Alternative sulphur metabolism in the fungal pathogen Candida parapsilosis.


Journal

Nature communications
ISSN: 2041-1723
Titre abrégé: Nat Commun
Pays: England
ID NLM: 101528555

Informations de publication

Date de publication:
24 Oct 2024
Historique:
received: 12 02 2024
accepted: 08 10 2024
medline: 25 10 2024
pubmed: 25 10 2024
entrez: 25 10 2024
Statut: epublish

Résumé

Candida parapsilosis is an opportunistic fungal pathogen commonly isolated from the environment and associated with nosocomial infection outbreaks worldwide. We describe here the construction of a large collection of gene disruptions, greatly increasing the molecular tools available for probing gene function in C. parapsilosis. We use these to identify transcription factors associated with multiple metabolic pathways, and in particular to dissect the network regulating the assimilation of sulphur. We find that, unlike in other yeasts and filamentous fungi, the transcription factor Met4 is not the main regulator of methionine synthesis. In C. parapsilosis, assimilation of inorganic sulphur (sulphate) and synthesis of cysteine and methionine is regulated by Met28, a paralog of Met4, whereas Met4 regulates expression of a wide array of transporters and enzymes involved in the assimilation of organosulfur compounds. Analysis of transcription factor binding sites suggests that Met4 is recruited by the DNA-binding protein Met32, and Met28 is recruited by Cbf1. Despite having different target genes, Met4 and Met28 have partial functional overlap, possibly because Met4 can contribute to assimilation of inorganic sulphur in the absence of Met28.

Identifiants

pubmed: 39448588
doi: 10.1038/s41467-024-53442-8
pii: 10.1038/s41467-024-53442-8
doi:

Substances chimiques

Sulfur 70FD1KFU70
Fungal Proteins 0
Transcription Factors 0
Methionine AE28F7PNPL
Cysteine K848JZ4886
Sulfates 0

Types de publication

Journal Article

Langues

eng

Sous-ensembles de citation

IM

Pagination

9190

Subventions

Organisme : Science Foundation Ireland (SFI)
ID : 19/FFP/6668

Informations de copyright

© 2024. The Author(s).

