER Ca2+ waves in plants.


Journal

Plant physiology
ISSN: 1532-2548
Titre abrégé: Plant Physiol
Pays: United States
ID NLM: 0401224

Informations de publication

Date de publication:
05 10 2021
Historique:
received: 02 02 2021
accepted: 12 05 2021
entrez: 5 10 2021
pubmed: 6 10 2021
medline: 1 3 2022
Statut: ppublish

Résumé

Calcium ions (Ca2+) play a key role in cell signaling across organisms. In plants, a plethora of environmental and developmental stimuli induce specific Ca2+ increases in the cytosol as well as in different cellular compartments including the endoplasmic reticulum (ER). The ER represents an intracellular Ca2+ store that actively accumulates Ca2+ taken up from the cytosol. By exploiting state-of-the-art genetically encoded Ca2+ indicators, specifically the ER-GCaMP6-210 and R-GECO1, we report the generation and characterization of an Arabidopsis (Arabidopsis thaliana) line that allows for simultaneous imaging of Ca2+ dynamics in both the ER and cytosol at different spatial scales. By performing analyses in single cells, we precisely quantified (1) the time required by the ER to import Ca2+ from the cytosol into the lumen and (2) the time required to observe a cytosolic Ca2+ increase upon the pharmacological inhibition of the ER-localized P-Type IIA Ca2+-ATPases. Furthermore, live imaging of mature, soil-grown plants revealed the existence of a wounding-induced, long-distance ER Ca2+ wave propagating in injured and systemic rosette leaves. This technology enhances high-resolution analyses of intracellular Ca2+ dynamics at the cellular level and in adult organisms and paves the way to develop new methodologies aimed at defining the contribution of subcellular compartments in Ca2+ homeostasis and signaling.

Identifiants

pubmed: 34608947
pii: 6288498
doi: 10.1093/plphys/kiab251
pmc: PMC8491065
doi:

Substances chimiques

Calcium SY7Q814VUP

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

603-617

Informations de copyright

© American Society of Plant Biologists 2021. All rights reserved. For permissions, please email: journals.permissions@oup.com.

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Auteurs

Francesca Resentini (F)

Department of Biosciences, University of Milan, Milan 20133, Italy.

Matteo Grenzi (M)

Department of Biosciences, University of Milan, Milan 20133, Italy.

Daniele Ancora (D)

Department of Physics, Politecnico di Milano, Milan 20133, Italy.

Mara Cademartori (M)

Department of Biosciences, University of Milan, Milan 20133, Italy.

Laura Luoni (L)

Department of Biosciences, University of Milan, Milan 20133, Italy.

Marianna Franco (M)

Department of Biosciences, University of Milan, Milan 20133, Italy.

Andrea Bassi (A)

Department of Physics, Politecnico di Milano, Milan 20133, Italy.

Maria Cristina Bonza (MC)

Department of Biosciences, University of Milan, Milan 20133, Italy.

Alex Costa (A)

Department of Biosciences, University of Milan, Milan 20133, Italy.
Institute of Biophysics, Consiglio Nazionale Delle Ricerche, Milan 20133, Italy.

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Classifications MeSH