Recent genetic connectivity and clinal variation in chimpanzees.


Journal

Communications biology
ISSN: 2399-3642
Titre abrégé: Commun Biol
Pays: England
ID NLM: 101719179

Informations de publication

Date de publication:
05 03 2021
Historique:
received: 25 06 2020
accepted: 04 02 2021
entrez: 6 3 2021
pubmed: 7 3 2021
medline: 11 8 2021
Statut: epublish

Résumé

Much like humans, chimpanzees occupy diverse habitats and exhibit extensive behavioural variability. However, chimpanzees are recognized as a discontinuous species, with four subspecies separated by historical geographic barriers. Nevertheless, their range-wide degree of genetic connectivity remains poorly resolved, mainly due to sampling limitations. By analyzing a geographically comprehensive sample set amplified at microsatellite markers that inform recent population history, we found that isolation by distance explains most of the range-wide genetic structure of chimpanzees. Furthermore, we did not identify spatial discontinuities corresponding with the recognized subspecies, suggesting that some of the subspecies-delineating geographic barriers were recently permeable to gene flow. Substantial range-wide genetic connectivity is consistent with the hypothesis that behavioural flexibility is a salient driver of chimpanzee responses to changing environmental conditions. Finally, our observation of strong local differentiation associated with recent anthropogenic pressures portends future loss of critical genetic diversity if habitat fragmentation and population isolation continue unabated.

Identifiants

pubmed: 33674780
doi: 10.1038/s42003-021-01806-x
pii: 10.1038/s42003-021-01806-x
pmc: PMC7935964
doi:

Types de publication

Comparative Study Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

283

Références

Serre, D. & Paabo, S. Evidence for gradients of human genetic diversity within and among continents. Genome Res. 14, 1679–1685 (2004).
pubmed: 15342553 pmcid: 515312 doi: 10.1101/gr.2529604
Ohta, T. Population size and rate of evolution. J. Mol. Evol. 1, 305–314 (1972).
pubmed: 24173491 doi: 10.1007/BF01653959
Slatkin, M. Gene flow and selection in a cline. Genetics 75, 733–756 (1973).
Potts, R. Variability selection in hominid evolution. Evol. Anthropol. 7, 81–96 (1998).
doi: 10.1002/(SICI)1520-6505(1998)7:3<81::AID-EVAN3>3.0.CO;2-A
Potts, R. Hominin evolution in settings of strong environmental variability. Quat. Sci. Rev. 73, 1–13 (2013).
doi: 10.1016/j.quascirev.2013.04.003
Prüfer, K. et al. The bonobo genome compared with the chimpanzee and human genomes. Nature 486, 527–531 (2012).
pubmed: 22722832 pmcid: 3498939 doi: 10.1038/nature11128
Hill, W. C. O. in The Nomenclature, Taxonomy and Distribution of Chimpanzees, Vol. 1 (ed. Bourne, G. H.) 22–49 (Karger, 1969).
Pruetz, J. D. & Bertolani, P. Chimpanzee (Pan troglodytes verus) behavioral responses to stresses associated with living in a savannah-mosaic environment: implications for hominin adaptations to open habitats. Paleoanthropology 2009, 252–262 (2009).
doi: 10.4207/PA.2009.ART33
Fünfstück, T. et al. The sampling scheme matters: Pan troglodytes troglodytes and P. t. schweinfurthii are characterized by clinal genetic variation rather than a strong subspecies break. Am. J. Phys. Anthropol. 156, 181–191 (2014).
pubmed: 25330245 pmcid: 4692250 doi: 10.1002/ajpa.22638
Fischer, A., Pollack, J., Thalmann, O., Nickel, B. & Paabo, S. Demographic history and genetic differentiation in apes. Curr. Biol. 16, 1133–1138 (2006).
pubmed: 16753568 doi: 10.1016/j.cub.2006.04.033
Gonder, M. K. et al. A new west African chimpanzee subspecies? Nature 388, 337 (1997).
pubmed: 9237749 doi: 10.1038/41005
Becquet, C., Patterson, N., Stone, A. C., Przeworski, M. & Reich, D. E. Genetic structure of chimpanzee populations. PLoS Genet. 3, e66 (2007).
pubmed: 17447846 pmcid: 1853122 doi: 10.1371/journal.pgen.0030066
Bowden, R. et al. Genomic tools for evolution and conservation in the chimpanzee: Pan troglodytes ellioti is a genetically distinct population. PLoS Genet. 8, e1002504 (2012).
pubmed: 22396655 pmcid: 3291532 doi: 10.1371/journal.pgen.1002504
Prado-Martinez, J. et al. Great ape genetic diversity and population history. Nature 499, 471–475 (2013).
pubmed: 23823723 pmcid: 3822165 doi: 10.1038/nature12228
de Manuel, M. et al. Chimpanzee genomic diversity reveals ancient admixture with bonobos. Science 354, 477–481 (2016).
pubmed: 27789843 pmcid: 5546212 doi: 10.1126/science.aag2602
Langergraber, K. E. et al. Genetic and ‘cultural’ similarity in wild chimpanzees. Proc. R. Soc. B. 278, 408–416 (2010).
pubmed: 20719777 doi: 10.1098/rspb.2010.1112 pmcid: 3013405
Garcin, Y. et al. Early anthropogenic impact on Western Central African rainforests 2,600 y ago. Proc. Natl Acad. Sci. USA 115, 3261–3266 (2018).
pubmed: 29483260 doi: 10.1073/pnas.1715336115 pmcid: 5879660
Vicente, M. & Schlebusch, C. M. African population history: an ancient DNA perspective. Curr. Opin. Genet. Dev. 62, 8–15 (2020).
pubmed: 32563853 doi: 10.1016/j.gde.2020.05.008
IUCN Red List. IUCN red list of threatened species. http://www.iucnredlist.org (2020).
Wentworth, C. K. Natural bridges and glaciation. Am. J. Sci. 26, 577–584 (1933).
doi: 10.2475/ajs.s5-26.156.577
Maley, J. The African rainforest: main characteristics of changes in vegetation and climate from the Upper Cretaceous to the Quaternary. Proc. R. Soc. Edinb. 104B, 31–73 (1996).
Langergraber, K. E. et al. Generation times in wild chimpanzees and gorillas suggest earlier divergence times in great ape and human evolution. Proc. Natl Acad. Sci. USA 109, 15716–15721 (2012).