Références

Brown, G. D. et al. Hidden killers: human fungal infections. Sci. Transl. Med. 4, 165rv113 (2012).
doi: 10.1126/scitranslmed.3004404
Rauseo, A. M., Coler-Reilly, A., Larson, L. & Spec, A. Hope on the horizon: Novel fungal treatments in development. Open Forum Infect. Dis. 7, ofaa016 (2020).
pubmed: 32099843 pmcid: 7031074 doi: 10.1093/ofid/ofaa016
Tóth, R. et al. Candida parapsilosis: from genes to the bedside. Clin. Microbiol Rev. 32, e00111–e00118 (2019).
pubmed: 30814115 pmcid: 6431126 doi: 10.1128/CMR.00111-18
Bassetti, M. et al. Incidence and outcome of invasive candidiasis in intensive care units (ICUs) in Europe: results of the EUCANDICU project. Crit. Care 23, 219 (2019).
pubmed: 31200780 pmcid: 6567430 doi: 10.1186/s13054-019-2497-3
Santos, M. A. S., Gomes, A. C., Santos, M. C., Carreto, L. C. & Moura, G. R. The genetic code of the fungal CTG clade. C. R. Biol. 334, 607–611 (2011).
pubmed: 21819941 doi: 10.1016/j.crvi.2011.05.008
Perez, J. C. Fungi of the human gut microbiota: Roles and significance. Int J. Med. Microbiol. 311, 151490 (2021).
pubmed: 33676239 doi: 10.1016/j.ijmm.2021.151490
Dogen, A. et al. Candida parapsilosis in domestic laundry machines. Med. Mycol. 55, 813–819 (2017).
pubmed: 28204594 doi: 10.1093/mmy/myx008
Trofa, D., Gácser, A. & Nosanchuk, J. D. Candida parapsilosis, an Emerging Fungal Pathogen. Clin. Microbiol. Rev. 21, 606–625 (2008).
pubmed: 18854483 pmcid: 2570155 doi: 10.1128/CMR.00013-08
Daneshnia, F. et al. Determinants of fluconazole resistance and echinocandin tolerance in C. parapsilosis isolates causing a large clonal candidemia outbreak among COVID-19 patients in a Brazilian ICU. Emerg. Microbes Infect. 1–33 https://doi.org/10.1080/22221751.2022.2117093 (2022).
Thomaz, D. Y. et al. A Brazilian inter-hospital candidemia outbreak caused by fluconazole-resistant Candida parapsilosis in the COVID-19 era. J. Fungi 8, 100 (2022).
doi: 10.3390/jof8020100
Guo, W. et al. An outbreak of Candida parapsilosis fungemia among preterm infants. Genet Mol. Res. 14, 18259–18267 (2015).
pubmed: 26782473 doi: 10.4238/2015.December.23.13
Zhai, B. et al. High-resolution mycobiota analysis reveals dynamic intestinal translocation preceding invasive candidiasis. Nat. Med. 26, 59–64 (2020).
pubmed: 31907459 pmcid: 7005909 doi: 10.1038/s41591-019-0709-7
Rolling, T. et al. Haematopoietic cell transplantation outcomes are linked to intestinal mycobiota dynamics and an expansion of Candida parapsilosis complex species. Nat. Microbiol. 6, 1505–1515 (2021).
pubmed: 34764444 pmcid: 8939874 doi: 10.1038/s41564-021-00989-7
Hsu, P. C. et al. Plastic rewiring of Sef1 transcriptional networks and the potential of nonfunctional transcription factor binding in facilitating adaptive evolution. Mol. Biol. Evol. 38, 4732–4747 (2021).
pubmed: 34175931 pmcid: 8557406 doi: 10.1093/molbev/msab192
Tsong, A. E., Miller, M. G., Raisner, R. M. & Johnson, A. D. Evolution of a combinatorial transcriptional circuit: a case study in yeasts. Cell 115, 389–399 (2003).
pubmed: 14622594 doi: 10.1016/S0092-8674(03)00885-7
Nocedal, I., Mancera, E. & Johnson, A. D. Gene regulatory network plasticity predates a switch in function of a conserved transcription regulator. Elife 6, e23250 (2017).
pubmed: 28327289 pmcid: 5391208 doi: 10.7554/eLife.23250
Alves, R. et al. Adapting to survive: How Candida overcomes host-imposed constraints during human colonization. PLoS Pathog. 16, e1008478 (2020).
pubmed: 32437438 pmcid: 7241708 doi: 10.1371/journal.ppat.1008478
Ding, C. et al. Conserved and divergent roles of Bcr1 and CFEM proteins in Candida parapsilosis and Candida albicans. PLoS ONE 6, e28151 (2011).
