Overexpression of thioredoxin m in chloroplasts alters carbon and nitrogen partitioning in tobacco.


Journal

Journal of experimental botany
ISSN: 1460-2431
Titre abrégé: J Exp Bot
Pays: England
ID NLM: 9882906

Informations de publication

Date de publication:
22 06 2021
Historique:
received: 19 01 2021
accepted: 27 04 2021
pubmed: 9 5 2021
medline: 10 8 2021
entrez: 8 5 2021
Statut: ppublish

Résumé

In plants, there is a complex interaction between carbon (C) and nitrogen (N) metabolism, and its coordination is fundamental for plant growth and development. Here, we studied the influence of thioredoxin (Trx) m on C and N partitioning using tobacco plants overexpressing Trx m from the chloroplast genome. The transgenic plants showed altered metabolism of C (lower leaf starch and soluble sugar accumulation) and N (with higher amounts of amino acids and soluble protein), which pointed to an activation of N metabolism at the expense of carbohydrates. To further delineate the effect of Trx m overexpression, metabolomic and enzymatic analyses were performed on these plants. These results showed an up-regulation of the glutamine synthetase-glutamate synthase pathway; specifically tobacco plants overexpressing Trx m displayed increased activity and stability of glutamine synthetase. Moreover, higher photorespiration and nitrate accumulation were observed in these plants relative to untransformed control plants, indicating that overexpression of Trx m favors the photorespiratory N cycle rather than primary nitrate assimilation. Taken together, our results reveal the importance of Trx m as a molecular mediator of N metabolism in plant chloroplasts.

Identifiants

pubmed: 33963398
pii: 6272276
doi: 10.1093/jxb/erab193
pmc: PMC8219043
doi:

Substances chimiques

Chloroplast Thioredoxins 0
Carbon 7440-44-0
Nitrogen N762921K75

Types de publication

Journal Article Research Support, Non-U.S. Gov't

Langues

eng

Sous-ensembles de citation

IM

Pagination

4949-4964

Informations de copyright

© The Author(s) 2021. Published by Oxford University Press on behalf of the Society for Experimental Biology.

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Auteurs

María Ancín (M)

Institute for Multidisciplinary Applied Biology (IMAB), Dpto. Agronomía, Biotecnología y Alimentación, Universidad Publica de Navarra (UPNA), Campus Arrosadia, 31006 Pamplona, Spain.

Luis Larraya (L)

Institute for Multidisciplinary Applied Biology (IMAB), Dpto. Agronomía, Biotecnología y Alimentación, Universidad Publica de Navarra (UPNA), Campus Arrosadia, 31006 Pamplona, Spain.

Igor Florez-Sarasa (I)

Max-Planck-Institut für Molekulare Pflanzenphysiologie, Am Mühlenberg 1, 14476 Potsdam-Golm, Germany.
Centre for Research in Agricultural Genomics (CRAG) CSIC-IRTA-UAB-UB, Campus UAB Bellaterra, 08193 Barcelona, Spain.

Camille Bénard (C)

UMR 1332 Biologie du Fruit et Pathologie and Plateforme Metabolome Bordeaux, INRA, Bordeaux University, 33882 Villenave d'Ornon, France.

Alicia Fernández-San Millán (A)

Institute for Multidisciplinary Applied Biology (IMAB), Dpto. Agronomía, Biotecnología y Alimentación, Universidad Publica de Navarra (UPNA), Campus Arrosadia, 31006 Pamplona, Spain.

Jon Veramendi (J)

Institute for Multidisciplinary Applied Biology (IMAB), Dpto. Agronomía, Biotecnología y Alimentación, Universidad Publica de Navarra (UPNA), Campus Arrosadia, 31006 Pamplona, Spain.

Yves Gibon (Y)

UMR 1332 Biologie du Fruit et Pathologie and Plateforme Metabolome Bordeaux, INRA, Bordeaux University, 33882 Villenave d'Ornon, France.

Alisdair R Fernie (AR)

Max-Planck-Institut für Molekulare Pflanzenphysiologie, Am Mühlenberg 1, 14476 Potsdam-Golm, Germany.

Iker Aranjuelo (I)

Instituto de Agrobiotecnología, CSIC-Gobierno de Navarra, Avda. Pamplona 123, 31192 Mutilva, Spain.

Inmaculada Farran (I)

Institute for Multidisciplinary Applied Biology (IMAB), Dpto. Agronomía, Biotecnología y Alimentación, Universidad Publica de Navarra (UPNA), Campus Arrosadia, 31006 Pamplona, Spain.

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Classifications MeSH