pubmed: 22891323 doi: 10.1073/pnas.1211740109 pmcid: 3465451
Shea, B. T. & Coolidge, H. J. Craniometric differentiation and systematics in the genus Pan. J. Hum. Evol. 13, 671–685 (1988).
doi: 10.1016/0047-2484(88)90024-3
Albrecht, G. H. & Miller, J. M. A. in Geographic Variation in Primates (eds Kimbel, W. H. & Martin, L. B.) 123–161 (Springer Science & Business Media, 2013).
Shea, B. T., Leigh, S. R. & Groves, C. P. in Multivariate Craniometric Variation in Chimpanzees (eds Kimbel, W. H. & Martin, L. B.) 265–296 (Springer Science & Business Media, 2013).
Kühl, H. S. et al. Chimpanzee accumulative stonethrowing. Sci. Rep. 6, 22219 (2016).
Wright, S. Isolation by distance. Genetics 28, 114–138 (1943).
pubmed: 17247074 pmcid: 1209196 doi: 10.1093/genetics/28.2.114
Pritchard, J. K., Stephens, M. & Donnelly, P. Inference of population structure using multilocus genotype data. Genetics 155, 945–959 (2000).
pubmed: 10835412 pmcid: 1461096 doi: 10.1093/genetics/155.2.945
Pritchard, J. K., Wen, X. & Falush, D. Documentation for structure version 2.3 software: version 2.3. 1–39. http://pritchardlab.stanford.edu/structure_software/release_versions/v2.3.4/structure_doc.pdf (2009).
Meirmans, P. G. The trouble with isolation by distance. Mol. Ecol. 21, 2839–2846 (2012).
pubmed: 22574758 doi: 10.1111/j.1365-294X.2012.05578.x
Perez, M. F. et al. Assessing population structure in the face of isolation by distance: are we neglecting the problem? Divers. Distrib. 24, 1883–1889 (2018).
doi: 10.1111/ddi.12816
Thalib, L., Kitching, R. L. & Bhatti, M. I. Principal component analysis for grouped data - a case study. Environmetrics 10, 565–574 (1999).
doi: 10.1002/(SICI)1099-095X(199909/10)10:5<565::AID-ENV360>3.0.CO;2-R
Fischer, A. et al. Bonobos fall within the genomic variation of chimpanzees. PLoS ONE 6, e21605 (2011).
pubmed: 21747915 pmcid: 3126833 doi: 10.1371/journal.pone.0021605
Frantz, A. C., Cellina, S., Krier, A., Schley, L. & Burke, T. Using spatial Bayesian methods to determine the genetic structure of a continuously distributed population: clusters or isolation by distance? J. Appl. Ecol. 46, 493–505 (2009).
doi: 10.1111/j.1365-2664.2008.01606.x
Kalinowski, S. T. The computer program STRUCTURE does not reliably identify the main genetic clusters within species: simulations and implications for human population structure. Heredity 106, 625–632 (2010).
pubmed: 20683484 pmcid: 3183908 doi: 10.1038/hdy.2010.95
Schwartz, M. K. & McKelvey, K. S. Why sampling scheme matters: the effect of sampling scheme on landscape genetic results. Conserv. Genet. 10, 441–452 (2008).
doi: 10.1007/s10592-008-9622-1
Meirmans, P. G. Seven common mistakes in population genetics and how to avoid them. Mol. Ecol. 24, 3223–3231 (2015).
pubmed: 25974103 doi: 10.1111/mec.13243
Arandjelovic M, et al. Pan African Programme—The cultured chimpanzee. Guidelines for research and data collection. http://panafrican.eva.mpg.de/english/approaches_and_methods.php (2014).
Arandjelovic, M. & Vigilant, L. Non-invasive genetic censusing and monitoring of primate populations. Am. J. Primatol. 80, e22743 (2018).
pubmed: 29457631 doi: 10.1002/ajp.22743
Arthofer, W., Heussler, C., Krapf, P., Schlick-Steiner, B. C. & Steiner, F. M. Identifying the minimum number of microsatellite loci needed to assess population genetic structure: a case study in fly culturing. Fly 12, 13–22 (2018).
Wenburg, J. K., Bentzen, P. & Foote, C. J. Microsatellite analysis of genetic population structure in an endangered salmonid: the coastal cutthroat trout (Oncorhyncus clarki clarki). Mol. Ecol. 7, 733–749 (1998).
doi: 10.1046/j.1365-294x.1998.00386.x
Guo, X.-Z. et al. Phylogeography and populationgenetics of Schizothorax o’connori: strong subdivision in the Yarlung Tsangpo River inferred from mtDNA and microsatellite markers. Sci. Rep. 6, 29821 (2016).