pubmed: 22145027 pmcid: 3228736 doi: 10.1371/journal.pone.0028151
Holland, L. M. et al. Comparative phenotypic analysis of the major fungal pathogens Candida parapsilosis and Candida albicans. PLoS Pathog. 10, e1004365 (2014).
pubmed: 25233198 pmcid: 4169492 doi: 10.1371/journal.ppat.1004365
Mancera, E. et al. Evolution of the complex transcription network controlling biofilm formation in Candida species. ELife 10, e64682 (2021).
pubmed: 33825680 pmcid: 8075579 doi: 10.7554/eLife.64682
Perez, J. C. The interplay between gut bacteria and the yeast Candida albicans. Gut Microbes 13, 1979877 (2021).
pubmed: 34586038 pmcid: 8489915 doi: 10.1080/19490976.2021.1979877
Brunke, S., Mogavero, S., Kasper, L. & Hube, B. Virulence factors in fungal pathogens of man. Curr. Opin. Microbiol. 32, 89–95 (2016).
pubmed: 27257746 doi: 10.1016/j.mib.2016.05.010
Padder, S. A., Ramzan, A., Tahir, I., Rehman, R. U. & Shah, A. H. Metabolic flexibility and extensive adaptability governing multiple drug resistance and enhanced virulence in Candida albicans. Crit. Rev. Microbiol. 48, 1–20 (2022).
pubmed: 34213983 doi: 10.1080/1040841X.2021.1935447
Amich, J., Krappmann, S. & Bachhawat, A. K. Editorial: Sulphur metabolism of Fungi - implications for virulence and opportunities for therapy. Front. Microbiol. 11, 583689 (2020).
pubmed: 33117323 pmcid: 7578252 doi: 10.3389/fmicb.2020.583689
Ljungdahl, P. O. & Daignan-Fornier, B. Regulation of amino acid, nucleotide, and phosphate metabolism in Saccharomyces cerevisiae. Genetics 190, 885–929 (2012).
pubmed: 22419079 pmcid: 3296254 doi: 10.1534/genetics.111.133306
Thomas, D. & Surdin-Kerjan, Y. Metabolism of sulfur amino acids in Saccharomyces cerevisiae. Microbiol. Mol. Biol. Rev. 61, 503–532 (1997).
pubmed: 9409150 pmcid: 232622
Lee, T. A. et al. Dissection of combinatorial control by the Met4 transcriptional complex. Mol. Biol. Cell 21, 456–469 (2010).
pubmed: 19940020 pmcid: 2814790 doi: 10.1091/mbc.e09-05-0420
Hebert, A., Casaregola, S. & Beckerich, J. M. Biodiversity in sulfur metabolism in hemiascomycetous yeasts. FEMS Yeast Res. 11, 366–378 (2011).
pubmed: 21348937 doi: 10.1111/j.1567-1364.2011.00725.x
Pacheco, D. et al. Transcription activation domains of the yeast factors Met4 and Ino2: tandem activation domains with properties similar to the yeast Gcn4 activator. Mol. Cell Biol. 38, e00038–18 (2018).
pubmed: 29507182 pmcid: 5954196 doi: 10.1128/MCB.00038-18
Kuras, L., Barbey, R. & Thomas, D. Assembly of a bZIP-bHLH transcription activation complex: formation of the yeast Cbf1-Met4-Met28 complex is regulated through Met28 stimulation of Cbf1 DNA binding. Embo j. 16, 2441–2451 (1997).
pubmed: 9171357 pmcid: 1169844 doi: 10.1093/emboj/16.9.2441
Blaiseau, P. L. & Thomas, D. Multiple transcriptional activation complexes tether the yeast activator Met4 to DNA. Embo j. 17, 6327–6336 (1998).
pubmed: 9799240 pmcid: 1170957 doi: 10.1093/emboj/17.21.6327
Shrivastava, M. et al. Modulation of the complex regulatory network for methionine biosynthesis in fungi. Genetics 217, iyaa049 (2021).
pubmed: 33724418 pmcid: 8045735 doi: 10.1093/genetics/iyaa049
Natorff, R., Sieńko, M., Brzywczy, J. & Paszewski, A. The Aspergillus nidulans metR gene encodes a bZIP protein which activates transcription of sulphur metabolism genes. Mol. Microbiol. 49, 1081–1094 (2003).
pubmed: 12890030 doi: 10.1046/j.1365-2958.2003.03617.x
Huberman, L. B. et al. Aspects of the Neurospora crassa sulfur starvation response are revealed by transcriptional profiling and DNA affinity purification sequencing. mSphere 6, e0056421 (2021). 10.1128/msphere.00564-00521.
pubmed: 34523983 doi: 10.1128/mSphere.00564-21
Lombardi, L., Oliveira-Pacheco, J. & Butler, G. Plasmid-based CRISPR-Cas9 gene editing in multiple Candida Species. mSphere 4, e00125–19 (2019).
pubmed: 30867327 pmcid: 6416365 doi: 10.1128/mSphere.00125-19
Hoot, S. J., Oliver, B. G. & White, T. C. Candida albicans UPC2 is transcriptionally induced in response to antifungal drugs and anaerobicity through Upc2p-dependent and -independent mechanisms. Microbiology 154, 2748–2756 (2008).
pubmed: 18757808 doi: 10.1099/mic.0.2008/017475-0
Homann, O. R., Dea, J., Noble, S. M. & Johnson, A. D. A phenotypic profile of the Candida albicans regulatory network. PLoS Genet. 5, e1000783 (2009).
pubmed: 20041210 pmcid: 2790342 doi: 10.1371/journal.pgen.1000783
Enjalbert, B. et al. Role of the Hog1 stress-activated protein kinase in the global transcriptional response to stress in the fungal pathogen Candida albicans. Mol. Biol. Cell 17, 1018–1032 (2006).
pubmed: 16339080 pmcid: 1356608 doi: 10.1091/mbc.e05-06-0501
Alonso-Monge, R. et al. The Hog1 mitogen-activated protein kinase is essential in the oxidative stress response and chlamydospore formation in Candida albicans. Eukaryot. Cell 2, 351–361 (2003).
pubmed: 12684384 pmcid: 154845 doi: 10.1128/EC.2.2.351-361.2003
Arana, D. M., Nombela, C., Alonso-Monge, R. & Pla, J. The Pbs2 MAP kinase kinase is essential for the oxidative-stress response in the fungal pathogen Candida albicans. Microbiology 151, 1033–1049 (2005).
pubmed: 15817773 doi: 10.1099/mic.0.27723-0
Jiang, L. et al. Cadmium-induced activation of high osmolarity glycerol pathway through its Sln1 branch is dependent on the MAP kinase kinase kinase Ssk2, but not its paralog Ssk22, in budding yeast. FEMS Yeast Res. 14, 1263–1272 (2014).
pubmed: 25331360 doi: 10.1111/1567-1364.12220
Kitanovic, A. et al. Phosphatidylinositol 3-kinase VPS34 of Candida albicans is involved in filamentous growth, secretion of aspartic proteases, and intracellular detoxification. FEMS Yeast Res. 5, 431–439 (2005).
pubmed: 15691748 doi: 10.1016/j.femsyr.2004.11.005
Blankenship, J. R., Fanning, S., Hamaker, J. J. & Mitchell, A. P. An extensive circuitry for cell wall regulation in Candida albicans. PLoS Pathog. 6, e1000752 (2010).
pubmed: 20140194 pmcid: 2816693 doi: 10.1371/journal.ppat.1000752
Rai, M. N., Sharma, V., Balusu, S. & Kaur, R. An essential role for phosphatidylinositol 3-kinase in the inhibition of phagosomal maturation, intracellular survival and virulence in Candida glabrata. Cell Microbiol. 17, 269–287 (2015).
pubmed: 25223215 doi: 10.1111/cmi.12364
Turner, S. A., Ma, Q., Ola, M., Martinez de San Vicente, K. & Butler, G. Dal81 Regulates expression of arginine metabolism genes in Candida parapsilosis. mSphere 3, e00028–18 (2018).
pubmed: 29564399 pmcid: 5853489 doi: 10.1128/mSphere.00028-18
Liao, W. L., Ramón, A. M. & Fonzi, W. A. GLN3 encodes a global regulator of nitrogen metabolism and virulence of C. albicans. Fungal Genet. Biol. 45, 514–526 (2008).
pubmed: 17950010 doi: 10.1016/j.fgb.2007.08.006
Tebung, W. A., Choudhury, B. I., Tebbji, F., Morschhäuser, J. & Whiteway, M. Rewiring of the Ppr1 zinc cluster transcription factor from purine catabolism to pyrimidine biogenesis in the saccharomycetaceae. Curr. Biol. 26, 1677–1687 (2016).
pubmed: 27321996 doi: 10.1016/j.cub.2016.04.064
Ghosh, S., Kebaara, B. W., Atkin, A. L. & Nickerson, K. W. Regulation of aromatic alcohol production in Candida albicans. Appl. Environ. Microbiol. 74, 7211–7218 (2008).
pubmed: 18836025 pmcid: 2592902 doi: 10.1128/AEM.01614-08
Masselot, M. & De Robichon-Szulmajster, H. Methionine biosynthesis in Saccharomyces cerevisiae. I. Genetical analysis of auxotrophic mutants. Mol. Gen. Genet. 139, 121–132 (1975).