Kleinhans, C. & Willows-Munro, S. Low genetic diversity andshallow population structure inthe endangered vulture, Gyps copotheres. Sci. Rep. 9, 5536 (2019).
pubmed: 30940898 pmcid: 6445149 doi: 10.1038/s41598-019-41755-4
Bonato, L. et al. Diversity among peripheral populations: genetic and evolutionary differentiation of Salamandra atraat the southern edge of the Alps. J. Zool. Syst. Evol. Res. 56, 533–548 (2018).
doi: 10.1111/jzs.12224
Balkenhol, N. et al. A multi-method approach for analyzing hierarchical genetic structures: a case study with cougars Puma concolor. Ecography 37, 552–563 (2014).
doi: 10.1111/j.1600-0587.2013.00462.x
Kobayashi, T. & Sota, T. Contrasting effects of habitat discontinuity on three closely related fungivorous beetle species with diverging host‐use patterns and dispersal ability. Ecol. Evol. 9, 2475–2486 (2019).
pubmed: 30891194 pmcid: 6405892 doi: 10.1002/ece3.4862
Rousset, F. Genetic differentiation and estimation of gene flow from F-statistics under isolation by distance. Genetics 145, 1219–1228 (1997).
Valdes, A. M., Slatkin, M. & Freimer, N. B. Allele frequencies at microsatellite loci: the stepwise mutation model revisited. Genetics 133, 737–749 (1993).
pubmed: 8454213 pmcid: 1205356 doi: 10.1093/genetics/133.3.737
Goldstein, D. B., Ruiz Linares, A., Cavalli-Sforza, L. L. & Feldman, M. W. An evaluation of genetic distances for use with microsatellite loci. Genetics 139, 463–471 (1995).
pubmed: 7705647 pmcid: 1206344 doi: 10.1093/genetics/139.1.463
Calabrese, P. P., Durrett, R. T. & Aquadro, C. F. Dynamics of microsatellite divergence under stepwise mutation and proportional slippage/point mutation models. Genetics 159, 839–852 (2001).
pubmed: 11606557 pmcid: 1461831 doi: 10.1093/genetics/159.2.839
Petkova, D., Novembre, J. & Stephens, M. Visualizing spatial population structure with estimated effective migration surfaces. Nat. Genet. 48, 94–100 (2015).
pubmed: 26642242 pmcid: 4696895 doi: 10.1038/ng.3464
Rich, A. M., Wasserman, M. D., Hunt, K. D. & Kaestle, F. A. Chimpanzee (Pan troglodytes schweinfurthii) population spans multiple protected areas in the Albertine Rift. Folia. Primatol. 91, 595–609 (2020).
Baldwin, P. J., McGrew, W. C. & Tutin, C. E. G. Wide-ranging chimpanzees at Mt. Assirik, Senegal. Int. J. Primatol. 3, 367–385 (1982).
doi: 10.1007/BF02693739
Lemoine, S. et al. Group dominance increases territory size and reduces neighbour pressure in wild chimpanzees. R. Soc. Open Sci. 7, 200577 (2020).
pubmed: 32537232 pmcid: 7277268 doi: 10.1098/rsos.200577
Newton-Fisher, N. E. The home range of the Sonso community of chimpanzees from the Budongo Forest, Uganda. Afr. J. Ecol. 41, 150–156 (2003).
doi: 10.1046/j.1365-2028.2003.00408.x
Allendorf, F. W. Genetic drift and the loss of alleles versus heterozygosity. Zoo. Biol. 5, 181–190 (1986).
doi: 10.1002/zoo.1430050212
Barratt, C. D., et al. Late Quaternary habitat suitability models for chimpanzees (Pan troglodytes) since the Last Interglacial (120,000 BP). Preprint at BioRxiv https://www.biorxiv.org/content/10.1101/2020.05.15.066662v1 (2019).
Bertola, L. D. et al. Phylogeographic patterns in Africa and high resolution delineation of genetic clades in the lion (Panthera leo). Sci. Rep. 6, 30807 (2016).
Marchesi, P., Marchesi, N., Fruth, B. & Boesch, C. Census and distribution of chimpanzees in Côte D’Ivoire. Primates 36, 591–607 (1995).
doi: 10.1007/BF02382880
Sommer, V., Adanu, J., Faucher, I. & Fowler, A. Nigerian chimpanzees (Pan troglodytes vellerosus) at Gashaka: two years of habituation efforts. Folia Primatol. 75, 295–316 (2004).
doi: 10.1159/000080208
Chancellor, R. L., Langergraber, K. E., Ramirez, S., Rundus, A. S. & Vigilant, L. Genetic sampling of unhabituated chimpanzees (Pan troglodytes schweinfurthii) in Gishwati Forest Reserve, an isolated forest fragment in western Rwanda. Int. J. Primatol. 33, 479–488 (2012).