pubmed: 1101032 doi: 10.1007/BF00264692
Yoo, S. J., Sohn, M. J., Jeong, D. M. & Kang, H. A. Short bZIP homologue of sulfur regulator Met4 from Ogataea parapolymorpha does not depend on DNA-binding cofactors for activating genes in sulfur starvation. Environ. Microbiol. 22, 310–328 (2020).
pubmed: 31680403 doi: 10.1111/1462-2920.14849
Kuras, L. & Thomas, D. Functional analysis of Met4, a yeast transcriptional activator responsive to S-adenosylmethionine. Mol. Cell Biol. 15, 208–216 (1995).
pubmed: 7799928 pmcid: 231936 doi: 10.1128/MCB.15.1.208
Byrne, K. P. & Wolfe, K. H. The Yeast Gene Order Browser: combining curated homology and syntenic context reveals gene fate in polyploid species. Genome Res. 15, 1456–1461 (2005).
pubmed: 16169922 pmcid: 1240090 doi: 10.1101/gr.3672305
Maguire, S. L. et al. Comparative genome analysis and gene finding in Candida species using CGOB. Mol. Biol. Evol. 30, 1281–1291 (2013).
pubmed: 23486613 pmcid: 3649674 doi: 10.1093/molbev/mst042
Gouy, M., Guindon, S. & Gascuel, O. SeaView version 4: A multiplatform graphical user interface for sequence alignment and phylogenetic tree building. Mol. Biol. Evol. 27, 221–224 (2010).
pubmed: 19854763 doi: 10.1093/molbev/msp259
Huang, M. Y., Woolford, C. A., May, G., McManus, C. J. & Mitchell, A. P. Circuit diversification in a biofilm regulatory network. PLOS Pathog. 15, e1007787 (2019).
pubmed: 31116789 pmcid: 6530872 doi: 10.1371/journal.ppat.1007787
Do, E. et al. Collaboration between antagonistic cell type regulators governs natural variation in the Candida albicans biofilm and hyphal gene expression network. mBio 13, e0193722 (2022).
pubmed: 35993746 doi: 10.1128/mbio.01937-22
Cravener, M. V. et al. Reinforcement amid genetic diversity in the Candida albicans biofilm regulatory network. PLoS Pathog. 19, e1011109 (2023).
pubmed: 36696432 pmcid: 9901766 doi: 10.1371/journal.ppat.1011109
Bergin, S. A. et al. Systematic analysis of copy number variations in the pathogenic yeast Candida parapsilosis identifies a gene amplification in RTA3 that is associated with drug resistance. mBio 0, e01777–01722 (2022).
Holt, S. et al. Major sulfonate transporter Soa1 in Saccharomyces cerevisiae and considerable substrate diversity in its fungal family. Nat. Commun. 8, 14247 (2017).
pubmed: 28165463 pmcid: 5303821 doi: 10.1038/ncomms14247
Isnard, A. D., Thomas, D. & Surdin-Kerjan, Y. The study of methionine uptake in Saccharomyces cerevisiae reveals a new family of amino acid permeases. J. Mol. Biol. 262, 473–484 (1996).
pubmed: 8893857 doi: 10.1006/jmbi.1996.0529
Linder, T. Genomics of alternative sulfur utilization in ascomycetous yeasts. Microbiology 158, 2585–2597 (2012).
pubmed: 22790398 doi: 10.1099/mic.0.060285-0
Linder, T. Assimilation of alternative sulfur sources in fungi. World J. Microbiol. Biotechnol. 34, 51 (2018).
pubmed: 29550883 pmcid: 5857272 doi: 10.1007/s11274-018-2435-6
Bailey, T. L., Johnson, J., Grant, C. E. & Noble, W. S. The MEME Suite. Nucleic Acids Res. 43, W39–W49 (2015).
pubmed: 25953851 pmcid: 4489269 doi: 10.1093/nar/gkv416
Thomas, P. D. et al. PANTHER: Making genome-scale phylogenetics accessible to all. Protein Sci. 31, 8–22 (2022).
pubmed: 34717010 doi: 10.1002/pro.4218
Lavoie, H. et al. Evolutionary tinkering with conserved components of a transcriptional regulatory network. PLOS Biol. 8, e1000329 (2010).
pubmed: 20231876 pmcid: 2834713 doi: 10.1371/journal.pbio.1000329
Schrevens, S. et al. Methionine is required for cAMP-PKA-mediated morphogenesis and virulence of Candida albicans. Mol. Microbiol. 108, 258–275 (2018).
pubmed: 29453849 doi: 10.1111/mmi.13933
Siggers, T., Duyzend, M. H., Reddy, J., Khan, S. & Bulyk, M. L. Non-DNA-binding cofactors enhance DNA-binding specificity of a transcriptional regulatory complex. Mol. Syst. Biol. 7, 555 (2011).
pubmed: 22146299 pmcid: 3737730 doi: 10.1038/msb.2011.89
Autry, A. R. & Fitzgerald, J. W. Sulfonate S: A major form of forest soil organic sulfur. Biol. Fertil. Soils 10, 50–56 (1990).
doi: 10.1007/BF00336124
Hall, C., Brachat, S. & Dietrich, F. S. Contribution of horizontal gene transfer to the evolution of Saccharomyces cerevisiae. Eukaryot. Cell 4, 1102–1115 (2005).
pubmed: 15947202 pmcid: 1151995 doi: 10.1128/EC.4.6.1102-1115.2005
Hogan, D. A., Auchtung, T. A. & Hausinger, R. P. Cloning and characterization of a sulfonate/alpha-ketoglutarate dioxygenase from saccharomyces cerevisiae. J. Bacteriol. 181, 5876–5879 (1999).
pubmed: 10482536 pmcid: 94115 doi: 10.1128/JB.181.18.5876-5879.1999
Teigen, L. M. et al. Dietary factors in sulfur metabolism and pathogenesis of Ulcerative Colitis. Nutrients 11, 931 (2019).
pubmed: 31027194 pmcid: 6521024 doi: 10.3390/nu11040931
Peng, D. & Tarleton, R. EuPaGDT: a web tool tailored to design CRISPR guide RNAs for eukaryotic pathogens. Micro. Genom. 1, e000033 (2015).
Lombardi, L. & Butler, G. Plasmid-based CRISPR-Cas9 editing in multiple Candida species. Methods Mol. Biol. 2542, 13–40 (2022).
pubmed: 36008654 doi: 10.1007/978-1-0716-2549-1_2
Cravener, M. V. & Mitchell, A. P. Candida albicans culture, cell harvesting, and total RNA extraction. Bio Protoc. 10, e3803 (2020).
pubmed: 33659457 pmcid: 7854013 doi: 10.21769/BioProtoc.3803
Guida, A. et al. Using RNA-seq to determine the transcriptional landscape and the hypoxic response of the pathogenic yeast Candida parapsilosis. BMC Genomics 12, 628 (2011).
pubmed: 22192698 pmcid: 3287387 doi: 10.1186/1471-2164-12-628
Dobin, A. et al. STAR: ultrafast universal RNA-seq aligner. Bioinformatics 29, 15–21 (2013).
pubmed: 23104886 doi: 10.1093/bioinformatics/bts635
Kuhn, D. M., Chandra, J., Mukherjee, P. K. & Ghannoum, M. A. Comparison of biofilms formed by Candida albicans and Candida parapsilosis on bioprosthetic surfaces. Infect. Immun. 70, 878–888 (2002).
pubmed: 11796623 pmcid: 127692 doi: 10.1128/IAI.70.2.878-888.2002
Anders, S. & Huber, W. Differential expression analysis for sequence count data. Genome Biol. 11, R106 (2010).
pubmed: 20979621 pmcid: 3218662 doi: 10.1186/gb-2010-11-10-r106

Auteurs

Lisa Lombardi (L)

School of Biomolecular and Biomedical Science, Conway Institute, University College Dublin, Belfield, Dublin, Ireland. lisa.lombardi@ucd.ie.

Letal I Salzberg (LI)

School of Medicine, Conway Institute, University College Dublin, Belfield, Dublin, Ireland.

Eoin Ó Cinnéide (EÓ)

School of Biomolecular and Biomedical Science, Conway Institute, University College Dublin, Belfield, Dublin, Ireland.

Caoimhe O'Brien (C)

School of Biomolecular and Biomedical Science, Conway Institute, University College Dublin, Belfield, Dublin, Ireland.

Florent Morio (F)

Nantes Université, CHU Nantes, Cibles et Médicaments des Infections et de l'Immunité, UR1155, Nantes, France.

Siobhán A Turner (SA)

School of Biomolecular and Biomedical Science, Conway Institute, University College Dublin, Belfield, Dublin, Ireland.

Kevin P Byrne (KP)

School of Medicine, Conway Institute, University College Dublin, Belfield, Dublin, Ireland.

Geraldine Butler (G)

School of Biomolecular and Biomedical Science, Conway Institute, University College Dublin, Belfield, Dublin, Ireland. gbutler@ucd.ie.

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