doi: 10.1007/s10764-012-9591-6
Piel, A. K. et al. Population status of chimpanzees in the Masito-Ugalla Ecosystem, Tanzania. Am. J. Primatol. 77, 1027–1035 (2015).
pubmed: 26119006 doi: 10.1002/ajp.22438
Wessling, E. G. et al. Seasonal variation in physiology challenges the notion of chimpanzees as a forest-adapted species. Front. Ecol. Evol. 6, 1–21 (2018).
doi: 10.3389/fevo.2018.00060
Whiten, A. et al. Cultures in chimpanzees. Nature 399, 682–685 (1999).
pubmed: 10385119 doi: 10.1038/21415
Kühl, H. S. et al. Human impact erodes chimpanzee behavioral diversity. Science 363, 1453–1455 (2019).
pubmed: 30846610 doi: 10.1126/science.aau4532
Luncz, L. V., Mundry, R. & Boesch, C. Evidence for cultural differences between neighboring chimpanzee communities. Curr. Biol. 22, 922–926 (2012).
pubmed: 22578420 doi: 10.1016/j.cub.2012.03.031
Boesch, C., Marchesi, P., Marchesi, N., Fruth, B. & Joulian, F. Is nut cracking in wild chimpanzees a cultural behaviour? J. Hum. Evol. 26, 325–338 (1994).
doi: 10.1006/jhev.1994.1020
Kalan, A. K. et al. Environmental variability supports chimpanzee behavioural diversity. Nat. Commun. 11, 4451 (2020).
Hockings, K. J. in Chimpanzees of Bossou and Nimba, 221–219 (Springer Science & Business Media, 2011).
McCarthy, M. S., Lester, J. D. & Stanford, C. B. Chimpanzees (Pan troglodytes) flexibly use introduced species for nesting and bark feeding in a human-dominated habitat. Int. J. Primatol. 38, 321–337 (2016).
pubmed: 28546652 pmcid: 5422490 doi: 10.1007/s10764-016-9916-y
McLennan, M. R. et al. Surviving at the extreme: chimpanzee ranging is not restricted in a deforested human‐dominated landscape in Uganda. Afr. J. Ecol. 8, e57872 (2020).
Junker, J. et al. Recent decline in suitable environmental conditions for African great apes. Divers. Distrib. 18, 1077–1091 (2012).
doi: 10.1111/ddi.12005
Kühl, H. S. et al. The Critically Endangered western chimpanzee declines by 80%. Am. J. Primatol. 79, e22681 (2017).
doi: 10.1002/ajp.22681
Walsh, P. D., Breuer, T., Sanz, C., Morgan, D. B. & Doran-Sheehy, D. M. Potential for Ebola transmission between gorilla and chimpanzee social groups. Am. Nat. 169, 684–689 (2007).
pubmed: 17427138 doi: 10.1086/513494
Baden, A. L. et al. Anthropogenic pressures drive population genetic structuring across a critically endangered lemur species range. Sci. Rep. 9, 16276 (2019).
pubmed: 31700150 pmcid: 6838192 doi: 10.1038/s41598-019-52689-2
Nsubuga, A. M. et al. Factors affecting the amount of genomic DNA extracted from ape faeces and the identification of an improved sample storage method. Mol. Ecol. 13, 2089–2094 (2004).
pubmed: 15189228 doi: 10.1111/j.1365-294X.2004.02207.x
Arandjelovic, M. et al. Two-step multiplex polymerase chain reaction improves the speed and accuracy of genotyping using DNA from noninvasive and museum samples. Mol. Ecol. Resour. 9, 28–36 (2009).
pubmed: 21564562 doi: 10.1111/j.1755-0998.2008.02387.x
Kalinowski, S. T., Taper, M. L. & Marshall, T. C. Revising how the computer program CERVUS accommodates genotyping error increases success in paternity assignment. Mol. Ecol. 16, 1099–1106 (2007).
pubmed: 17305863 doi: 10.1111/j.1365-294X.2007.03089.x
Waits, L. P., Luikart, G. & Taberlet, P. Estimating the probability of identity among genotypes in natural populations: cautions and guidelines. Mol. Ecol. 10, 249–256 (2001).
pubmed: 11251803 doi: 10.1046/j.1365-294X.2001.01185.x
Meirmans, P. G. & Hedrick, P. W. Assessing population structure: FST and related measures. Mol. Ecol. Resour. 11, 5–18 (2010).
pubmed: 21429096 doi: 10.1111/j.1755-0998.2010.02927.x
Mondol, S. et al. New evidence for hybrid zones of forest and savanna elephants in Central and West Africa. Mol. Ecol. 24, 6134–6147 (2015).
pubmed: 26577954 doi: 10.1111/mec.13472

Auteurs

Jack D Lester (JD)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany. jack_lester@eva.mpg.de.

Linda Vigilant (L)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Paolo Gratton (P)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Maureen S McCarthy (MS)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Christopher D Barratt (CD)

German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Leipzig, Germany.

Paula Dieguez (P)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Anthony Agbor (A)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Paula Álvarez-Varona (P)

Jane Goodall Institute Spain and Senegal, Dindefelo Biological Station, Dindefelo, Kedougou, Senegal.

Samuel Angedakin (S)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Emmanuel Ayuk Ayimisin (EA)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Emma Bailey (E)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Mattia Bessone (M)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Gregory Brazzola (G)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Rebecca Chancellor (R)

West Chester University, Depts of Anthropology & Sociology and Psychology, West Chester, PA, USA.

Heather Cohen (H)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Emmanuel Danquah (E)

Department of Wildlife and Range Management, Faculty of Renewable Natural Resources, Kwame Nkrumah University of Science and Technology, Kumasi, Ghana.

Tobias Deschner (T)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Villard Ebot Egbe (VE)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Manasseh Eno-Nku (M)

WWF Cameroon Country Programme Office, Yaoundé, Cameroon.

Annemarie Goedmakers (A)

Chimbo Foundation, Amsterdam, Netherlands.

Anne-Céline Granjon (AC)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Josephine Head (J)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Daniela Hedwig (D)

Elephant Listening Project, Center for Conservation Bioacoustics, Cornell Lab of Ornithology, Cornell University, Ithaca, NY, USA.

R Adriana Hernandez-Aguilar (RA)

Jane Goodall Institute Spain and Senegal, Dindefelo Biological Station, Dindefelo, Kedougou, Senegal.
Department of Social Psychology and Quantitative Psychology, Faculty of Psychology, University of Barcelona, Barcelona, Spain.

Kathryn J Jeffery (KJ)

Biological and Environmental Sciences, Faculty of Natural Sciences, University of Stirling, Stirling, UK.

Sorrel Jones (S)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Jessica Junker (J)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Parag Kadam (P)

University of Cambridge, Cambridge, UK.

Michael Kaiser (M)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Ammie K Kalan (AK)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Laura Kehoe (L)

Wild Chimpanzee Foundation (WCF), Leipzig, Germany.

Ivonne Kienast (I)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Kevin E Langergraber (KE)

School of Human Evolution and Social Change, Arizona State University, 900 Cady Mall, Tempe, AZ 85287 Arizona State University, Tempe, AZ, USA.

Juan Lapuente (J)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.
Comoé Chimpanzee Conservation Project, Comoé National Park, Kakpin, Côte d'Ivoire.

Anne Laudisoit (A)

Ecohealth Alliance, New York, NY, USA.
University of Antwerp, Campus Drie Eiken, lokaal D.133, Universiteitsplein 1 - 2610, Antwerpen, Belgium.

Kevin Lee (K)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Sergio Marrocoli (S)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Vianet Mihindou (V)

Agence National des Parcs Nationaux (ANPN) Batterie 4, Libreville, Gabon.
Ministère des Eaux, des Forêts, de la Mer, de l'Environnement, Chargé du Plan Climat, des Objectifs de Développement Durable et du Plan d'Affectation des Terres, Libreville, Gabon.

David Morgan (D)

Lester E. Fisher Center for the Study and Conservation of Apes, Lincoln Park Zoo, Chicago, IL, USA.

Geoffrey Muhanguzi (G)

Budongo Conservation Field Station, Masindi, Uganda.

Emily Neil (E)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Sonia Nicholl (S)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Christopher Orbell (C)

Panthera, New York, NY, USA.

Lucy Jayne Ormsby (LJ)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Liliana Pacheco (L)

Jane Goodall Institute Spain and Senegal, Dindefelo Biological Station, Dindefelo, Kedougou, Senegal.

Alex Piel (A)

Department of Anthropology, University College London, London, UK.

Martha M Robbins (MM)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Aaron Rundus (A)

West Chester University, Department of Psychology, West Chester, PA, USA.

Crickette Sanz (C)

Washington University in Saint Louis, Department of Anthropology, One Brookings Drive, St. Louis, MO, USA.
Wildlife Conservation Society, Congo Program, Brazzaville, Republic of Congo.

Lilah Sciaky (L)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Alhaji M Siaka (AM)

National Protected Area Authority, Freetown, Sierra Leone.

Veronika Städele (V)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Fiona Stewart (F)

School of Biological & Environmental Sciences, Liverpool John Moores University, Liverpool, UK.

Nikki Tagg (N)

KMDA, Centre for Research and Conservation, Royal Zoological Society of Antwerp, Antwerp, Belgium.

Els Ton (E)

Chimbo Foundation, Amsterdam, Netherlands.

Joost van Schijndel (J)

Chimbo Foundation, Amsterdam, Netherlands.

Magloire Kambale Vyalengerera (MK)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Erin G Wessling (EG)

Department of Human Evolutionary Biology, Harvard University, Cambridge, MA, USA.

Jacob Willie (J)

KMDA, Centre for Research and Conservation, Royal Zoological Society of Antwerp, Antwerp, Belgium.

Roman M Wittig (RM)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.
Taï Chimpanzee Project, Centre Suisse de Recherches Scientifiques, Abidjan, Côte d'Ivoire.

Yisa Ginath Yuh (YG)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Kyle Yurkiw (K)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.
Pan Verus Project Outamba-Kilimi National Park, Freetown, Sierra Leone.

Klaus Zuberbuehler (K)

Budongo Conservation Field Station, Masindi, Uganda.
Université de Neuchâtel, Institut de Biologie, Neuchâtel, Switzerland.
School of Psychology and Neuroscience, University of St Andrews, St Andrews, UK.

Christophe Boesch (C)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.

Hjalmar S Kühl (HS)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany.
German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Leipzig, Germany.

Mimi Arandjelovic (M)

Max Planck Institute for Evolutionary Anthropology (MPI EVAN), Leipzig, Germany. arandjel@eva.mpg.de.

Articles similaires

Genome, Chloroplast Phylogeny Genetic Markers Base Composition High-Throughput Nucleotide Sequencing
Robotic Surgical Procedures Animals Humans Telemedicine Models, Animal

Odour generalisation and detection dog training.

Lyn Caldicott, Thomas W Pike, Helen E Zulch et al.
1.00
Animals Odorants Dogs Generalization, Psychological Smell
Animals TOR Serine-Threonine Kinases Colorectal Neoplasms Colitis Mice

Classifications MeSH